CATALOGUE OF THE MAMMALS OF ETHIOPIA

Transcription

1 Monitore Zoologico Italiano. Supplemento ISSN: (Print) (Online) Journal homepage: CATALOGUE OF THE MAMMALS OF ETHIOPIA M. J. Largen, D. Kock & D. W. Yalden To cite this article: M. J. Largen, D. Kock & D. W. Yalden (1974) CATALOGUE OF THE MAMMALS OF ETHIOPIA, Monitore Zoologico Italiano. Supplemento, 5:1, To link to this article: Published online: 12 Jul Submit your article to this journal Article views: 114 View related articles Citing articles: 7 View citing articles Full Terms & Conditions of access and use can be found at Download by: [ ] Date: 02 January 2018, At: 10:17

2 Monitore Zoologico Italiano ITALIAN JOURNAL OF ZOOLOGY PUBBLICATO DALLA UNIVERSITJI. DEGLI STUDI Dl FIRENZE CON IL CONTRIBUTO DEL CONSIGLIO NAZIONALE DELLE RICERCHE N. S. SUPPLEMENTO V NO. 16 : CATALOGUE OF THE MAMMALS OF ETHIOPIA 1. CHIROPTERA ( PUBBLICAZIONI DEL CENTRO Dl STUDIO PER LA FAUNISTICA ED ECOLOGIA TROPICAL! DEL C.N.R.: LXXXIII) M. J. LARGEN, D. KocK and D. W. YALDEN Natural History Museum, Haile Selassie I University, Addis Ababa Forschungsinstitut Senckenberg, Frankfurt a.m. Department of Zoology, University of Manchester Received 12 April 1974 I. Introduction page 223 II. Suborder Megachiroptera» 225 A. Family Pteropidae.» Hypsignathus monstrosus H. Allen, 1861 >> Epomophorus minor Dobson, » Epomophorus labiatus labiatus (Temminck, 1837) >> Epomophorus labiatus anurus Heuglin, >> Epomophorus gambianus gambianus (Ogilby, 1835).» Micropteropus pusillus (Peters, 1868) >> Eidolon helvum (Kerr, 1792)... >> Rousettus aegyptiacus (E. Geoffroy, 1810) >> Rousettus angolensis (Bocage, 1898) >> Rousettus lanosus kempi Thomas, 1909 >> 230 III. Suborder Microchiroptera >> 230 A. Family Rhinopomatidae. >> Rhinopoma hardwickei sennaariense Fitzinger, >> Rhinopoma muscatellum Thomas, 1903 >> 230 B. Family Emballonuridae. >> Taphozous perforatus E. Geoffroy, >> Taphozous nudiventris Cretzschmar, >> Taphozous mauritianus E. Geoffroy, 1818 >> Coleura afra (Peters, 1852) >> 232 C. Family Nycteridae. >> Nycteris thebaica E. Geoffroy, 1813 )) Nycteris hispida hispida (Schreber, 1775). >> Nycteris aethiopica Dobson, 1878 >>

3 222 M. J. LARGEN, D. KOCK and D. W. YALDEN D. Family Megadermatidae. 1. Lavia frons (E. Geoffroy, 1810) 2. Cardioderma cor (Peters, 1872) E. Family Rhinolophidae 1. Rhinolophus clivosus acrotis Heuglin, Rhinolophus clivosus?andersoni Thomas, Rhinolophus landeri dobsoni Thomas, Rhinolophus hipposideros minimus Heuglin, Rhinolophus simulator Andersen, Rhinolophus blasii andreinii Senna, Rhinolophus fumigatus Riippell, Rhinolophus hildebrandti Peters, 1878 F. Family Hipposideridae 1. Hipposideros ca/fer ca/fer (Sundevall, 1846) 2. Hipposideros ruber centralis Andersen, Hipposideros commersoni marungensis (Noack, 1887) 4. Hipposideros fuliginosus (T~mminck, 1853) 5. Hipposideros megalotis (Heuglin, 1861) 6. Triaenops persicus afer Peters, Asellia tridens tridens (E. Geoffroy, 1813) 8. Asellia patrizii De Beaux, 1931 G. Family Vespertilionidae. 1. Eptesicus somalicus somalicus (Thomas, 1901) 2. Eptesicus capensis (A. Smith, 1829) 3. Eptesicus guineensis (Bocage, 1889) 4. Pipistrellus nanus (Peters, 1852) 5. Pipistrellus kuhli fuscatus Thomas, Pipistrellus rusticus marrensis Thomas & Hinton, Pipistrellus rueppelli rueppelli (J. B. Fischer, 1829) 8. Mimetillus moloneyi (Thomas, 1891). 9. Glauconycteris variegata (Tomes, 1861). 10. Laephotis wintoni Thomas, Plecotus austriacus (J. B. Fischer, 1829) 12. Barbastella barbastellus leucomelas (Cretzschmar, 1830) 13. Miniopterus inflatus africanus Sanborn, Miniopterus schreibersi arenarius Heller, Nycticeius schlieffeni schlieffeni Peters, Nycticeius (Scotoecus) hindei (Thomas, 1901) 17. Nycticeius (Scotoecus) hirundo (De Winton, 1899) 18. Myotis (Selysius) tricolor (Temminck, 1832). 19. Myotis (Selysius) scotti Tl)omas, Myotis (Selysius) morrisi Hill, Myotis (Chrysopteron) welwitschi (Gray, 1866) 22. Scotophilus nigrita flavigaster (Heuglin, 1861) 23. Scotophilus leucogaster (Cretzschmar, 1830) 24. Kerivoula harrisoni harrisoni Thomas, Kerivoula eriophora (Heuglin, 1877) H. Family Molossidae 1. Otomops martiensseni martiensseni (Matschie, 1897) 2. Platymops setiger macmillani Thomas, Tadarida (Chaerephon) pumila pumila (Cretzschmar, 1830) 4. Tadarida (Chaerephon) nigeriae nigeriae (Thomas, 1913) 5. Tadarida (Chaerephon) bivittata (Heuglin, 1861) 6. Tadarida (Mops) midas midas (Sundevall, 1843) 7. Tadarida (Mops) condylura (A. Smith, 1833) 8. Tadarida (Mops) nanula (J. A. Allen, 1917). 9. Tadarida (Tadarida) ansorgei (Thomas, 1913) page 234» 234 >> 235 >> 235» 235 >> 236 >> 236 >> 237» 237 >> 237 >> 238 >> 239 >> 239 >> 239 >> 240 >> 240 >> 240 >>» » 241» 242» 242 >> )) >> 243 >> 243 >> 244» 244» 245 >> 245 >> 245 >> 245 >> 246 >> 246 >> 246 >> 246» 247 >> 247 >> 247 >> 248» 248 >> 248 >> 248» 249» 249 >> 250 >> 250 >> 250 >> 250 >> 250 >> 251 >> 252 >> 252 >> 252 >> 252 >> 253 >> 253

4 MAMMALS OF ETHIOPIA. I. CHIROPTERA Tadarida (Tadarida) aegyptiaca (E. Geoffroy, 1818) 11. Tadarida (Tadarida) ventralis (Heuglin, 1861) Tadarida (Mormopterus) acetabulosus natalensis (A. Smith, 1847) IV. Zoogeographical aspects of the bat fauna of Ethiopia. A. The Palaearctic element B. The Saharo-Sindian element. C. The African element V. The habitat and faunal composition at some major collecting sites VI. Gazeteer of collection localities Summary Riassunto References I. INTRODUCTION page 253» 253» 254 )) 254» 254» 255 >> 255, 257 )) 287» 293» 293 )) 294 The mammals of Ethiopia, in common with most other aspects of the fauna, are relatively poorly known. Little has been published on the subject in recent years, older reports are widely scattered throughout the zoological literature and the conclusions of past workers have often been limited by a lack of comparative material. However, a considerable amount of collecting has been done in Ethiopia during the past 10 years and the present work has been undertaken both to document this recent material and as an attempt to summarise the known distribution of Ethiopian species. It is hoped that this review of the literature omits few, if any, records and that any references which may have been overlooked will prove to be of minor importance. The collections of Ethiopian Chiroptera in the British Museum (Natural History), London; the Forschungsinstitut und Natur-Museum Senckenberg, Frankfurt; the Museum Alexander Koenig, Bonn; the Staatliches Museum fiir Naturkunde, Stuttgart; the J. W. Goethe-University, Frankfurt and the Natural History Museum, Addis Ababa have been thoroughly examined. It has not been possible to undertake a similar treatment of material in other collections and, inevitably, some specimens may have been omitted. If this is so, it is to be hoped that the present work will provoke a re-examination of such neglected material and the authors would welcome information concerning additional records. As far as possible, all collecting localities in Ethiopia have been gazetted and this record is supplemented with distribution maps for every species. It is hoped that such information will be of value to subsequent workers engaged tipon much-needed taxonomic revisions, who may be hampered by a limited knowledge of Ethiopia. In addition to localities within Ethiopia itself, it seems instructive to include certain collecting sites lying on or close to the Ethiopian border. Places which

5 224 M. J. LARGEN, D. KOCK and D. W. YALDEN have been treated in this way are French Somaliland (French Territory of Afars and Issas); Dolo and Jiffa Medir, Somalia; Elolo, Maddo Erelle Wells and Sancurar, Kenya; Gallabat and Kibbish Wells, Sudan. No attempt has been made to provide information for the determination of species. The standard work on this subject, «The mammals of Africa. An identification manual», MEESTER & SETZER (Edits) 1971, is so recently published that such repetition appears unjustified. However, in some cases, brief explanatory notes are provided to elucidate taxonomic uncertainties or recent changes in taxonomic concepts. Most of the recently acquired material, here reported for the first time, has been identified by Mr J. E. HILL of the British Museum (Natural History), whose assistance and advice is gratefully acknowledged. We are also indebted to Dr K. F. KooPMAN of the American Museum of Natural History, New York, for allowing us to include his records of Ethiopian Chiroptera in the museums of the United States and Europe and we thank Dr F. DIETERLEN, Staatliches Museum fur Naturkunde, Stuttgart, and Prof. E. VON LEHMANN, Museum Alexander Koenig, Bonn, for the loan of specimens. Prof. D. STARCK of the J. W. Goethe-University, Frankfurt has kindly allowed us to study the bats collected during his Ethiopia Expedition , which was undertaken with financial support from the Deutsche Forschungsgemeinschaft. A number of Ethiopian Chiroptera, recently acquired by the Museo Zoologico dell'universita di Firenze, were obtained by Dr M. L. AzzAROLI, Mr E. GRANCHI and Prof. B. LANZA during a visit to the Awash National Park (8-19.VI.1971) financed by the Consiglio Nazionale delle Ricerche (Centro di Studio per la Faunistica ed Ecologia Tropicali del C.N.R. Director: Prof. L. Pardi). A considerable debt is owed to the various individuals who have, in recent years, helped to build up the collections of Ethiopian bats; in particular, Dr J. S. AsH, who is responsible for most of the valuable series of specimens from Gambela, the Didessa River, Lake Abiata and Bahadu. The contributions of Dr R. AsHFORD, Messers M. BoLTON, R. 0. S. CLARKE, T. HAKANSSON, G. NIKOLAus, H. RuPP and Dr P. A. MORRIS are also acknowledged. Finally, thanks are due to Mrs S. E. LARGEN for her secretarial work and other assistance extending over a period of several years. The following abbreviations are used in this work: A.M.N.H. A.N.S.P. American Museum of Natural History, New York. = Academy of Natural Sciences, Philadelphia.

6 MAMMALS OF ETHIOPIA. I. CHIROPTERA 225 B.M. D.S.F. F.M.N.H. M.F. M.K.B. M.N.S. M.S.N.G. M.S.N.M. N.H.M.A.A. S.M.F. U.S.N.M. Z.M.B. British Museum (Natural History), London. Prof. D. Starck, University of Frankfurt, Ethiopia Expedition Field Museum of Natural History, Chicago. Museo Zoologico dell'universita di Firenze. Museum Alexander Koenig, Bonn. Staatliches Museum fiir Naturkunde, Stuttgart. Museo Civico di Storia Naturale «Giacomo Doria», Genova. Museo Civico di Storia Naturale, Milano. Natural History Museum, Addis Ababa. Senckenberg Museum, Frankfurt a.m. United States National Museum, Washington D.C. Zoologisches Museum der Humboldt Universitat, Berlin. II. SUBORDER MEGACHIROPTERA A. FAMILY PTEROPIDAE 1. Hypsignathus monstrosus H. Allen, 1861 Hypsignathus monstrosus H. ALLEN, 1861, Proc. Acad. nat. Sci. Philad.: 157. Gabon. Records. - ANDRAL et al., 1968: Manera Forest. Note. - A forest form extending from West Africa to Uganda and western Kenya; it is likely that the species is restricted, in Ethiopia, to areas of tropical deciduous forest in the south-western part of the country. 2. Epomophorus minor Dobson, 1880 Epomophorus minor DOBSON, 1880, Proc. zoo). Soc. Lond.: 715. Zanzibar. Records.- MoNTICELLI, 1887; ANDERSEN, 1912a; HARRISON, 1959: Farre.- MoNTICELLI, 1887: Let Marefia.- }ENTINK, 1888; ANDERSEN, 1912a; HARRISON, 1959; KocK, 1969: Shoa. - DE BEAux, 1922: Dolo (E. anurus). - DE BEAUX, 1914: Dolo (E. labiatus minor: correction of previous record). - BEADLES & INGERSOL, 1968: Errer valley, at bridge on Jigjiga road; Errer valley, 5 km E of bridge on Jigjiga road. - INGERSOL, 1968: Errer valley. - FuNAIOLI & LANZA, 1968: Dolo. - KocK, 1969: Let Marefia (labiatus: here corrected).

7 226 M. J. LARGEN, D. KOCK and D. W. YALDEN Additional material. - B.M.: Bahadu, (d', <;? ), E shore of Lake Abiata, (5 d' d', 3 <;? <;? ), X So Omar, ( <;? ), 29.XII km SW of Gidole, ( <;? ), 13.VIII N.H.M.A.A.: Bahadu, M (d', <;?), I.l971.- E shore of Lake Abiata, M (2 <;? <;? ), X S.M.F.: Bahadu, (8 d' d', 14 <;? <;? ), 3-7.V Koka, , 44231,44257 (d', 4 <;? <;?, 1 juv.), 5-18.III.l971; (d'), 2.IV U.S.N.M.: Bahadu, (3.5 ex.), XI.l971, 29.IV-3.V Koka, (25 ex.), XII.1971, 26.III-8.IV E shore of Lake Abiata, (8 ex.), X Note. -The genus Epomophorus is represented in Ethiopia by four forms. The smallest (minor) is a southern species which apparently reaches its northern limits in central Ethiopia; the rather larger E. labiatus is represented by the small nominate race, which is perhaps confined to northern Ethiopia and adjacent Sudan, and by the larger E. l. anurus which extends to the south and west. The largest species, E. gambianus, is, like Micropteropus pu.sillus, a West African element which reaches its eastern limits of distribution in Ethiopia. 3. Epomophorus labiatus labiatus (Temminck, 1837) Pteropus labiatus TEMMINCK, 1837, Mon. Mammalog. 2: 83, pl. 39. Sennar, Sudan (fide ANDERSEN 1912a). Pteropus schoensis RUPPELL, 1842, Mus. Senckenberg. 3 (2): 131. Shoa, Ethiopia. Epomophorus schovanus HEUGLIN, 1877, Reise in Nordost-Afrika 2: 18. (Emendation for schoensis Riippell). Epomophorus doriae MATSCHIE, 1899, Megachiroptera Berlin Mus.: 54. Bogos-Land, Ethiopia. Records. - RuPPELL, 1842; ANDERSEN, 1912a; KocK, 1969: Shoa (schoensis). - FrTZINGER, 1866 (schoensis); HEDGLIN, 1877 (schovanus): Bellegaz valley. - MoNTICELLI, 1887; SENNA, 1905 (doriae); ANDERSEN, 1912a (anurus); DE BEAUX, 1939 (anurusi; KocK, 1969; PuscHER, 1972 (anurus): Keren. - MATSCHIE, 1899: Bogos (doriae). - ANDERSEN, 1912a: Simien (anurus); Let Marefia. - HARRISON, 1959: Shoa; Let Marefia. - KocK, 1969: Simien; Ghinda; Zeriba Ambarab (the record from Let Marefia is here corrected to E. minor). Additional material. - B.M.: Gondar, D.S.F.: Keren, (7 d' d', 4 <;? <;? ), 26.II Note. - The distribution of E. labiatus in NE Africa is very poorly understood and is at present under review elsewhere (HILL, in litt.; KocK, in preparation). The occurence of E. l. anurus in the southern part

8 MAMMALS OF ETHIOPIA. I. CHIROPTERA 227 of Ethiopia seems established but records from further north, in nearly all cases based upon small samples, often of immature or female specimens, are here tentatively referred to E. l. labiatus. A much better understanding of the taxonomy of medium-sized Epomophorus is likely to be obtained when further specimens of anurus from its typical locality are available. 4. Epomophorus labiatus anurus Heuglin, 1864 Epomophorus anurus HEUGLIN, 1864, Nova Acta Acad. Caesar Leop. Care~!., Halle 31 (7): 12. Bongo, Bahr el Ghazal, S Sudan. Records. -ANDERSEN 1912a: Gofa. -KERSHAW, 1924: Bahr der Giorgis. -?ANDRAL et al., 1968: Manera Forest (limbatus, sic!). - HILL & MoRRIS, 1971: Azir River (mouth). - DoRST & PREVOST, 1972: W shore of Lake Abaya; 7 km from Ghinir. Additional material. - B.M.: Bulcha Forest, (d'), 13.XII. 1969; ( d' ), 22.XI.l970; (2 d' d', 4 <;? <;? ), 4-U.VI D.S.F.: Ambo, Am.II.165 ( <;.> ), 18.II M.S.N.G.: Jimma (KooPMAN, in litt.). N.H.M.A.A.: E shore of Lake Awasa, M (2 <;? <;? ), 1l.X Bulcha Forest, M. 558 ( <;? ), 13.XII.l969. S.M.F.: Arba Minch, (3 <;? <;? ). - E shore of Lake Awasa, (6 d'd', 6 <;? <;? ), 2.IX Epomophorus gambianus gambianus (Ogilby, 1835) Pteropus gambianus OGILBY, 1835, Proc. zoo!. Soc. Lond.: 100. Gambia. Records. -ANDERSEN, 1912a: Djiren; Djala. - HILL & MORRIS, 1971: Azir River (mouth) (E.? gambianus). - DoRST & PREVOST, 1972: 10 km from Agaro; Ghimbi. Additional material. - B.M.: Didessa River bridge, , ( d', 4 <;? <;? ), 11-2l.III.l Sidamo (Allison coil.). - Bulcha Forest, ( d', 4 <;? <;? ), 9-13.VI.l971. M.K.B.: Gambela, ( <;.> ), 27.VI.l972; ( <;? ), 2.VIII.l972. N.H.M.A.A.: Didessa River bridge, M ( d', <;? ), 27.VII Gambela, M ( d', <;.> ), 8-16.VIII S.M.F.: Didessa River bridge, (4 <;? <;? ), 4-9.VII Gambela, ( <;? ), 29.VII E shore of Lake Awasa, (4d'd'), 2.IX.1973.

9 228 M. J. LARGEN, D. KOCK and D. W. YALDEN U.S.N.M.: Didessa River bridge, (2 ex.), Gambela, (6 ex.), 27.VII.1972, 4-12.XII Bulcha Forest, (21 ex.), 27.II-8. III.l972, 1-ll.VII.1972, XI.l Micropteropus pusillus (Peters, 1868) Epomophorus pusillus PETERS, 1868, Mber. K. preuss. Akad. Wiss. Berlin for 1867: 870. Gambia. Records. - HILL & MoRRIS, 1971: Donkam River; Sirba. - DoRST & PREVOST, 1972: 10 km from Agaro; Didessa (River bridge). Additional material.- B.M.: Didessa River bridge, (2 d d, 2 ~ ~ ), 15.III-26.V.1970; ( 15 d d, 22 ~ ~ ), 11-2l.III.l970, V.1970; (4 dd, 17 S? ~ ), II.1971, 3-U.VII Gambela, (3 d d, 2 ~ ~ ), 27.XII.1969, 13.VI.l970; (15 dd, 21 ~ ~), l.i-17.ii.1970, 13.VI.l971; (3 ~ ~ ), III.l970; (14 dd, 7 ~ ~ ). - Bulcha Forest, ( ~ ), 6.III.l972. M.K.B.: Gambela, (3 d d, 6 ~ ~, 1? sex), VI.l972, 3.VII.l972, 28.VII.l972. N.H.M.A.A.: Gambela, M. 552 ( ~ ), ; M. 555, 559 (d, ~ ), 6-10.!1.1970; M ( d, 'i' ), VI.l970. U.S.N.M.: Didessa River bridge, (13 ex.), 27.I-9.II.1972, Bulcha Forest, (1 ex.), 7.VII Eidolon helvum (Kerr, 1792) Vespertilio vampyrus helvus KERR, 1792, Linnaeus' Animal Kingdom 17: 91. No locality, restricted to Senegal by ANDERSEN (1907 a). Records. - HAYMAN & HILL, 1971: Ethiopia (a single specimen from Dech Island, Lake Tana: B.M , Cheeseman coli.). Additional material. - M.K.B.: Gambela, (2 d d), N.H.M.A.A.: Gambela, M. 687 (d), 12.XII U.S.N.M.: Gambela, (1 ex.), 13.XII.l Rousettus aegyptiacus (E. Geoffroy, 1810) Pteropus egyptiacus E. GEOFFROY, 1810, Annis Mus. natn. Hist. nat. Paris IS: 96 (misprint); corrected to aegyptiacus in: Descript. de I'Egypte, 2: 134, Great Pyramid, Gizeh, Egypt. Pteropus leachii A. SMITH, 1829, Zool. J. 4: 433. Cape Town, South Africa. Rousettus arabicus ANDERSON & DE WINTON, 1902, Zool. Egypt, Mamm.: 86, 83. Lahej, South Yemen. Records. -DoBSON, 1878; ANDERSON & DE WINTON, 1902: Abyssinia. - SoRDELLI, 1902; SENNA, 1905: Addi Sciaddi. - HAYMAN

10 MAMMALS OF ETHIOPIA. I. CHIROPTERA 229 & HILL, 1971: Harar (arabicus). - DoRST & PREVOST, 1972: W shore of Lake Abaya; Koffole; 10 km from Agaro; Ghimbi. Additional material: Rousettus aegyptiacus leachi B.M.: Didessa River bridge, , (4 cfcf, 13 <;? <;? ), III M.K.B.: Gambela, ( cf, <;? ), 6.VIII.l972; ( <;? ), 8.VIII Gololcha, ( cf ), 15.IX N.H.M.A.A.: Didessa River bridge, M ( cf, <;? ), III S.M.F.: Didessa River bridge, (2 <;? <;? ), 14.III.l Gambela, ( <;? ), 6.VIII Jimma, ( cf ), 4.VIII Gojeb River, (cf), 17.VIII Rousettus aegyptiacus ssp. B.M.: E shore of Lake Abiata, ( <;? ), 10.X Bulcha Forest, ( <;? ), 2.XII.1969; (2 cf cf ), 11.XII Arba Minch, ( <;? ). N.H.M.A.A.: Koka, M. 560 ( cf ), 12.VII E shore of Lake Abiata, M. 542 ( <;? ), 27.X Bulcha Forest, M ( cf, <;? ), 13.XII So Omar, M (3 <;? <;? ), XII U.S.N.M.: Bulcha Forest, (1 ex.), 7.III Note. - This species is represented in Ethiopia by two races: arabicus, of small size but with heavy teeth, in the east (Harar) and leachi, with larger dimensions but weaker dentition, in the west. The geographical limits of the two forms have yet to be determined and the subject is at present under review (HILL, in litt.). 9. Rousettus angolensis (Bocage, 1898) Cynonycteris angolensis BocAGE, 1898, J. Sci. math. phys. nat., Lis boa (2) 5: 133, 138. Pungo Andongo, Cahata, Quibula, Angola. Rousettus angolensis ruwenzorii E!SENTRAUT, 1965, Bonn. Zoo!. Beitr. 16: 3. Ruwenzori East (Mubuku valley), Uganda. Material. - B.M.: Didessa River bridge, ( <;? ), 13.III.l970; ( cf ), 27.VII.1970; (2 cf cf, <;? ), 5-12.VII Nur Mohammed, ( cf ), 28.III N.H.M.A.A.: Didessa River bridge, M. 561 ( cf), 16.II U.S.N.M.: Didessa River bridge, (2 ex.), 28.!.1972,

11 230 M. J. LARGEN, D. KOCK and D. W. YALDEN Note. - This is the first record of the species in Ethiopia; the specimens agree closely with the East African race R. a. ruwenzorii, (HILL, in litt. ). 10. Rousettus lanosus kempi Thomas, 1909 Rousettus kempi THOMAS, 1909, Ann. Mag. nat. Hist. (8) 4: 543. Kirui, Mt. E1gon, Kenya. Records. - ANDERSEN, 1907a, 1912a: Shoa. Additional material. - M.B.: Beletta Forest, (2 9 9 ), 8.II.l972. III. SuBORDER MICROCHIROPTERA A. FAMILY RHINOPOMATIDAE 1. Rhinopoma hardwickei sennaariense Fitzinger, 1866 Rhinopoma sennaariense FrTZINGER, 1866, Sber. Akad. Wiss. Wien 54 (1): 547. Sennaar and Fazoglo ( = Fazughli), Sudan. Records. -HILL & MoRRIS, 1971: NW slope of Mount Fantalle; N bank of Awash River; Metahara. -DE BLASE et al., 1973: Djibouti. Additional mataial.- M.F.: Metahara, 6537 (d'), (3 9 9 ), (3 9 9 ), (2 9 9 ), 8-19.IV (3 9 ), (2 9 ), 8-19.IV N.H.M.A.A.: W slope of Mount Fantalle, M. 18 ( d' ), 28.IX Metahara, M (d', 9 ), 10.IV.1971; M. 303 ( 9 ), 2l.XII Awash Falls, M. 415 ( 9 ), 25.IX S.M.F.: W of Metahara, (2 d' d', 9 ), 1l.IX Rhinopoma muscatellum Thomas, 1903 Rhinopoma muscatellum THOMAS, 1903, Ann. Mag. nat. Hist. (7) 11: 493. Wadi Bani Ruha, Muscat, Arabia. Records. -? SENNA, 1905: Assab. - KocK, 1969 (R. hardwickei macinnesi): Assab (l.viii.1906). Note. -DE BLASE et al. (1973) have demonstrated that the smallest Asian representatives of Rhinopoma should be treated as a separate species living sympatrically with the larger hardwickei, to which two very small examples from Assab are now assigned. SENNA's ( 1905) specimens have not been re-examined but are presumed to belong here.

12 MAMMALS OF ETHIOPIA. I. CHIROPTERA 231 B. FAMILY EMBALLONURIDAE 1. Taphozous perforatus E. Geoffroy, 1818 Taphozous perforatus E. GEOFFROY, 1818, Descript. de l'egypte 2: 126. Kom Ombo, Egypt. Taphozous perforatus haedinus THOMAS, 1915, J. Bombay nat. Hist. Soc. 24: 62. Chanter Falls, N Guaso Nyiro, Kenya. Records. - DE BEAUX, 1939: Agordat. - HARRISON, 1962: Kibbish Wells (T. p. haedinus). - BEADLES & lngersol, 1968: 15 km SE of the Jigjiga road (T. sudani). - lngersol, 1968: Dacata valley. - KocK, 1969: Agordat cr. p. perforatus ). - HILL & MORRIS' 1971: 10 km W of Mabil; N bank of Awash River (T. p. haedinus). Additional material: T aphozous perforatus haedinus B.M.: Beleka, ( d' ), 14.IV W of Metahara, (d'), 29.XII W shore of Lake Langano, ( ' ), 14.III. 1969; ( ' ), 8.V E shore of Lake Shalla, (d'), 22.III.l969. N.H.M.A.A.: 10 km W of Mabil, M ( d', ' ), 19.IX W shore of Lake Langano, M. 7-8 (2 ' ' ), 14.III.1969; M. 36 ( ' ), 8.V E shore of Lake Shalla, M (2 d'd'), 22.III Note. - In Ethiopia this species is possibly represented only by the dark form haedinus, which is replaced to the north-west by paler perforatus and to the south-west by white-winged sudani. However, the exact distribution of these three races is not clearly established and it is to be expected that intermediate populations will be discovered. 2. Taphozous nudiventris Cretzschmar, 1830 Taphozous nudiventris CRETZSCHMAR, 1830, in: Senck. naturf. Ges.: Atlas Reise nord!. Afrika E. Riippell, Sliugeth.: 70, fig. 27 b. Gizeh Pyramids, Egypt. Taphozous perforatus var. assabensis MONTICELLI, 1885, Ann. Accad. aspir. Nat. Napoli 1: 78, pl. 7. Assab, Ethiopia. Records. - MoNTICELLI, 1885: Assab. - SENNA, 1905: Buia ( = Assab). - KocK, 1969: Kibbish Wells. Additional material. - S.M.F.: Abyssinia, 1313 (?sex), 1827, Riippell coll. 3. T aphozous mauritianus E. Geoffroy, 1818 Taphozous mauritianus E. GEOFFROY, 1818, Descript. de I'Egypte 2: 127. Mauritius. Material.- B.M.: Gambela, (d'), 1.! M.K.B.: Gambela, (d'), 3.VII.1972.

13 232 M. J. LARGEN, D. KOCK and D. W. YALDEN Note. - This appears to be the first record of the species in Ethiopia, although previously known from Somalia and central Sudan. Emballonura afra PETERS, Mozambique. 4. Coleura afra (Peters, 1852) 1852, Reise nach Mossambique, Siiugeth.: 51, pl Tette, Records. - KocK, 1969: Kibbish Wells. - HAYMAN & HILL, 1971: Ethiopia (apparently based upon a specimen from Dobleh in the British Museum (Natural History)), Additional material. - B.M.: Lake Hertale, ( cf ), 19.II S.M.F.: E shore of Lake Awasa, ( <:i? ), 2.IX W shore of Lake Chamo, ( <:i? ), 4.VIII.l973. c. FAMILY NYCTERIDAE 1. Nycteris thebaica E. Geoffroy, 1813 Nycteris thebaicus E. GEOFFROY, 1813, Annis Mus. natn. Hist. nat. Paris 20: 20. Egypt. Nycteris labia/a HEUGLIN, 1861, Nova Acta Acad. Caesar Leop. Carol., Halle 29 (8): 4, 5. Near Keren, Ethiopia. Nycteris revoilii RoBIN, 1881, Bull. Soc. philom. Paris (7) 5: 90. Somalia north of 10 N. Petalia damarensis brockmani ANDERSEN, 1912, Ann. Mag. nat. Hist. (8) 10: 548, line 10. Upper Sheikh, N Somalia. Petalia damarensis media ANDERSEN, 1912, Ann. Mag. nat. Hist. (8) 10: 548, line 28. Harar, Ethiopia. Records. - HEDGLIN, 1861a, 1877: near Keren (N. labiata). - PETERS, 1871; SENNA, 1905; DE BEAux, 1923: Keren. - MoNTICELLI, 1887; DE BEAUX, 1923: Nora Island. - GIGLIOLI, 1888: Assab. - }ENTINK, 1888: Azam River (N. labiata). - SENNA, 1905; DE BEAUX 1923, 1930: Agordat (labiata fide KooPMAN, in litt.). - SENNA, 1905: Godolefassi; Adi Guhebo; Adi Ugri; Adi Caie (all labiata fide KooPMAN, in litt.). - ANDERSEN, 1912b; DE BEAUX, 1923: Harar (N. damarensis media). - DE BEAUX, 1923: Setit (River); Rorum plains (the latter labiata fide KooPMAN, in litt.). - DE BEAUX, 1923: Saganeiti (N. damarensis brockmani). - KERSHAW, 1924: Baht der Giorgis (revoili). - RADFORD, 1954: Asmara (N. damarensis). - lngersol, 1968: 4 km S of Dire Dawa (N. thebaica media fide KooPMAN, in litt.). - KocK, 1969: Lake Langano.- HILL & MoRRIS, 1971: Keren; Mabil; 10 km W of Mabil (all labiata); Gondar; Ghibbey valley (as N. t. thebaica but both labiata fide KooPMAN, in litt.); Gallabat (N. t. thebaica); NE of Lake Chamo.

14 MAMMALS OF ETHIOPIA. I. CHIROPTERA 233 Additional material: Nycteris thebaica labiata A.N.S.P.: Gura (KOOPMAN, in litt.). B.M.: E shore of Lake Abiata, (juv.), 14.III.l W shore of Lake Langano, (juv. d'), 13.I'II.1969; (d'), 8.V Abella, ( ~ ), 25.V N. shore of Lake Abaya, ( ~ ), 25.V D.S.F.: Ambo, Am.II.52 ( d' ), F.M.N.H.: 6 miles W of Asmara.- Sembel (both KooPMAN, in litt.). M.S.N.G.: Dorfu (KooPMAN, in litt.). M.S.N.M.: Tarat (KooPMAN, in litt.). N.H.M.A.A.: W shore of Lake Langano, M. 2-3 (2 ~ ~ ), 13.III.l969; M. 37 (d'), 8.V N shore of Lake Abaya, M. 40 (d'), 25.V S.M.F.: Abyssinia, 1322, Riippell coli. Nycteris thebaica brockmani B.M. Metahara, (1 ex.), 29.XII Nycteris thebaica ssp. B.M.: Didessa River bridge, ( ~ ), 13.Il Note. - The subspecies distribution of N. thebaica is poorly understood and apparently complex. It seems that labiata (of which revoili is a synonym) is widely distributed in western and northern Ethiopia, while thebaica may be found in the vicinity of the Sudanese border. In the eastern part of the country labiata is apparently replaced by a larger form, brockmani (with media as a possible synonym). This confusion results from the fact that new names have often been based upon a very small number of specimens and upon characters, such as colouration and the position of a considerably reduced lower premolar, which are now known to be unreliable. The problem it at present under review (KOOPMAN, in litt.). 2. Nycteris hispida hispida (Schreber, 1775) Vespertilio hispidus ScnREBER, 1775, Sliugethiere 1: 169, 188, pl. 56. Senegal. Material. - B.M.: Bahadu, ( ~ ), Filwoha, (d'), 26.XII Awash Falls, (d'), 25.XII Gambela, (d'), Bulcha Forest, (d'), 11.XII km N of Lake Rudolf, ( d' ), 8.VIH M.K.B.: Gambela, 73.69, (3 d' d', ~ ), 1972; 73.70, (3 d' d' ), 26.VI-28.VII.1972.

15 234 M. J. LARGEN, D. KOCK and D. W. YALDEN N.H.M.A.A.: Pokwo, M ( d', ~ ), 23.VIII S.M.F.: W shore of Lake Abaya, ( d' ), 29.VII shore of Lake Chamo, ( d' ), 4.VIII.l973. U.S.N.M.: Filwoha, (1 ex.), 25.X Note. - It seems that this species has not previously been recorded from Ethiopia, although widely distributed in sub-saharan Africa. The two specimens in the Senckenberg Museum, although from similar habitats separated by only 30 km, are markedly different in size. Some charactensttc measurements are (Lake Abaya: Lake Chamo) - forearm 39.1 : 43.1; ear 20.0: 24.5; condylocanine length 13.8: 15.3; upper toothrow 4.9 : 6.0. This size difference perhaps supports KooPMAN's opinion (in HAYMAN & HILL, 1971) that in parts of eastern Africa a smaller form (hispida) appears to co-exist with a larger one (aurita). 3. Nycteris aethiopica Dobson, 1878 Nycteris aethiopica DoBSON, 1878, Cat. Chiropt. Br. Mus.: 165. Kordofan, Sudan. Records. - RoDE, 1933: Gondar. -DE BEAUX, 1943: Gondaraba (N. aethiopica luteola ). Additional material. - A.N.S.P.: 30 miles W of Alghe. B.M.: Robi. - Didessa River bridge, ( ~ ), 9.VII Gambela, ( ~ ), 13.II M.K.B.: Gambela, ( d' ), 20.XII Note. - It seems very probable that this form should be regarded only as a subspecies of N. macrotis Dobson, 1876 (type locality: Sierra Leone), unless the supposed specific differences can be demonstrated in sympatric populations (see KocK, 1969). D. FAMILY MEGADERMATIDAE W 1. Lavia frons (E. Geoffroy, 1810) Megaderma frons E. GEOFFROY, 1810, Annis Mus. natn. Hist. nat. Paris IS: 192. Senegal. Records. - DoBSON, 1878: Abyssinia. - MAGRETTI, 1884; SENNA, 1905; DE BEAUX, 1923: Barca (River). - THOMAS, 1898; DE BEAUX, 1923: between Badditu and Dime. - DE BEAUX, 1923: Agordat; Setit (River). - DE BEAUX, 1943: Sagan River; Murle; Elolo. - DoRST & PREVOST, 1972: W shore of Lake Abaya. Additional material. - A.N.S.P.: Sassabone. - A.M.N.H.: Lake Abaya. NW shore of Lake Stephanie.

16 MAMMALS OF ETHIOPIA. I. CHIROPTERA 235 B.M.: Abyssinia, , Zaphiro coli. - Gambela, ( S? ), 28.XII.1969; (2 c:f c:f), 9.II.l970, 16.II Dobleh. - Near Mega. M.K.B.: Gambela, (?sex), 12.IV.1972; 73.37, ( c:f, 2 S? S? ), VII M.S.N.M.: Odar. N.H.M.A.A.: Murle, M. 226 (1 ex.), 28.VI km W of Hoddo, M. 227 (1 ex.), 26.VI.l969. S.M.F.: Arba Minch, ( c:f, S? ), 25.IX U.S.N.M.: Gambela, (c:f), 12.VI.l970; (2 c:fc:f), VIII.l970; (c:f, 2 S? S? ), 25.VII-2.VIII.l971; (11 ex), 12.VI.l970, 20-2l.VIII.1970, 25.VII-2.VIII.l971, 26.VII.1972, 4-13.XII Cardioderma cor (Peters, 1872) Megaderma cor PETERS, 1872, Mber. K. preuss. Akad. Wiss. Berlin: 194. Abyssinia. Records. - PETERS, 1872: Abyssinia. - THOMAS, 1902: Dhar Ala, Danakil. - THOMAS, 1904: Gerlogobi. - HoLLISTER, 1918: Ogaden. - ]OBLING, 1939: Solomona Habesh. - HAYMAN & HILL, 1971: just N of Lake Stephanie. - HILL & MoRRIS, 1971: Awash National Park. - DoRST & PREVOST, 1972: W shore of Lake Abaya; 7 km from Ghinir. Additional material. - A.N.S.P.: Near Amar Cocche. B.M.: Wiah, (3 S? S? ). - Jebel Geddem, ( c:f, 2 S? S? ), 8.XII Metahara, (5 c:f c:f ), 21.XII.1970; ( S? ), 25.XII M.P.: Girghiret.- Hop Saloa mine.- Metahara, (7 c:f c:f), 8-19.IV M.K.B.: Auramalka, (?sex), N.H.M.A.A.: Metahara, M. 312, 314 (2 c:f c:f ), 21.XII S.M.F.: Arba Minch, (c:f), 28.VII U.S.N.M.: Bahadu, ( c:f, 2 S? S? ), l.v.1970; (2 c:f c:f, 2 S? S? ), IV.1972; (7 ex.), l.v.1971, IV E. FAMILY RHINOLOPHIDAE 1. Rhinolophus clivosus acrotis Heuglin, 1861 Rhinolophus acrotis HEUGLIN, 1861, Nova Acta Acad. Caesar Leop. Carol., Halle 29 (8): 4, 10. Near Keren, Ethiopia. Records. - HEUGLIN, 1861a: near Keren. - HEUGLIN, 1877: Bogos. - SENNA, 1905 (R. antinorii): Coatit; Assab. - ANDERSEN,

17 236 M. J. LARGEN. D. KOCK and D. W. YALDEN 1906: Keren; Assab; Harar. -DE BEAUX, 1931: Assab; Serae; Barentu; Harar; Gold mine of Uallega. - BEADLES & lngersol, 1968; lngersol, 1968: Alemaya. -HILL & MoRRIS, 1971: Awash National Park. - DoRST & HILL, 1972: Harar region. - DoRST & PREVOST, 1972: Dinshu; near Hirna. Additional material. - B.M.: Western Abyssinia, Gondar, Dangila. - Gara Hakim, (2 Cj) '?, 1 newborn), --.IV Didessa River bridge, ( cj, 5 '? '? ), V.1970; ( 11 cj cj, 15 '? '? ), 24.VII.1970, II. 1971, 3-9.VII.1971, 28.I-3.II Ueb River, Dinshu, (2 cfcf, S? ), 8-9.! F.M.N.H.: Sembel miles NE of Gondar (both KooPMAN, in!itt.). N.H.M.A.A.: Didessa River bridge, M ( cj, 3 S? S? ), V Note. - The forearm lengths and horseshoe widths recorded by SENNA ( 1905) for his specimens of R. antinorii ( = fumigatus ), indicate that they are misidentified R. c. acrotis. 2. Rhinolophus clivosus?andersoni Thomas, 1904 Rhinolophus andersoni THOMAS, 1904, Ann. Mag. nat. Hist. (7) 14: 156. Eastern Desert, in the vicinity of Wadi Alagi, 22'N 35 E, Sudan. Material. - B.M.: Gambela, (1 ex.), 7.II Note. - This specimen agrees in most respects with R. c. andersoni, (HILL, in litt.). 3. Rhinolophus landeri dobsoni Thomas, 1904 Rhinolophus dobsoni THOMAS, 1904, Ann. Mag. nat. Hist. (7) 14: 156. Kocdofan, Sudan. Records. - THOMAS, 1897: Sancurrar (Hipposideros euryale). ANDERSEN, 1906: Sancurrar (R. lobatus). - RoDE, 1933: Gondar. HILL & MoRRIS, 1971; Azir River (mouth). - DoRST & PREVOST, 1972: Neghelli. Additional material. - B.M.: Gallabat, ,10, Barye, W Ethiopia. - Koka, ( Cj) ), 20.XII Gambela, (cf), 9.II.1970; , 2212 (2 S? S? ), 2.!.1970, 11.II.1970; (5 cj cj, 3 Cj) Cj) ), VI.l So Omar, (4 cfcf, S? ), 26.XII Bulcha Forest, (cf), 16.XII.1969; (cf), 24.XI.1970.

18 MAMMALS OF ETHIOPIA. I. CHIROPTERA 237 N.H.M.A.A.: Gambela, M (2 9 Cj! ), , 8.II Bulcha Forest, M. 448 (d'), 17.XII Note. - It remains to be determined whether dobsoni is actually distinguishable from R. landeri lobatus Peters, 1852 of eastern and southern Africa (type locality: Sena, Mozambique). 4. Rhinolophus hipposideros minimus Heuglin, 1861 Rhinolophus minimus HEUGLIN, 1861, Nova Acta Acad. Caesar Leop. Carol., Halle 29 (8): 4, 6. Keren, Ethiopia. Records. - HEUGLIN 1861a, 1877: Keren. - HILL & MORRIS, 1971: Lake Baa-sa-ka. Additional material. - B.M.: Between Garibaldi Pass and Awash National Park, ( Cj! ), 19.XII Rhinolophus simulator Andersen, 1904 Rhinolophus simulator ANDERSEN, 1904, Ann. Mag. nat. Hist. (7) 14: 384. Mazoe, Mashonaland, Rhodesia. Records. - HILL & MoRRIS, 1971: 3 miles S of Goba. Additional material. - B.M.: Koka, ( Cj! ), 6.III E shore of Lake Abiata, (d'), 26.X So Omar, (d', 4 Cj! Cj! ), --.III.1971; (4 d'd', 3 Cj! Cj! ), XII N.H.M.A.A.: So Omar, M (2 ex.), --.III.l971. S.M.F.: Sof Omar, (2 9 9 ), --.III.l971. Note. - Specimens recorded from Ethiopia are perhaps referable to a small, undescribed montane subspecies (HILL & MoRRIS, 1971 ). In neighbouring territories, the nearest localities from which the species has been recorded are in southern Sudan. 6. Rhinolophus blasii andreinii Senna, 1905 Rhinolophus andreinii SENNA, 1905, Archo zool. ital. 2: 256, pl. 16, f. 1; pl. 18, f Adi Ugri, Ethiopia. Records. -SENNA, 1905: Adi Ugri.- ANDERSEN, 1906: Harar. DoRST & HILL, 1972: So Omar. Additional material. - A.M.N.H.: Harar (KooPMAN, in litt.). 17

19 238 M. J. LARGEN, D. KOCK and D. W. YALDEN B.M.: Koka, (2 ~ ~ ), 8-12.III.l Sidamo (Province), 140 miles S of Addis Ababa, So Omar, ( d' ), 26.XII Z.M.B.: Dire Dawa (KooPMAN, in litt.). 7. Rhinolophus fumigatus Riippell, 1842 Rhinolophus fumigatus RUPPELL, 1842, Mus. Senckenberg. 3: 132. Shoa, Ethiopia (Type: S.M.F. 4372). Rhinolophus macrocephalus HEDGLIN, 1877, Reise in Nordost-Afrika, 2: ~2. Adowa, Ethiopia. Rhinolophus antinorii DoBSON, 1885, Annali Mus. civ. Stor. nat. Giacomo Doria (2) 2: 16. Daimbi, Shoa, Ethiopia. Rhinolophus fumigatus exsul ANDERSEN, 1905, Ann. Mag. nat. Hist. (7) 15: 74. Kitui, Kenya. Records. - Ri.iPPELL, 1842: Shoa. - HEUGLIN, 1877: Adowa. - DOBSON, 1885: Daimbi.- THOMAS, 1898: Dass Wells. -SENNA, 1905 (R. aethiops): Adi Ugri; Godolefassi; Debaroa; near Adi Caie; Zeban Zighib.- DE BEAUX, 1923, 1925: Gondar.- DE BEAux, 1924: Asmara. - KERSHAW, 1924: Bahr der Giorgis. - HILL & MoRRIS, 1971: Fincha River (mouth); 30 km SE of Portuguese Bridge. - DoRST & HILL, 1972; DoRST & PREVOST, 1972: 10 km from Ghinir. Additional material. - B.M.: Gondar, Dangila. - Didessa River bridge, (juv.), 18.V.1970; ( d', 4 ~ ~ ), 3.VII.1971, Gambela (R. f. exsul), ( ~ ), 13.II E shore of Lake Abiata, ( ~ ), 10.X W shore of Lake Langano, ( d', 2 ~ ~ ), 9.III Sof Omar, (8 d' d', ~ ), XII E of Mega, (1 ex.), 28.II M.K.B.: Gambela (R. f. exsul), ( ~ ), M.S.N.M.: Torat. - Gondar (both KooPMAN, in litt.). N.H.M.A.A.: W shore of Lake Langano, M (2 d' d', ~ ), 9.III.l So Omar, M. 593 ( d' ), 28.XII S.M.F.: Shoa, (?sex), Gambela (R. f. exsul), ( d' ), 6.XI U.S.N.M.: Didessa River bridge, (3 ex.), Note. - Until very recently it was considered that R. f. fumigatus was the only representative of this species in Ethiopia, but the present recognition of specimens from Gambela as exsul suggests that a re examination of material from other localities in southern and western Ethiopia would be desirable. It is not yet clear whether R. aethiops Peters, 1868 should be regarded as a larger race of fumigatus or as a separate species. However, in the case of SENNA's (1905) records of aethiops from Ethiopia, a

20 MAMMALS OF ETHIOPIA. I. CHIROPTERA 239 straightforward misidentification of R. f. fumigatus seems very likely, particularly since his R. antinorii ( = fumigatus) prove to belong to the smaller R. clivosus. 8. Rhinolophus hildebrandti Peters, 1878 Rhinolophus hildebrandtii PETERS, 1878, Mber. K. preuss. Akad. Wiss. Berlin: 195. Ndi, Taita Hills, Kenya. Records. - DoRST & PREVOST, 1962: Koffole. Additional material. - B.M.: Melkay Manna, (2 ~ ~ ), 22.III.l972. Note. - Although only recently recorded from Ethiopia, this species is widely distributed in the eastern half of Africa, from Somalia and southern Sudan to the Transvaal. F. FAMILY HIPPOSIDERIDAE 1. Hipposideros calfer calfer (Sundevall, 1846) Rhinolophus caffer SUNDEVALL, 1846, Ofvers. K. svenska VetenskAkad. Forb. 3 (4): 118. Durban, Natal, South Africa. Phyllorrhina bicornis HEUGLIN, 1861, Nova Acta Acad. Caesar Leop. Carol., Halle 29 (8): 4, 7. Near Keren, Ethiopia. Records. - HEDGLIN, 1861a, 1877: near Keren. - SoRDELLI, 1902; DE BEAUX, 1930: Agordat. - SENNA, 1905: Keren; Agordat. - ANDERSEN, 1907b: Monkullo; Ghinda; Agordat; Harar. - HILL & MoRRIS, 1971: 10 km W of Mabil. Additional material. - B.M.: Tserea Wukro, ( d' ), 9.V Gallabat, , 20, Filwoha, ( ~ ), 26.XII Metahara, ( d' ), 25.XII Awash Falls, (2 ~ ~ ), 25.XII Woliso. - Gambela, (d'), 27.XII. 1969; (d'), 26.VI.l Adamatuli. - Sof Omar, (d'), 28.XII D.S.F.: Errer valley at Harar, HA.VIII (3 ~ ~ ), 2.IV M.F.: Cheren. M.K.B.: Gambela, ( d' ), N.H.M.A.A.: Awash Falls, M. 334 ( d'), 27.XII S.M.F.: Arba Minch, ( 14 d' d' ), 28.VII-9.VIII.l973. Z.M.B.: Dire Dawa. Note. - For the present, H. c. calfer is considered to be the only representative of the species in Ethiopia. It may be noted, however, that

21 240 M. J. LARGEN, D. KOCK and D. W. YALDEN the three female specimens from near Harar (D.S.F.) are rather similar to Sudanese examples of H. c. tephrus Cabrera, 1906 (type locality: Mogador, Marocco). 2. Hipposideros ruber centralis Andersen, 1906 Hipposideros catfer centralis ANDERSEN, 1906, Ann. Mag. nat. Hist. (7) 17: 275, 277. Entebbe, Uganda. Records. - HILL & MoRRIS, 1971: Fincha River (mouth); 10 km W of Mabil; Didessa River (mouth); SW corner of Lake Abaya. Additional material. - B.M.: Didessa River bridge, ( ~ ), 18.III.l970; ( d' ), 9.VII Gambela, ( ~ ), 28.XII.1969; (d'), 14.II.l970; (d'), 26.VII M.K.B.: Gambela, (? sex), 1972; ( d', ~ ), VI.l972. N.H.M.A.A.: 10 km W of Mabil, M. 404 ( ~ ), 19.IX Didessa River bridge, M.447 ( ~ ), 20.V Arba Mondi, M.29 (1 ex.), -.VIII.l966. Note. - Where this species is sympatric with H. caffer it is always distinctly larger, although other external characters may seem identical. HAYMAN & HILL ( 1971) characterise ruber as having a forearm length greater than 48 mm while that of caffer is less than 48 mm. In Ethiopia, however, the forearm in adult ruber exceeds 53 while that of caffer is never more than Hipposideros commersoni marungensis (Noack, 1887) Phyllorhina commersonii var. marungensis!'\oack, 1887, Zoo!. Jb. (Syst.) 2: 272, pl. 10, f Qua Mpala, Marungu, Zaire. Records. - THOMAS, 1895: Upper Ganale (river). - DE BEAUX, 1922, 1924: Dolo.- HAYMAN, 1967; HAYMAN & HILL, 1971: Southern Ethiopia. Additional material. - N.H.M.A.A.: Awash National Park, M.24 (1 ex.). - Koka, M.717 ( d' ), 6.III.l Hipposideros fuliginosus (Temminck, 1853) Phyllorrhina fuliginosa TEMMI"CK, 1853, Esq. Zoo!. Cote de Guine: 77. Guinea coast. Material. - S.M.F.: Gaba River bridge, (3 ~ ~ ), 24. VIII.1973.

22 MAMMALS OF ETHIOPIA. I. CHIROPTERA 241 Note. - This is the first recorded occurrence of H. fuliginosus in Ethiopia and the only collection of this forest species to be obtained east of the Ituri region, Zaire. 5. Hipposideros megalotis (Heuglin, 1861) Phyllorrhina megalotis HEDGLIN, 1861, Nova Acta Acad. Caesar Leop. Carol., Halle 29 (8): 4, 8. Bogos Land, Ethiopia. Records. - HEUGLIN, 1861a; DoBSON, 1878: Bogos. - HEUGLIN, 1877: near Keren. - SENNA, 1905: Keren. - lngersol, 1968: Alemaya; Era Gota. - HAYMAN, 1960, 1967; HAYMAN & HILL, 1971: Sidamo (Province), 140 miles S of Addis Ababa. Note. - The species is known only from Ethiopia and a few localities in central Kenya. 6. Triaenops persicus afer Peters, 1877 Triaenops afer PETERS, 1877, Mber. K. preuss. Akad. Wiss. Berlin for 1876: 913, f. 2. Mombasa, Kenya. Records.- THOMAS 1897; FuNAIOLI & LANZA, 1968: Maddo Erelle Wells. - DE WINTON, 1898; FuNAIOLI & LANZA, 1968: Jiffa Medir. - HILL & MoRRIS, 1971: Didessa River (mouth); Awash National Park. Additional material. - B.M.: Bahadu, (3 ~ ~ ), , 8.V.1971, 23.XI Beleka, ( cf, ~ ), 14.IV Koka, (1 ex.), 19.XII So Omar, (11 ~ ~ ), XII Boran-Galla country (Borana region), , 5. N.H.M.A.A.: Between Addis Ababa and Lake Langano, M.19 (1 ex.), -.X So Omar, M.592 ( ~ ), 28.XII Asellia tridens tridens (E. Geoffroy, 1813) Rhinolophus tridens E. GEOFFROY, 1813, Annis Mus. natn. Hist. nat., Paris 20: 260, pl. 5. Egypt, restricted to Thebes, Luxor. Records. - SoRDELLI, 1902; SENNA, 1905: Agordat. -DE BEAUX, 1922, 1924: Dolo. -DE BEAux, 1931, 1934: Dolo (A. t. italosomalica). -HILL & MoRRIS, 1971: Awash National Park. Additional material. - B.M.: Metahara, (2 cf cf, ~ ), XII.1970.

23 242 M. J. LARGEN, D. KOCK and D. W. YALDEN 8. Asellia patrizii De Beaux, Asellia patrizii DE BEAUX, 1931, Annali Mus. civ. Stor. nat. Giacomo Doria 55: 1f6. Gaarre, Ethiopia. Records.- DE BEAUX, 1931: Gaarre; Assab.- HARRISON, 1965; HAYMAN & HILL, 1971: Entedebir Island; Assab. -HAYMAN & HILL, 1971; HILL & MoRRIS, 1971: Awash National Park. Additional material. - B.M.: Nocra Island, (3 9 9 ), 27.XII Wongabo. - Filwoha, (2 9 9 ), 23.XII W of Metahara, (7 cf cf, ), XII M.F.: Metahara, (3 cf cf ), 6548 ( 9 ), 8-19.IV N.H.M.A.A.: W of Metahara, M (cf, 9), 28.XII S.M.F.: W of Metahara, (cf, ), 11.IX Note. - The species is, at present, known only from Ethiopia. G. FAMILY VESPERTILIONIDAE 1. Eptesicus somalicus somalicus (Thomas, 1901) Vespertilio minutus somalicus THOMAS, 1901, Ann. Mag. nat. Hist. (7) 8: 32. HargeLa, N Somalia. Records. - DE BEAUX, 1923: Asmara. - INGERSOL, 1968: Errer valley; Dacata valley.- HILL & MoRRIS, 1971: Fincha River (mouth); 10 km W of Mabil. Additional material.- A.M.N.H.: Dire Dawa (KoOPMAN, in litt.). A.N.S.P.: Sheik Hussein (KooPMAN, in litt.). B.M.:Didessa River bridge, (3 cfcf, ), 16.II.1971, 3-13.VII Koka, ( 9 ), 6.IV E shore of Lake Abiata, (2 cf cf), 17.X.1970, 21.X Wadere, Lake Abaya, Bulcha Forest, (4 cf cf, ), 24.XI.l970, 3-7.VII Arba Minch, (cf), M.K.B.: Lake Harairobi, uncatalogued (cf, ), 5.II Z.M.B.: Dire Dawa (KooPMAN, in litt.). 2. Eptesicus capensis (A. Smith, 1829) Vespertilio capensis A. SMITH, 1829, Zoo!. J. 4: 435. Cape Colony, South Africa. Records. -? SENNA, 1905: Adi Ugri (E. somalicus). - HILL & MoRRIS, 1971: NE of Lake Abaya. Additional material. - B.M.: Didessa River bridge, ( 9 ). 18.V Koka, ( 9 ), 17.III.l E shore of Lake

24 MAMMALS OF ETHIOPIA. I. CHIROPTERA 243 Abiata, ( ~ ), 16.X km S of Neghelli, ( ~ ), 21.X M.F.: Adi Ugri (KooPMAN, in litt., this appears to be the specimen recorded as somalicus by Senna, 1905 ). M.K.B.: Lake Harairobi, uncatalogued (2 ~ ~ ), 5.II Gambela, (d'), 22.VI.l972. Z.M.B.: Gurairobi See (KOOPMAN, in litt.). 3. Eptesicus guineensis (Bccage, 1889) Vesperus guineensis BOCAGE, 1889, J. Sci. math. phys. nat., Lisboa (2) 1: 6. Bissau, Pcr:uguese Guinea. Material. - B.M.: Gambela, ( d', ~ ), 27.XII.1969, l.i. 1970; (3 d'd', 4 ~ ~), 6-14.II.1970; (d', ~), VII M.K.B.: Gambela, (2 ~ ~ ), 6.XI.l972. N.H.M.A.A.: Gambela, M.459, (4 d'd'), 7-17.Il S.M.F.: Gambela, ( d', ~ ), Note. - This form has often been regarded as a race of E. capensis. Its specific status is indicated by the fact that small guineensis and larger capensis appear to be sympatric at Gambela. 4. Pipistrellus nanus (Peters, 1852) Vespertilio pipistrellus var. africana Ri.iPPELL, 1842, Mus. Senckenberg. 3: 156. Shoa. Ethiopia. Vespertilio nanus PETERS, 1852, Reise nach Mossambique, Sliugeth.: 63, pl. 16, f. 2. Inhambane. Mozambique. Records. - RiiPPELL, 1842: Shoa (africana).- MoNTICELLI, 1887: Let Marefia.- OusTALET, 1894: Djibouti. - DE BEAUX, 1922, 1924: Dolo. - DE BEAUX, 1925: Gondar. - DE BEAUX, 1930: Agordat. - THEODOR, 1957: Harar.- HILL & MoRRIS, 1971: Sabeta; Ghimbi. DoRST & PREVOST, 1972: W shore of Lake Abaya; 7 km from Ghinir. Additional material.- B.M.: Gondar, Dangila.- Didessa River bridge, (2 ~ ~ ), III.l970; ( ~ ), 2l.V Gambela, ( ~ ), 5.!.1970; (2 d' d', ~ ), VI.l970, 25.VII Ghibie River bridge, ( d' ), 3.! D.S.F.: Ambo, Am.II.14, 24 5, 40-1 (4 d' d', ~ ), M.K.B.: Gambela, ( ~ ), 22.VI.l972; 73.29, ( d', ~ ), 3-4.VIII.1972; (d'), 6.XI.l972; ( ~ ), N.H.M.A.A.: Meta Abo, M.20-1 (2 ex.), 4.IX Gambela, M.456 ( ~ ), 28.XII S.M.F.: Bahar Dar, (5d'd'), 6.VI.l971.

25 244 M. J. LARGEN, D. KOCK and D. W. YALDEN U.S.N.M.: Didessa River bridge, ( 1 ex.), Bulcha Forest, (5 ex.), 3-5.VI.l972, 10.XI Note. - KocK (1969) considered africana Riippell, 1842 to be conspecific with Pipistrellus kuhli (Kuhl, 1819) and a probable prior name for the race fuscatus Thomas, This synonymy cannot be maintained following an examination of the cranial remains of Ri.iPPELL's material, which indicates that africana is a composite. The lectotype (S.M.F. 4306) is found to be Pipistrellus nanus (Peters, 1852) for which africana is, therefore, an earlier name; a former syntype (S.M.F ) has proved to be a specimen of kuhli. In order to avoid the widespread confusion which would result from changing the name of one of the commonest African bats, we recommend that the name africana Riippell, 1842 be suppressed in accordance with Article 23 of the International Rules of Zoological Nomenclature. 5. Pipistrellus kuhli fuscatus Thomas, 1901 Pipistrellus kuhli fuscatus THOMAS, 1901, Ann. Mag. nat. Hist. (7) 8: 34. Naivasha, Kenya. Records. - Ri.iPPELL, 1842: Shoa (africana). - DoBSON, 1878; DoRIA, 1887: Senafe. -DoRIA, 1887: Let Marefia. - }ENTINK, 1888: Shoa. - SoRDELLI, 1902: Keren; Senafe. -DE BEAUX, 1925: Gondar. - HAYMAN, 1967 (P. aero); HILL, 1968; HAYMAN & HILL, 1971: S of Lake Tana (these records are based upon specimens from Dangila: B.M ; ). -HILL & MoRRIS, 1971: Ghimbi. Additional material. - A.M.N.H.: Addis Ababa. B.M.: Bulcha Forest, (2 d' d', 7 <.i? <.i? ), 5-7.VI.l971. F.M.N.H.: Near Debre Marcos. - Addis Ababa. M.K.B.: Addis Ababa, uncatalogued ( d' ), 25.X Pipistrellus rusticus marrensis Thomas & Hinton, 1923 Pipistrellus marrensis THOMAS & HINTON, 1923, Proc. zool. Soc. Lond.: 249. Foothills of southern Jebel Marra, Darfur, Sudan. Records. - HAYMAN & HrLL, 1971: NW Ethiopia (based upon a specimen from Wahina, Amhara in the Field Museum, Chicago). - LANZA, 1972: Aware Melca (P. nanus). Additional material. - B.M.: Didessa River bridge, ( <.i? ), 22.V Gambela, (2 d' d', 2 <.i? <.i? ), ; (10 d' d', 14 <.i? <.i? ), 4-16.II.1970; (2 <.i? <.i? ), 20.VIII. 1970, 25.VII.1971.

26 MAMMALS OF ETHIOPIA. I. CHIROPTERA 245 M.K.B.: Gambela, ( cf, <:( ), 30.VI.l972, l.vii.1972; (? sex), N.H.M.A.A.: Gambela, M.246, 455, 457, (5 <:( <:( ), XII. 1969, 8-9.II S.M.F.: Gambela, (2 cfcf), Pipistrellus rueppelli rueppelli (]. B. Fischer, 1829) Vespertilio riippellii J. B. FISCHER, 1829, Synopsis Mamm.: 109. Dongola, Sudan. (Nomen 12ovum for V. temmincki Cretzschmar, 1826: preoccupied). Records. - FITZINGER, 1866; HEUGLIN, 1877; ANDERSON & DE WINTON, 1902: Gallabat. - DoBSON, 1878: Abyssinia. Additional material. - B.M.: Bahadu, (2 cf cf, <:( ), S.M.F.: W shore of Lake Chamo, ( <:( ), 4.VIII Mimetillus moloneyi (Thomas, 1891) Vesperugo (Vesperus) moloneyi THOMAS, 1891, Ann. Mag. nat. Hist. (6) 7: 528. Lagos, Nigeria. Material. - M.K.B.: Gambela, (? <:( ), Note. - This is the first record of the genus for Ethiopia; the nearest localities from which other specimens have been obtained are West Pokot District (Kenya), West Nile District (Uganda) and Imatong Mountains (S Sudan). This material from NE Africa is still insufficient to allow subspecific determination. 9. Glauconycteris variegata (Tomes, 1861) Scotophilus variegatus TOMES, 1861, Proc. zool. Soc. Lond.: 36. Otjoro, SW Africa. Records.- HAYMAN & HILL, 1971; HILL & MoRRIS, 1971: 10 km W of Mabil (G. v. variegata). Additional material. - M.K.B.: Gambela, ( <:( ), 7.X Note. - Although the specimen from near Mabil has been referred to the typical race, that from Gambela agrees better in size of teeth with G. v. phalaena (Thomas, 1915) from Sudan. The racial variation of this species is poorly known, but colouration of the head, back and belly are not reliable characters. 10. Laephotis wintoni Thomas, 1901 Laephotis wintoni THOMAS, 1901, Ann. Mag. nat. Hist. (7) 7: 460. Kitui, Kenya. Records. - HILL (in press): Koka.

27 246 M. J. LARGEN, D. KOCK and D. W. YALDEN 11. Plecotus austriacus (J. B. Fischer, 1829) Vespertilio auritus austriacus J. B. FISCHER, 1829, Synopsis Mamm.: 117. Vienna, Austria. Records. - RuPPELL, 1842; PETERS, 1866: Shoa (P. auritus) - SoRDELLI, 1902; SENNA, 1905: Asmara (P. auritus). Note. - Ethiopian representatives of this genus are readily assignable to austriacus because of the large auditory bullae. They probably belong to the subspecies christiei Gray, 1838, but more specimens are needed to confirm this point. 12. Barbastella barbastellus leucomelas (Cretzschmar, 1830) Vespertilio leucomelas CRETZSCHMAR, 1830, in: Senck. naturf. Ges.: Atlas Reise nord!. Afrika E. Riippell, Saugeth.: 73, pl. 28 b. Sinai. Records. - HEUGLIN, 1861b, 1877; FITZINGER, 1866: Massawa. - HAYMAN & HILL, 1971: Eritrea (based upon a specimen in the Museum of Comparative Zoology, Harvard). 13. Miniopterus inflatus africanus Sanborn, 1936 Miniopterus africanus SANBORN, 1936, Zoo!. Ser. Field Mus. Nat. Hist. 20: 111. Sanford's Ranch, Mulu, Ethiopia. Records. - SANBORN, 1936: Mulu. - INGERSOL, 1968: Alemaya; Errer valley.- HILL & MoRRIS, 1971: Awash National Park.- DoRST & HILL, 1972: So Omar. Additional material. - B.M.: Suha River, Danka River, near Dinshu, (2 cf cf, 40 Cjl Cjl ), 19.XII So Omar, (3 cf cf, 7 Cjl Cjl ), 27.VI.l970. N.H.M.A.A.: Danka River, near Dinshu, M (2 Cjl Cjl ), 19.XII So Omar, M (3 cf cf, 4 Cjl Cjl ), 27.VI S.M.F.: So Omar, (cf, Cjl ), 27.VI.l Miniopterus schreibersi arenarius Heller, 1912 Miniopterus natalensis arenarius HELlER, 1912, Smilhson. misc. Collns 60 (12): 2. Guaso Nyuki, N Guaso Nyiro, Kenya. Records. - HEDGLIN, 1877: Gorgora caves. - THOMAS, 1898; FuNAIOLt & LANZA, 1968: Dass Wells. - INGERSOL, 1968: Gara Mullata. Additional material.- B.M.: Gondar.- Nur Mohammed, (2 cfcf), 28.III.l972.- So Omar, (cf, 3 Cjl Cjl ), XII Arba Minch, (cf), 16.VIII F.M.N.H.: Mulu (KooPMAN, in litt.).

28 MAMMALS OF ETHIOPIA. I. CHIROPTERA 247 N.H.M.A.A.: N end of Lake Abaya, M.677 ( <.;? ), -.VII S.M.F.: W shore of Lake Abaya, ( d', 2 <.;? <.;? ), 30.VII. 1973; ( d', <.;? ), 6.VIII Note. - In Ethiopia this species is distinguished by having a forearm length of less than 45 mm, while that of adult M. inflatus exceeds 48 mm. HAYMAN & HILL ( 1971) record a range of for schreibersi and for inflatus, but these figures include measurements from allopatric populations. 15. Nj'Cticeius schlieffeni schlieffeni Peters, 1859 Nycticejus schlieffenii PETERS, 1859, Mber. K. preuss. Akad. Wiss. Berlin: 224. Cairo, Egypt. N(ycticejus) adovanus HEUGLIN. 1877, Reise in Nordost Afrika, 2: 34. Asam River, Adowa, Ethiopia. Records. - HEUGLIN, 1877: Asam River. - THOMAS & DoRIA, 1886: Massawa (Scotophilus pallidus). - THOMAS, 1890; DE BEAUX, 1923: Massawa. - ANDERSON & DE WINTON, 1902: Tigre (Province); Assab. - DE BEAUX, 1922, 1923, 1924: Dolo. - DE BEAUX, 1923: Assab. - HARRISON, 1961a: Kibbish Wells. Additional material. - B.M.: Bahadu, ( <.;? ), 22.IV Filwoha, ( d' ), 24.XII Awash Falls, ( d' ), 28.IV M.K.B.: Gambela, ( <.;? ), 23.VI.l972. M.S.N.G.: Gonderaba. 16. Nycticeius (Scotoecus) hindei (Thomas, 1901) Scotoecus hindei THOMAS, 1901, Ann. Mag. nat. Hist. (7) 7: 264. Kitui, Kenya. Material. - B.M.: Didessa River bridge, ( d' ), 6.VII Bulcha Forest, (d'), 30.XI.1969; (d'), 8.VI.l971. M.N.S.: Ethiopia (probably Eritrea), 1057 ( d' ), 1863, Heuglin coll. 17. Nycticeius (Scotoecus) hirundo (De Winton, 1899) Scotophilus hirundo DE WINTON, 1899, Ann. Mag. nat. Hist. (7) 4: 355. Gambaga, Ghana. Material. - B.M.: Didessa River bridge, ( <.;? ), 6.VI.l Gambela, (2 d'd', 6 <.;? <.;?), 6-17.II.1970; (<.;?), 1.!.1970; ( <.;? ), 16.VI.l970. M.K.B.: Gambela, (2 <.;? <.;? ), VI.l972; ( <.;? ), 6.VIII.l972; 73.10, 12, 14-16, 67 (3 d' d', 3 <.;? <.;? ), N.H.M.A.A.: Gambela, M (2 d'd', 3 <.;? <.;? ), 8-17.II Didessa River bridge, M.444 ( <.;? ), 20.III.l970. S.M.F.: Gambela, ( d' ), 8.II.1970; ( d', <.;? ), 1972.

29 248 M. J. LARGEN, D. KOCK and D. W. YALDEN Note. - HILL (in litt.) thinks it possible that the dark-winged members of this genus may represent two species; the smaller ( hirundo) having a forearm less than 33 and condylobasallength less than 13.2, the larger (hindei) with the forearm above 33 and condylobasallength greater than Although the two forms are here listed separately, further specimens are needed to establish this difference. 18. Myotis (Selysius) tricolor (Temminck, 1832) Vespertilio tricolor TE;.,IMINCK, 1832, in Smuts: Enum. Mamm. Capensium: 106. Cape Town, South Africa. Records.- HAYMAN, 1967; HILL & MoRRIS, 1971: Ethiopia (based upon a specimen from Dangila: B.M ). Additional material. - B.M.: E shore of Lake Abiata, ( d' ), 16.X Nur Mohammed, ( ~ ), 28.III Myotis (Selysius) scotti Thomas, 1927 Myotis scotti THOMAS, 1927, Ann. Mag. nat. Hist. (9) 19: 554. Djemdjem Forest, Ethiopia. Records. - THOMAS, 1927: Djemdjem Forest. - HAYMAN, 1960: Woliso. - HAYMAN, 1967; HAYMAN & HILL, 1971: 70 miles SW of Addis Ababa ( = Woliso). - DoRST & PREVOST, 1972: W shore of Lake Abaya; Addis Alem. Additional material. - B.M.: Gabba River bridge, ( d' ), km E of Shashamane, ( d' ), 13.X Note. - The species is known only from Ethiopia. 20. Myotis (Selysius) morrisi Hill, 1971 Myotis morrisi HILL, 1971, in Hill & Morris: Bull. Br. Mus. nat. Hist. (Zool.) 21 (2): 43, pl. 1, 2. Didessa River (mouth), Ethiopia. Note. - The species is, at present, known only from the type. 21. Myotis (Chrysopteron) welwitschi (Gray, 1866) Scotophilus welwitschii GRAY, 1866, Proc. zool. Soc. Lond.: 211, pl. 24. Angola. Records. - RoDE, 1933: Gondar. N.B. - Myotis (Myotis) blythi (Tomes, 1857) has been recorded from «Abyssinia» by DoBSON, 1878 (under the name Vespertilio murinus). It appears that DoBSON's purchased specimen has never been re-examined, but the occurrence of this species in Ethiopia seems most unlikely.

30 MAMMALS OF ETHIOPIA. I. CHIROPTERA Scotophilus nigrita flavigaster (Heuglin, 1861) Nycticejus flavigaster HEUGLIN, 1861, Nova Acta Acad. Caesar Leop. Carol., Halle 29 (8): 5, line 14. Near Keren, Ethiopia. Nycticejus murino-flavus HEUGLIN, 1861, Nova Acta Acad. Caesar Leop. Carol., Halle 29 (8): 5, line 15. M'kullu (= Moncullo), Ethiopia. Records. - HEDGLIN, 1861a, 1877: Moncullo; Gondar (murinoflavus). - HEDGLIN, 1861a, 1877: near Keren (flavigaster). - DoBSON, 1878: Abyssinia (borbonicus). - }ENTINK, 1888: Bogos. -DE WINTON, 1900: Bilo. - SoRDELLI, 1902: Sia-ha (borbonicus). - THOMAS, 1902: Harar. - SENNA, 1905: Sia-ha; Keren. - DRAKE-BROCKMAN, 1910: Harar. - DE BEAUX, 1922: Harar (borbonicus). - BEADLES & INGER SOL, 1968: Alemaya. - lngersol, 1968: Urso River; Dacata valley; Errer valley; between the Dacata and Fafan valleys. Additional material. - B.M.: Dangila.- Bahadu, ( <;? ), 2.V Ghimbi, {d').- Koka, (d'), 8.IV Gambela, ( d' ), 28.XII E shore of Lake Abiata, (6 d' d', 3 <;? <;? ), X.1970, X Dobleh.- Wadere km N of Yavello, ( <;? ), 3.III Hara Digallu, Borana. M.K.B.: Gambela, ( d' ), M.N.S.: Keren, A ( d', <;? ), 1862, Heuglin coli. S.M.F.: W shore of Lake Abaya, ( <;? ), 7.VIII.l973. U.S.N.M.: Bulcha Forest (3 ex.), 3.VII.1972, 14.XI Z.M.B.: Dire Dawa. Note. - S. n. flavigaster is considered to include colias Thomas, Scotophilus leucogaster (Cretzschmar, 1830) Nycticejus leucogaster CRETZSCHMAR, 1R30, in Senck. naturf. Ges.: Atlas Reise nord!. Afrika E. Ruppel!, Saugeth.: 71, pl. 28 a. Bir Nedger, Kordofan, Sudan. Material.- B.M.: Gondar.- Adur, 20 miles SW of Lake Tana. Gambela, (2 d' d', 2 <;? <;? ), 28.XII.1969; ( d', <;? ), ; ( d', <;? ), 5-9.II.1970; (5 d' d', 6 <;? <;? ), VI.1970, 23.VIII.1970, 31.VII-5.VIII.l971. F.M.N.H.: Near Dungulbar. M.K.B.: Gambela area, {d'), Gambela, , , (5 d' d', 6 <;? <;? ), VI.1972, 2-14.VII.1972, 2-24.VIII. 1972; , (7 d' d', 3 <;? <;? ), VIII-IX N.H.M.A.A.: Gambela, M.423-6, (5 d' d', 3 <;? <;? ), 26.XII ; M.432 5, (5 d' d', <;? ), 6-1l.II S.M.F.: Gambela, ( d', <;? ), 29.XII.1969, 10.II U.S.N.M.: Bulcha Forest, (5 ex.), 5.VII.1972, VII.1972.

31 250 M. J. LARGEN, D. KOCK and D. W. YALDEN Note. - It has been suggested that leucogaster might be regarded as a smaller race of nigrita, the two being difficult to distinguish in NE Africa although readily separable elsewhere (HAYMAN & Hill, 1971). 24. Kerivoula harrisoni harrisoni Thomas, 1901 Kerivoula harrisoni THOMAS, 1901, Proc. zoo!. Soc. Lond.: 802, footnote. Walamo (region), between Lakes Zwai and Margherita, Ethiopia. Note. - In Ethiopia the species is known only from the type locality, although widely distributed elsewhere in eastern Africa. 25. Kerivoula eriophora (Heuglin, 1877) Nycticejus eriophorus FITZINGER, 1866, Sber. K. Akad. Wiss. Wien 54 (!): 547. Bellegas valley, Ethiopia (nomen nudum). Nycticejus eriophorus HEUGLIN, 1877, Reise in Nordost-Afrika 2: 34. Belegaz valley, Ethiopia. Note. - The species is known only from the type description since no specimen can be found in the Museum fiir Naturkunde, Stuttgart. It is possible that this name might eventually replace africana Dobson, H. FAMILY MOLOSSIDAE 1. Otomops martiensseni martiensseni (Matschie, 1897) Nyctinomus martiensseni MATSCHIE, 1897, Arch. Naturgesch. 63 (I): 84. Magrotto plantation, near Tanga, Tanzania. Material. - B.M.: So Omar, (5 d d), -.III.l971; (d, 4<;?<;?), XII N.H.M.A.A.: So Omar, M (2 d d), -.III.1971; M.637 ( <;? ), 27.XII S.M.F.: So Omar, (2 d d), -.III Note. - This is the first record of the genus in Ethiopia, connecting known localities in Kenya with a single record from French Somaliland (l1 46'N 42 39'E, alt m) by HILL & MoRRIS (1971). 2. Platymops setiger macmillani Thomas, 1906 Platymops macmillani THOMAS, 1906, Ann. Mag. nat. Hist. (7) 17: 500. Between Addis Ababa and Lake Rudolf, Ethiopia. Records. -THOMAS, 1906: between Addis Ababa and Lake Rudolf. -DE BEAUX, 1943: Caschei. Additional material. - B.M.: Mui, (d), 10.IV.1972.

32 MAMMALS OF ETHIOPIA. I. CHIROPTERA Tadarida (Chaerephon) pumila pumila (Cretzschmar, 1830) Dysopes pumilus CRETZSCHMAR, 1830, in: Senck. naturf. Ges.: Atlas Rei e nord!. Afrika E. Riippell, Saugeth.: 69, pl. 27. Massawa, Ethiopia. Records. - CRETZSCHMAR, 1830; FITZINGER, 1866; HEUGLIN, 1861a, 1877; THOMAS & DoRIA, 1886; HOLLISTER, 1918: Massawa. - HEUGLIN, 1861a: Moncullo. - THOMAS & DORIA, 1886: Shoa. - SENNA, 1905: Abd elkader. -KLATT, 1913: Mai AtaL- HoLLISTER, 1916, 1918: Saaita. -FLOWER, 1932: Gallabat. - BRAESTRUP, 1933: Harar, Galab ( = Gallabat); Dangila. - DE BEAUX, 1939: Om Ager. - SMIT, 1957: Harar.- ANDRAL et al, 1968: Manera Forest (T. limbata). - BEADLES & lngersol, 1968: Errer valley, at bridge on Jigjiga road. - lngersol, 1968: Alemaya; Errer valley; Gota River. - KocK, 1969: Kibbish Wells. - HILL & MoRRIS, 1971: Awash National Park; Ghimbi. Additional material. - B.M.: Gondar. - Piccolo Abbai, (2 d' d', 7 <i? <j? ). - Backo, (3 d' d', 10 <j? <j? ), VII Aware Melca, ( <j? ), 22.XII N bank of Awash River, (2 d' d', <j? ), 29.XII Gambela, (2 <j? <j? ), ; (2 <;? <;? ), 5-14.II.1970; (2 d'd'), VI km S of Zwai village, ( d', <j? ), 7JCI.l km S of Lake Zwai, ( <j? ), 7.XI.l969.- E shore of Lake Abiata, (9 d' d', 5 <j? <j? ), X.1970, 19.X W shore of Lake Langano, (2 d' d', 2 <j? <j? ), D.S.F.: Bishoftu, Bi.IV.3 ( <;? ), 13.III.l956. F.M.N.H.: Near Metemma (KooPMAN, in litt.). M.K.B.: Gambela, ( <j? ), 25.VI.l972; (d'), 6.VII.1972; ( d', <j? ), VII.1972; (3 d' d', 2 <j? <j?, 1? sex), M.S.N.G.: Massawa.- Ghinda.- Dorfu.- Zeghie. (All KooPMAN, in litt. ). N.H.M.A.A.: Backo, M. 111, 113-4, 117, (d', 6 <;? <;? ), VII N bank of Awash River, M. 416 ( <j? ), 26.IX Gambela, M ( d', <j? ), 8-10.II W shore of Lake Langano, M ( d', 3 <;? <;? ), 27.II S.M.F.: Backo, (2 d' d' ), 24.VII E shore of Lake Awasa, (5 d' d' ), 2.IX W shore of Lake Abaya, (21 d' d', 50 <;? <i? ), 7.VIII U.S.N.M.: Harar (KooPMAN, in 1itt.). Note. - A single, recently acquired specimen (U.S.N.M.: Bulcha Forest, d', ) is tentatively referred to Tadarida (Chaerephon) chapini (J. A. Allen, 1917); a little-known form whose status has yet to be satisfactorily established. HAYMAN & HILL (1971) accept the validity

33 252 M. J. LARGEN, D. KOCK and D. W. YALDEN of this species and the long, distinctly bicoloured inter-aural crest of the present specimen seems to make it readily distinguishable from Ethiopian examples oft. pumila. However, KocK (1969) suggests, on the basis of published measurements of the skull, that chapini and pumila are conspecific. 4. Tadarida (Chaerephon) nigeriae nigeriae (Thomas, 1913) Chaerephon nigeriae THOMAS, 1913, Ann. Mag. nat. Hist. (8) 11: 319. Zaria Province, N Nigeria. Records. - lngersol, 1968: Gota River. - HILL & MoRRIS, 1971: Awash National Park (N bank of Awash River). 5. Tadarida (Chaerephon) bivittata (Heuglin, 1861) Nyctinomus bivittatus HEUGLIN, 1861, Nova Acta Acad. Caesar Leop. Carol., Halle 29 (8): 4, 13. Near Keren, Ethiopia. Records. - HElJGLIN, 1861a; HAYMAN & HARRISON, 1966: near Keren.- HEDGLIN, 1877: between the Anseba River and Keren; Atirba River. - lngersol, 1968: Doha River; Dacata River. 6. Tadarida (Mops) midas midas (Sundevall, 1843) Dysopes midas SUNDEVALL, 1843, K. svenska Vetensk-Acad. Handl. for 1842: 207. Bahr el Abiad (= White Nile), Sudan. Records. - THOMAS & DORIA, 1886; DoRIA, 1887: Moncullo (Nyctinomus cestonii). - SENNA, 1905: Massawa. 7. Tadarida (Mops) condylura (A. Smith, 1833) Nyctinomus condylurus A. SMITH, 1833, S. Afr. q. J. 2: 54. Durban, South Africa. Material. - B.M.: Koka, ( ~ ), 13.III km S of Zwai village, (4cfcf, 7 ~~), 7.XI.l969.- E shore of Lake Abiata, ( cf ), 27.X M.K.B.: Gambela, 73.46, 61-2 (2 cf cf, ~ ), N.H.M.A.A.: 1 km S of Lake Zwai, M (2 cf cf, 6 ~ ~ ), 7.XI.l969. S.M.F.: 7 km S of Zwai village, (cf, ~ ), 7.XI.l969. Note. - Although common and widely distributed in sub-saharan Africa, this appears to be the first record of the species in Ethiopia.

34 MAMMALS OF ETHIOPIA. I. CHIROPTERA Tadarida (Mops) nanula (J. A. Allen, 1917) Mops (Allomops) nanulus J. A. ALLEN, 1917, Bull. Am. Mus. nat. Hist. 37: 477. Niangara, Zaire. Material. - B.M.: Gambela, ( d' ), 13.VI.l970. M.K.B.: Gambela, ( d' ), 2.VIII Note. - A forest form extending from West Africa to Uganda and Kenya but not previously recorded from Ethiopia. 9. Tadarida (Tadarida) ansorgei (Thomas, 1913) Nyctinomus ansorgei THOMAS, 1913, Ann. Mag. nat. Hist. (8) 11: 318. Malange, Angola. Records. - HARRISON, 1959: Kibbish Wells. Additional material. - B.M.: Addis Ababa, ( 9 ), 24.VI E shore of Lake Abiata, ( 9 ), 20.X M.K.B.: Gambela area, ( 9 ), Tadarida (Tadarida) aegyptiaca (E. Geoffroy, 1818) Nyctinomus aegyptiacus E. GEOFFROY, 1818, Descript. de I'Egypte, 2: 128. Egypt. Records. - SORDELLI, 1902: Keren. 11. Tadarida (Tadarida) ventralis (Heuglin, 1861) Nyctinomus ventralis HEDGLIN, 1861, Nova Acta Acad. Caesar Leop. Carol., Halle 29 (8): 4, 11. Near Keren, Ethiopia. Nyctinomus africanus DoBSON, 1876, Ann. Mag. nat. Hist. (4) 17: 348. Transvaal, South Africa. Records. - HEUGLIN, 1861a: near Keren. - HEUGLIN, 1877: Anseba River, near Keren. - JABLONOWSKI, 1898: Keren (africanus). - THOMAS, 1928: Lake Tana region (ventralis). - HAYMAN, 1967; HAYMAN & HILL, 1971: 100 miles S of Lake Tana (africana). - HAR RISON, 1971; HILL & MoRRIS, 1971: Fatam River (africana).- HAYMAN & HILL, 1971; HILL & MoRRIS, 1971:?vicinity of Addis Ababa (africana). Note. - DoBSON ( 1878) synonymised both ventralis Heuglin, 1861 and midas Sundevall, 1843 with cestonii Savi, 1825 ( =teniotis Rafinesque, 1814). HEUGLIN (1861a, 1877) writes of several specimens of his ventralis being identified as cestonii upon their arrival at the Stuttgart Museum, but insisted that these were distinct from teniotis. Subsequently, it became apparent that midas and teniotis represent different taxa (DE WINTON, 1901) but ventralis was not re-examined and has since been 18

35 254 M. J. LARGEN, D. KOCK and D. W. YALDEN generally regarded as a synonym of midas, although not, apparently, by THOMAS (1928). The present study has involved a re-examination of all bats collected by HEDGLIN and now in the Museum fiir Naturkunde, Stuttgart. A single alcohol-preserved specimen with the skull in situ (M.N.S. 982) was found to be labelled «ventralis» but this appears to be the individual identified by JABLONOWSKI ( 1898) as africanus. Its short ears distinguish it from lobata Thomas, 1891 and the large cranial dimensions from fulminans Thomas, The specimen lies close to the lower end of the africanacluster in HARRISON's (1971) scatter diagram and the two would appear to be conspecific. 12. Tadarida (Mormopterus) acetabulosus natalensis (A. Smith, 1847) Dysopes natalensis A. SMITH, 1847, Illustr. Zool. South Africa, Mamm., pl. 49. Near Durban, South Africa. Records. - HAYMAN, 1967; HAYMAN & HILL, 1971; HILL & MoRRIS, 1971: between Shoa and Lake Rudolf. IV. ZOOGEOGRAPHICAL ASPECTS OF THE BAT FAUNA OF ETHIOPIA* The present work lists 72 species of Chiroptera from Ethiopia, of which only five appear to be represented by more than one subspecies. For a country of such diverse habitats, the number of species might be considered low, but the figure corresponds well with those for other, comparable regions of Africa (KocK, 1973 ), despite the fact that intensive collecting in Ethiopia has only recently begun. The bat fauna, whose composition is summarised in Table 1, may be considered to represent three main zoogeographical elements. A. The Palaearctic element This element is divisible into two subgroups: 1. Species widely distributed in the African faunal region and represented by subspecies different from those in the Palaearctic. Rhinotophus blasii, Pipistrellus kuhli, Miniopterus schreibersi. 2. Species represented only marginally in the African region by the same subspecies as are found in the Palaearctic. Rhinolophus hipposideros, Plecotus austriacus, Barbastella barbastellus. * In this account the term «Ethiopian» has been deliberately avoided. In order to eliminate confusion, the zoogeographical region usually referred to as Ethiopian is here termed the African region.

36 MAMMALS OF ETHIOPIA. I. CHIROPTERA 255 B. The Saharo-Sindian element In this group are placed four species which are found in the arid areas of the African, Palaearctic and Oriental faunal regions but cannot readily be assigned to any one of these. Rhinopoma hardwickei, R. muscatellum, T aphozous nudiventris, Asellia tridens. C. The African element The majority of species found in Ethiopia (86%) are essentially African in distribution. A number of subgroups are recognisable: 1. Species found also in the Madagascan region. Eidolon helvum, Rousettus lanosus, T aphozous mauritianus, Hipposideros commersoni, Scotophilus nigrita, S. leucogaster, Otomops martiensseni, T adarida pumila, T. midas, T. condylura, T. acetabulosus. It is of interest that 45% of these species are molossids while the remainder are relatively large forms, which perhaps indicates the correlation between superior powers of flight and the ability to cross water barriers. In addition to these species, it is possible that Nycteris thebaica also reaches the Madagascan region (HAYMAN & HILL, 1971). 2. Species which are widely distributed in Africa but also penetrate into the arid Saharo-Sindian belt and thus reach the southern Palaearctic or even western Oriental regions. Rousettus aegyptiacus, T aphozous perforatus, Rhinolophus clivosus, Triaenops persicus, Pipistrellus rueppelli, T adarida aegyptiaca. 3. Species which are essentially African but also occur as relicts in the non-arid areas of the Palaearctic. Nycteris thebaica, Hipposideros calfer, Nycticeius schlielfeni. 4. Species typical of the arid areas of NE Africa, termed the Somali arid zone. (Because of climatic similarities, this region also harbours most of the species found in the Saharo-Sindian belt). Cardioderma cor, Platymops setiger. A further species, Asellia patrizii, is restricted to the Somali arid zone but, since apparently endemic to Ethiopia, is considered separately. 5. Species typical of the East African highlands. At present this group seems to contain only Hipposideros megalotis since Rousettus lanosus, which is otherwise characteristic of this region, occurs also in Madagascar. 6. Species typical of the Central African lowland forests. Hypsignathus monstrosus, Rousettus angolensis, Hipposideros fuliginosus,? Mimetillus moloneyi, T adarida nanula.

37 256 M. J. LARGEN, D. KOCK and D. W. YALDEN 7. Species at present considered endemic to Ethiopia. Asellia patrtzzt, Myotis morrisi, M. scotti, Kerivoula eriophora. It may be noted that A. tridens, the only other representative of Asellia, has a Saharo Sindian distribution. Table 1. The zoogeographical composition of the bat fauna of Ethiopia. Zoogeographical category No. of species % of total A. Palaearctic element 1. Widely distributed in the African region 2. Marginally distributed in the African region B. Saharo-Sindian element C. African element 1. Found also in the Madagascan region 2. Extending also into the Saharo Sindian 3. Marginal in the Palaearctic 4. Somali arid 5. East African montane 6. Central African forest 7. Endemic 8. Unclassified TOTAL ( + 1 endemic) (4.2) The remaining species of bat found in Ethiopia ( 42% of the total) are considered to be too poorly known to be attributed with certainty to any particular geographical region. However, it seems likely that at least some of these may be confined to the savanna areas of East Africa. This brief analysis is sufficient to demonstrate the zoogeographical diversity of the Chiroptera of Ethiopia. Considering the great variety of habitats in this country, the very slight subspecific diversity is, perhaps, surprising and suggests a precise niche selection by bats. However, both this feature and the relatively low percentage of endemic forms is likely to be correlated with the comparative mobility of these flying animals.

38 MAMMALS OF ETHIOPIA. I. CHIROPTERA 257 CoRBET & YALDEN (1972), utilising the figures of CARPENTER (1935), CARCASSON (1964) and URBAN & BROWN (1971), calculate an endemicity in Ethiopia of approximately 2% of the butterfly fauna and 3.5% amongst birds. They record 11 species of endemic mammal, other than bats and marine forms, of which six are rodents. The addition of two further rodent species, subsequently described by PETTER ( 1972 ), indicates an endemicity of about 9.6% amongst such forms. V. THE HABITAT AND FAUNAL COMPOSITION AT SOME MAJOR COLLECTING SITES Although much remains to be learned of the Ethiopian bat fauna and its distribution, a number of localities, largely as a result of intensive study within the past few years, are considered to be relatively well known. Most new material, here recorded for the first time, has been obtained at these sites. Awash National Park ca 08 54'N 39 55'E A region of semi-desert grassland with areas of Acacia scrub and volcanic rubble, most of which lies between 900m and 1100m, but including Mount Fantalle, a dormant volcano which rises to 1700m. The park includes a major river, the Awash, and its smaller tributary, the Cassam, both of which are fringed by a narrow zone of riverine vegetation with large Acacia or Ficus trees. A feature which undoubtedly has an important influence upon the bat fauna is the presence of numerous lava ' blisters '; hollow mounds originally formed as gas pockets in molten lava, many of which have since perforated to form caves which provide roosts for a large bat population (see also HILL & MoRRIS, 1971; LANZA, 1972). 16 species: Rhinopoma hardwickei sennaariense, Taphozous perforatus haedinus, Nycteris thebaica brockmani, N. h. hispida, Cardioderma cor, Rhinolophus clivosus acrotis, R. hipposideros minimus, Hipposideros c. caffer, H. commersoni marungensis, Asellia t. tridens, A. patrizii, Nycticeius s. schlieffeni, Pipistrellus rusticus marrensis, Miniopterus in flatus africanus, Tadarida p. pumila, T. n. nigeriae. Bahadu A small area of scattered Ficus and shrubs in the open flood-plain of the Awash River, surrounded by semi-desert scrub not greatly different from the Awash National Park but rather more arid and rocky. The relatively small number of bat species, as compared with the Awash Park, may perhaps reflect a paucity of suitable roosting sites.

39 258 M. J. LARGEN, D. KOCK and D. W. YALDEN 8 species: Epomophorus minor, Coleura afra, Nycteris h. hispida, Cardioderma cor, Triaenops persicus afer, Pipistrellus r. rueppelli, Nycticeius s. schlieffeni, Scotophilus nigrita flavigaster. Sof Omar 06Q54'N 40 48'E 1340m An extensive limestone cave system carved by the Ueb River which here flows underground for a distance of 1.5 km. Outside the cave the steep, rocky sides of the river gorge are covered by dense mixed Acacia/ broad-leaved scrub, while a narrow zone of more luxuriant vegetation fringes the river itself. All bats have been collected at roost in the cave with the exception of Epomophorus, Hipposideros and Triaenops, and the latter probably utilises some unvisited part of the system since large numbers were found flying at dusk near the cave entrance (see also MoNon & MoRTON, 1972). 11 species: Epomophorus minor, Rousettus aegyptiacus, Rhinolophus landeri dobsoni, R. simulator, R. blasii andreinii, R. f. fumigatus, Hipposideros c. caffer, Triaenops persicus afer, Miniopterus inflatus africanus, M. schreibersi arenarius, Otomops m. martiensseni. 1700m A large Rift Valley reservoir formed by the damming of the A wash River and including the former Lake Arairobi. The surrounding area consists of Acacia woodland extensively degraded to provide open farmland. 9 species: Epomophorus minor, Rousettus aegyptiacus, Rhinolophus landeri dobsoni, R. simulator, R. blasii andreinii, Triaenops persicus afer, Laephotis wintoni, Eptesicus s. somalicus, E. capensis. Lake Abiata 07 37'N 38 39'E 1590m A rather strongly alkaline Rift Valley lake with open, heavily-grazed shores surrounded by Acacia woodland. A few scattered Ficus trees border a nearby small river and low cliffs provide caves in which Nycteris roosts. 13 species (Taphozous has been recorded from adjacent Lake Shalla): Epomophorus minor, Rousettus aegyptiacus, T aphozous perforatus haedinus, Nycteris thebaica labiata, Rhinolophus simulator, R. f. fumigatus, Eptesicus s. somalicus, E. capensis, Myotis tricolor, Scotophilus nigrita flavigaster, Tadarida p. pumila, T. condylura, T. ansorgei. Arba Minch 1400m A sprawling town over-looking the isthmus between Lakes Abaya and Chama in the southern Rift Valley. The dominant vegetation in the area is Acacia scrub but the Collufu River, which flows between the two lakes, is fringed by well developed riverine forest with very large Ficus

40 MAMMALS OF ETHIOPIA. I. CHIROPTERA 259 trees. The lakes themselves, at an altitude of about 1300m, support extensive reed beds at their margins. 11 species: Epomophorus labiatus anurus, Rousettus aegyptiacus, Nycteris h. hispida, Lavia frons, Cardioderma cor, Hipposideros c. caffer, H. ruber centralis, Eptesicus s. somalicus, Miniopterus schreibersi arenarius, Scotophilus nigrita flavigaster, T adarida p. pumila. Bulcha Forest 1800m This collecting site is close to a small river flowing into the east side of Lake Abaya, at the transition zone between riverine forest and Acacia woodland. 9 species: Epomophorus labiatus anurus, E. g. gambianus, Micropteropus pusillus, Rousettus aegyptiacus, Nycteris h. hispida, Rhinolophus landeri dobsoni, Eptesicus s. somalicus, Pipistrellus kuhli fuscatus, Nycticeius hindei. Didessa River bridge 09 02'N 36 09'E 1200m This locality is situated on the edge of the Didessa River gorge at the transition zone between three main habitat types; dense, luxuriant tropical deciduous forest, Combretum /T erminalia woodland, and savanna/ farmland derived from the two. 15 species: Epomophorus g. gambianus, Micropteropus pusillus, Rousettus aegyptiacus leachi, R. angolensis, Nycteris thebaica, N. aethiopica, Rhinolophus clivosus acrotis, R. f. fumigatus, Hipposideros ruber centralis, Eptesicus s. somalicus, E. capensis, Pipistrellus nanus, P. rusticus marrensis, Nycticeius hindei, N. hirundo. 515m Hot, humid, marshy grassland with sorghum and maize plantations bordering the Baro River, and Combretum/Terminalia woodland on the surrounding higher ground. Most specimens have been collected close to a small patch of riverine forest which perhaps serves as a focus for fruit bats. The remarkable richness of the fauna in this area is illustrated not only by the number of bat species but also in other groups which have been studied, including birds, Amphibia and Coleoptera. 27 species: Epomophorus g. gambianus, Micropteropus pusillus, Eidolon helvum, Rousettus aegypiiacus leachi, T aphozous mauritianus, Nycteris h. hispida, N. aethiopica, Lavia frons, Rhinolophus clivosus, R. landeri dobsoni, R. fumigatus exsul, Hipposideros c. caffer, H. ruber centralis, Eptesicus guineensis, E. capensis, Pipistrellus nanus, P. rusticus marrensis, Glauconycteris variegata, Mimetillus moloneyi, Nycticeius s. schlieffeni, N. hirundo, Scotophilus nigrita flavigaster, S. leucogaster, Tadarida p. pumila, T. nanula, T. condylura, T. ansorgei.

41 260 M. J. LARGEN, D. KOCK and D. W. YALDEN ' -- 1\ I ERITREA \~~ Sea! 15''-t !----f--t---t V-I-\ "-.+~ ' Suda1 --,--t_ / ---.,- 'z._ ~~ I i f-t-- 1 HARAR I~ ILUBABOR / o ARUSI -~~~- --, 0 g a d e n / 1\ A 'KAFFA ; c:;j /-- / "\:... I <".. \ { / \',,----' ),~ '1, BALE ',, I ~~~~ ~~,., / ' \ ' -..,, --., / ' SIDAMO -- '\ s'lh---!l/~~~t~~i~~~~b~~-,~~:t~~- =-- - ~B~o-rr_a nt--a~~-~---~r---i~--~~~ -=-~--1-so_m_a_ltia---ir~ -...-A. \., I Fig Figs ~... \70~0 ~ Ethiopia: Provinces and other regions not included in the gazeteer. Symbols: Locality precisely identified, identification of specimen accepted. Locality precisely identified; identification of specimen open to doubt. Locality approximate or doubtful; identification of specimen accepted. Type locality.

42 MAMMALS OF ETHIOPIA. I. CHIROPTERA 261 A E. labiotus anurus E. gambianus gambianus Fig The recorded distribution of Epomophorus labiatus labiatus, E. labiatus anurus and E. gambianus gambianus in Ethiopia.

43 262 M. J. LARGEN, D. KOCK and D. W. YALDEN e Micropteropus pusillus Epomophorus minor 9 Hypsignathus monstrosus Eidolon helvum Fig The recorded distribution of Micropteropus pusillus, Epomophorus minor, Hypsignathus monstrosus and Eidolon helvum in Ethiopia.

44 MAMMALS OF ETHIOPIA. I. CHIROPTERA 263 e Rousettus aegyptiocus.a R. angolensis R. Ia nos us kempi ~- 0 ~") o Fig The recorded distribution of Rousettus aegyptiacus, R. angolensis and R. lanosus kempi in Ethiopia.

45 264 M. J. LARGEN, D. KOCK and D. W. YALDEN e Rhinopoma hardwickei sennaa riense 6 R. muscatel I urn 9. Taphozous mauritianus Coleura afro Fig The recorded distribution of Rhinopoma hardwickei sennaariense, R. muscatellum, Taphozous mauritianus and Coleura afra in Ethiopia.

46 MAMMALS OF ETHIOPIA. I. CHIROPTERA 265 e Taphozous perloratus haedinu A T. perforatus? perforatus 9 T. nudiventris Fig The recorded distribution of Taphozous perforatus haedinus, T. perforatus? perforatus and T. nudiventris in Ethiopia.

47 266 M. J. LARGEN, D. KOCK and D. W. YALDEN e Nycteris thebaica sspp. A N.t. labiata N.t. brockmani N.t. media CJ ') 0 ~~ 0 Fig The recorded distribution of Nycteris thebaica sspp. in Ethiopia. The type localities of N. t. labiata, N. t. brockmani and N. t. media are indicated.

48 MA.M.MALS OF ETHIOPIA. I. CHIROPTERA 267 e _ Nycteris hispida hispida A Nycteris oethiopico Fig The recorded distribution of Nycteris hispida hispida and Nycteris aethiopica in Ethiopia.

49 268 M. J. LARGEN, D. KOCK and D. W. YALDEN e Lavio frons Cordioderma cor Fig The recorded distribution of Lavia frons and Cardioderma cor in Ethiopia.

50 MAMMALS OF ETHIOPIA. I. CHIROPTERA 269 e Rhinolophus clivosus ocrotis.r. clivosus? andersoni.& R. blasii andreinii Fig The recorded distribution of Rhinolophus clivosus acrotis, R. clivosus? andersoni and R. blasii andreinii in Ethiopia. 19

51 270 M. J. LARGEN, D. KOCK and D. W. YALDEN e Rhinolophus landeri dobsoni R. simulator R. hipposideros minimus Fig The recorded distribution of Rhinolophus landeri dobsoni, R. simulator and R. hipposideros minimus in Ethiopia.

52 MAMMALS OF ETHIOPIA. I. CHIROPTERA 271 e Rhinolophus fumigatus R. hildebrondti Fig The recorded distribution of Rhinolophus fumigatus and R. hildebrandti in Ethiopia.

53 272 M. J. LARGEN, D. KOCK and D. W. YALDEN e Hipposideros coffer coffer H. ruber centrolis Fig The recorded distribution of Hipposideros caffer caffer and H. ruber centra/is in Ethiopia.

54 MAMMALS OF ETHIOPIA. I. CHIROPTERA 273 Hipposideros commersoni marungensis Hipposideros megalotis e Trioenops persicus afer Y Hipposideros fuliginosus Fig The recorded distribution of Hipposideros commersoni marungensis, H. mega/otis, Triaenops persicus afer and H. fuliginosus in Ethiopia.

55 274 M. J. LARGEN, D. KOCK and D. W. YALDEN e. Asellio tridens tridens 6 Asellio potrizii Fig The recorded distribution of Asellia tridens tridens and A. patrizii in Ethiopia.

56 MAMMALS OF ETHIOPIA. I. CHIROPTERA 275 e Eptesicus somalicus somalicus,& E. capensis E. guineensis Fig The recorded distribution of Eptesicus somalicus somalicus, E. capensis and E. guineensis in Ethiopia.

57 276 M. J. LARGEN, D. KOCK and D. W. YALDEN e Pipistrellus nanus Fig The recorded distribution of Pipistrellus nanus in Ethiopia.

58 MAMMALS OF ETHIOPIA. I. CHIROPTERA 277 e Pipistrellus kuhli fuscatus A P. rusticus marrensis P. rueppell i rueppelli 9 Mimetillus moloneyi Fig The recorded distribution of Pipistrellus kuhli fuscatus, P. rusticus marrensis, P. rueppelli rueppelli and Mimetillus moloneyi in Ethiopia.

59 278 M. J. LARGEN, D. KOCK and D. W. YALDEN e Glouconycteris variegate Plecotus austriacus Barbastella barbastellus leucomelas 9 Loephotis wintoni Fig The recorded distribution of Glauconycteris variegata, Plecotus austriacus, Barbastella barbastellus leucomelas and Laephotis wintoni in Ethiopia.

60 MAMMALS OF ETHIOPIA. I. CHIROPTERA 279 e Miniopterus schreibersi. arena nus 6 M. inflatus african us v a 0 ~ 0 Fig The recorded distribution of Miniopterus schreibersi arenarius and }Vf. inflatus africanus in Ethiopia.

61 280 M. J. LARGEN, D. KOCK and D. W. YALDEN (J -~ e Nycticeius schlieffeni schlieffeni.a. N. hirundo N. hindei Fig The recorded distribution of Nycticeius schlielfeni schlielfeni, N. hirundo and N. hindei in Ethiopia.

62 MAMMALS OF ETHIOPIA. I. CHIROPTERA 281 Myotis tricolor.. M. scotti M. morrisi M. welwitschi Fig The recorded distribution of Myotis tricolor, M. scotti, M. morrisi and M. welwitschi in Ethiopia.

63 282 M. J. LARGEN, D. KOCK and D. W. YALDEN e Scotophilus nigrito. flavigaster A S.leucogaster Fig The recorded distribution of Scotophilus nigrita flavigaster and S. leucogaster in Ethiopia.

64 MAMMALS OF ETHIOPIA. I. CHIROPTERA Kerivoula eriophora K. harrisoni 9 Otomops. martiensseni martiensseni e Plotymops setiger moc.milloni (j 60 0 Fig The recorded distribution of Kerivoula eriophora, K. harrisoni, Otomops martiensseni martiensseni and Platymops setiger macmillani in Ethiopia.

65 284 M. J. LARGEN, D. KOCK and D. W. YALDEN Tadarida? chapini e T. pumila pumila. T. bivittata Fig The recorded distribution of T adarida? chapini, T. pumila pumila and T. bivittata in Ethiopia.

66 MAMMALS OF ETHIOPIA. I. CHIROPTERA 285 e Tadarida nigeriae nigeriae A Tadarido midas midas Tadarida condylura Fig The recorded distribution of Tadarida nigeriae nigeriae, T. midas midas and T. condylura in Ethiopia. 20

67 286 M. J. LARGEN, D. KOCK and D. W. YALDEN e Todorida ansorgei A T. ventralis T. oegyptioco T. nonulo Fig The recorded distribution of Tadarida ansorgei, T. ventralis, T. aegyptiaca and T. nanula in Ethiopia.

68 MAMMALS OF ETHIOPIA. I. CHIROPTERA 287 VI. GAZETEER OF COLLECTION LOCALITIES The co-ordinates cited in the following gazeteer are based mainly upon the 1: and 1: maps, series GSGS 2465 and 4646, published by the British War Office during the years This series, to a great extent compiled from earlier Italian surveys, is known to include many inaccuracies, but these are the only large-scale maps of Ethiopia at present available and their age is often an advantage when attempting to identify the older place-names mentioned in the zoological literature. A number of older maps and expedition journals have also been consulted, and the gazeteers provided by MoHR (1962) and DAVIS & MISONNE (1964) have been found useful, although the latter, almost inevitably, contains a number of major errors. There remain certain records which it has not been possible to trace; either because a village has subsequently disappeared or because its name has since been changed or, in some cases, because a collector may have used local information which has never appeared on a map. It is possible, however, that some of these localities may later be identified when work on the literature of other mammalian orders is completed. Latitude and longitude are given as accurately as possible for towns and villages. Unless more precise information is available, co-ordinates are given for the approximate centre of lakes, mountain ranges and plains; and for rivers the mouth, or place where the Ethiopian border is crossed, is pin-pointed. Heights in italics are those provided by collectors or for other reasons considered to be relatively accurate; the remaining references to altitude are approximate only. Locality Abaya, Lake Abaya, Lake, N shore Abaya, Lake, NE of Abaya, Lake, W shore Abaya, Lake, SW corner Abd el Kadr Abella Abiata, Lake, E shore Adamatuli Addis Ababa Addis Ababa, 70 miles SW of Addis Alem Addi Sciaddi Co-ordinates 06 15'N 37 55'E 06 36'N 37 56'E 06 30'N 38 05'E 06 13'N 37 49'E 06 04'N 37 45'E 15 38'N 39 28'E 06 42'N 3r56'E 07 37'N 38 39'E 07 51'N 38 42'E 09 02'N 38 45'E see Woliso 09 03'N 38 23'E 1Y39'N 38 38'E Altitude ( m) sea level

69 288 M. J. LARGEN, D. KOCK and D. W. YALDEN Locality Co-ordinates Altitude ( m) Adi Caie 14 51'N 39 23'E 2400 Adi Guhebo near Adi Ugri Adi Ugri 14 53'N 38 49'E 1900 Adowa 14 10'N 38 54'E 1900 Adur (? Aduri Mariam ll 0 25'N 36 53'E) 1800 Agaro, 10 km from ca 07 50'N 36 40'E 1650 Agordat 15 33'N 3J053'E 600 Alemaya 09 26'N 'E 2000 Alghe, 30 miles W of ca 05 30'N 38 00'E Amar Cocche 05 04'N 36 47'E 2000 Ambo 08 57'N 37 57'E 2140 Anseba (River) valley l7 00'N 37 24'E Arba Minch 06 03'N 37 38'E 1400 Arba Mondi 06 00'N 37 45'E 1300 Asam River see Adowa As mara 15 20'N 38 55'E 2300 Asmara, 6 miles W of 15 22'N 39 00'E 2000 Assab 13 01'N 42 43'E sea level Atirba River (? Ataba River 13 38'N 38 39'E) Auramalka see Aware Melca Aware Melca 09 08'N 39 58'E 900 Awasa, Lake, E shore 07 05'N 38 27'E 1700 Awash Falls 08 50'N 40 01'E 960 Awash National Park ca 08 54'N 39 55'E 1000 Awash River, N bank 08 52'N 40 01'E 960 Azir River 10 29'N 36 25'E 1000 Baa-sa-ka, Lake 08 52'N 39 52'E 1000 Backo, W ollega 09 07'N 37 03'E 1750 Badditu ca 06 00'N 38 00'E Bahadu 10 06'N 40 36'E 600 Bahar Dar ll 0 35'N 37 25'E 1850 Bahr der Giorgis see Bahar Dar Barca River l7 19'N 37 31'E Barentu 15 09'N 37 38'E 980 Barye (? Barri 09 02'N 36 12'E 1400) Belegaz (River) valley ca 12 50'N 38 20'E Beleka 09 33'N 41 37'E 1455 Beletta Forest 07 33'N 36 34'E 2200 Bilo 08 53'N 37 00'E 1700 Bishoftu (Debre Zeit) 08 44'N 38 59'E 2000 Bogos (region) 15 46'N 38 30'E

70 MAMMALS OF ETHIOPIA. I. CHIROPTERA 289 Locality Co-ordinates Altitude (m) Buia see Assab Bulcha Forest 06 1l'N 38 10'E 1800 Caschei 04 48'N 36 27'E 900 Chamo, Lake, W shore 05 50'N 37 32'E 1200 Chamo, Lake, NE of 05 58'N 37 55'E 1500 Cheren see Keren Coatit 14 49'N 39 13'E 1900 Dacata valley 09 08'N 42 25'E 1300 Daimbi Shoa Dan gila 11 17'N 36 55'E 2000 Danka River, near Dinshu 07 00'N 39 45'E 3300 Dass Wells 04 05'N 38 45'E 1500 Debaroa 15 06'N 38 51'E 2000 Debre Marcos 10 21'N 37 43'E 2500 Dech Island l1 54'N 37 17'E 1900 Dhar-ala (? Dabala l1 57'N 40 08'E 900) Didessa River mouth 10 05'N 35 38'E 900 Didessa River bridge 09 02'N 36 09'E 1200 Dime 06 18'N 36 16'E Din shu 07 06'N 39 47'E 3200 Dire Dawa 09 35'N 41 52'E 1200 Dire Dawa, 4 km S of 09 32'N 41 53'E 1250 Djala 06 20'N 36 50'E 2000 Djemdjem Forest 09 00'N 38 12'E 2500 Djibouti l1 35'N 43 09'E sea level Djiren 07 42'N 36 53'E 1900 Doha River 09 42'N 40 50'E 900 Dobleh ca OJGOO'N 45 00'E (collector's reference) Dolo, Somalia 04 ll'n 42 05'E 200 Donkam River 10 40'N 36 32'E 1500 Dorfu 15 10'N 39 03'E 2000 Dungulbar l1 58'N 37 01'E 1850 Elolo, Kenya 04 23'N 36 14'E 400 Entedebir Island 15 43'N 39 54'E sea level Erer Gota 09 31'N 41 2l'E 1200 Errer valley 09 05'N 42 17'E 1200 Errer valley at Harrar ca 09 15'N 42 10'E 1500 Errer valley, bridge on Jigjiga road 09 15'N 42 16'E 1500 Errer valley, 5 km E of bridge on Jigjiga road 09 14'N 42 17'E 1500

71 290 M. J. LARGEN, D. KOCK and D. W. YALDEN Locality Co-ordinates Altitude ( m) F afan valley 09 1l'N 42 38'E 1500 Fantalle, Mount, NW slope 08 58'N 39 54'E 1000 Fantalle, Mount, W slope 08 54'N 39 56'E 1000 Farre (? Fare 09 38'N 39 53'E 2000) Fatam River 10 25'N 37 00'E 1800 Filwoha 09 00'N 39 58'E 790 Fincha River 10 03'N 3J020'E 1000 Gaarre 13 13'N 42 07' 150 Gabba River 08 15'N 36 10'E 2000 Gara Hakim 09 17'N 42 07'E 2060 Gara Mullata 09 16'N 41 44'E 2700 Gallabat, Sudan 12 57'N 36 09'E 770 Gam bela 08 15'N 34 35'E 515 Ganale River 04 10'N 42 05'E Garibaldi Pass 08 48'N 39 42'E 1200 Geddem see Jebel Geddem Gerlogobi 06 52'N 45 02'E 700 Ghibie River bridge 08 14'N 3J035'E 1400 Ghimbi 09 10'N 35 50'E 2150 Ghinda 15 12'N 39 03'E 1000 Ghinir, 7-10 km from ca 07 10'N 40 42'E 1400 Ghirghiret 15 37'N 39 03'E 500 Gidole, 70 km SW of 05 20'N 37 10'E 1000 Goba, 3 miles S of 06 58'N 39 59'E 3000 Godolefassi 14 53'N 38 51'E 1900 Gofa (district) 06 15'N 36 40'E Gojeb River 07 30'N 36 25'E 1500 Gold mine of Uallega (Yubdo) 08 52'N 35 26'E 1500 Gololcha 08 10'N 40 04'E 1500 Gondar l2 37'N 37 27'E 2200 Gondar, 15 miles NE of 12 45'N 37 42'E 3000 Gondaraba 04 58'N 36 48'E 450 Gorgora l2 15'N 37 20'E 2000 Gota River 09 40'N 41 20' 1100 Gura 15 01'N 39 04'E 2000 Gurairobi See (the former Lake Arairobi, now obliterated by Koka reservoir) 08 19'N 39 05'E 1700 Hamar Koke see Amar Cocche Hara Digallu (? Digalu 04 42'N 38 23'E) 1400 Harairobi, Lake see Gurairobi See

72 MAMMALS OF ETHIOPIA. I. CHIROPTERA 291 Locality Co-ordinates Altitude (m) Harar 09 18'N 42 08'E 2000 Hertale, Lake 09 57'N 40 27'E 600 Hirna, near ca 09 12'N 41 01'E 2520 Hoddo, 22 km W of 04 43'N 36 02'E 400 Hop Saloa mine not located, probably Eritrea Jebel Geddem 15 20'N 39 32'E 30 Jjgjiga road, 15 km SE of see Dacata valley Jimma 07 40'N 36 50'E 1800 Jiffa Medir, Somalia 09 45'N 43 15'E 1500 Keren 15 46'N 38 27'E 1400 Kibbish Wells, Sudan 05 14'N 35 51'E 400 Koffole 07 05'N 38 47'E 2500 Koka 08 27'N 39 06'E 1700 Langano, Lake 07 35'N 38 45'E 1600 Langano, Lake, W shore 07 37'N 38 42'E 1600 Let Marefia 09 39'N 39 46'E 3000 Little Abbai see Piccolo Abbai Mabil 10 20'N 36 45'E 1200 Mabil, 10 km W of 10 19'N 36 45'E 1000 Maddo Erelle Wells, Kenya 03 54'N 41 05'E 500 Mai Atal 15 32'N 39 13'E 1300 Manera Forest 06 54'N 36 42'E 1220 Margharita, Lake see Abaya, Lake Massawa 15 35'N 39 26'E sea level Mega, near ca 04 05'N 38 19'E 1200 Mega, E of 04 05'N 38 19'E 1800 Melkay Manna 06 45'N 40 45'E 1500 Meta Abo 08 55'N 'E 2500 Metahara 08 54'N 39 55'E 1000 Metahara, W of 08 53'N 39 55'E 1000 Metemma 12 57'N 36 10'E 770 Monkullo 15 32'N 39 26'E sea level Mui 05 50'N 35 45'E 700 Mulu 09 20'N 38 41'E 2500 Murle 05 10'N 36 13'E 400 Neghelli 05 20'N 39 35'E 1500 Neghelli, 54 km S of 05 00'N 39 28'E 1000 Nocra Island 15 45'N 39 57'E sea level Nora Island 16 02'N 40 03'E sea level Nur Mohammed 06 47'N 40 22'E 2600 Odar Eritrea

73 292 M. J. LARGEN, D. KOCK and D. W. YALDEN Locality Co-ordinates Altitude ( m) Om Ager 14 18'N 36 38'E 500 Piccolo Abbai 11 32'N 37 03'E 1900 Pokwo 08 15'N 34 25'E 500 Portuguese bridge, 30 km SE of 11 10'N 38 10'E 1300 Robi, Shoa 10 15'N 40 00'E 1200 Rorum Plains 13 25'N 40 45'E ± sea level Rudolf, Lake 04 37'N 36 08'E Saaita see Saati Saati 15 35'N 39 16'E 200 Sa beta 08 55'N 38 40'E 2500 Sagan River 04 51'N 35 56'E Saganeiti 1Y04'N 39 12'E 2200 Sanford's Ranch see Mulu Sancurrar, Kenya 04 02'N 40 08'E 1100 Sassabane 07 53'N 43 40'E 900 Sembel 15 15'N 38 53'E 2300 Senafe 14 42'N 39 25'E 2450 Serae 15 00'N 38 30'E 1500 Setit (River) 14 15'N 36 34'E Shalla, Lake, E shore 07 27'N 38 38'E 1600 Shashamane, 11 km E of 07 10'N 38 40'E 2230 Sheik Hussein OJ036'N 40 40'E 1500 Sia'ha between Saganeiti and Godolefassi Sidamo (Province), 140 miles S of Addis Ababa 07 00'N 38 30'E 1800 Simien Mountains 13 15'N 38 20'E Sirba 10 05'N 35 30'E 800 So Omar 06 54'N 40 48'E 1340 Solomona Habesh ( Salomona) ca 1Y42'N 38 50'E 800 Stephanie, Lake, N of 04 55'N 36 55'E 500 Stephanie, Lake, NW shore 04 50'N 36 45'E 500 Suha River 10 08'N 38 20'E 2500 Tana, Lake 12 00'N 3J020'E 1830 Tana, Lake, 100 miles S of see Fatam River Torat (region) ca 13 10'N 37 15'E Tserea Wukro 13 49'N 39 37'E 2030 Ueb River, near Dinshu 07 06'N 39 46'E 3000 Ursa River 09 38'N 41 38'E 1000 Wadere 06 59'N 45 20'E 600 Wahina, Amhara (? Uahini 11 35'N 39 15'E 2500) Wiah 15 18'N 39 31'E 700

74 MAMMALS OF ETHIOPIA. I. CHIROPTERA 293 Locality Co-ordinates Altitude (m) Woliso 08 32'N 37 59'E 2000 Wongabo (region) ca 15 00'N 39 52'E 200 Yavello, 25 km N of 05 03'N 38 12'E 2000 Zeban Zighib 14 51'N 39 22'E 2400 Zeghie l1 42'N 37 21'E 1900 Zeriba Ambarab 15 10'N 36 55'E 800 Zwai, Lake, 1 km S of } 07 55'N 38 43'E 1600 Zwai village, 7 km S of SUMMARY The present work summarises the known distribution of Chiroptera in Ethiopia. Seventy-two species are listed, of which 12 represent new records for the country: Rousettus angolensis (Bocage, 1898), Taphozous mauritianus E. Geoffroy, 1818, Nycteris hispida hispida (Schreber, 1775), Hipposideros fuliginosus (Temminck, 1853), Eptesicus guineensis (Bocage, 1889), Mimetillus moloneyi (Thomas, 1891), Nycticeius hindei (Thomas, 1901), Nycticeius hirundo (De Winton, 1899), Scotophilus leucogaster (Cretzschmar, 1830), Otomops martiensseni martiensseni (Matschie, 1897), Tadarida condylura (A. Smith, 1833), Tadarida nanula (J. A. Allen, 1917). As far as possible, all collecting localities in Ethiopia have been gazetted and this record is supplemented with distribution maps for every species. Sixty-two of the species recorded from Ethiopia are essentially African in distribution, six of the remainder are Palaearctic forms and four are associated with the arid Saharo-Sindian belt. Only five species appear to be represented by more than one race and only four are potentially endemic to Ethiopia: Asellia patrizii De Beaux, 1931, Myotis scotti Thomas, 1927, Myotis morrisi Hill, 1971, Kerivoula eriophora (Heuglin, 1877). These figures low compared with those for other groups of small mammals and are, presumably, correlated with the comparative mobility of these flying animals. RIASSUNTO La chirotterofauna dell'etiopia annovera attualmente 72 specie, 12 delle quali sono qui ricordate per Ia prima volta come presenti nella regione considerata: Rousettus angolensis (Bocage, 1898 ), Taphozous mauritianus E. Geoffroy, 1818, Nycteris hispida hispida (Schreber, 1775), Hipposideros fuliginosus (Temminck, 1853 ), Eptesicus guineensis (Bocage,

75 294 M. J. LARGEN, D. KOCK and D. W. YALDEN 1889), Mimetillus moloneyi (Thomas, 1891 ), Nycticeius hindei (Thomas, 1901), Nycticeius hirundo (De Winton, 1899), Scotophylus leucogaster (Cretzschmar, 1830), Otomops martiensseni martiensseni (Matschie, 1897), Tadarida condylura (A. Smith, 1833), Tadarida nanula (].A. Allen, 1917). Gli Autori forniscono un elenco il pili possibile completo delle localita, con loro posizione geogra6ca, in cui sono stati sinora raccolti pipistrelli e forniscono una carta di distribuzione di tutte le specie note dell'etiopia. Di tali specie, 62 hanna una distribuzione essenzialmente africana e 10 sono paleartiche; di quest'ultime 4 sono proprie della fascia saharosindica. Solo 5 specie sono rappresentate nel territorio da piu di una razza e solo 4 sono potenzialmente endemiche dell'etiopia: Asellia patrizii De Beaux, 1931, Myotis scotti Thomas, 1927, Myotis morrisi Hill, 1971, Kerivoula eriophora (Heuglin, 1877). II tasso d'endemismo e basso rispetto a quello riscontrabile in altri ordini della stessa classe e presumibilmente cio e da mettere in relazione con Ia vagilita di questa ordine di mammiferi. REFERENCES ANDERSEN, K On some new or little-known bats of the genus Rhinolophus in the collection of the Museo Civico, Genoa. Annali Mus. civ. Stor. nat. Giacomo Doria (3a) 2: ANDERSEN, K. 1907a. On Pterocyon, Rousettus and Myonycteris. Ann. Mag. nat. Hist. (7) 19: ANDERSEN, K. 1907b. Chiropteran notes. Annali Mus. civ. Stor. nat. Giacomo Doria (3a) 3: ANDERSEN, K. 1912a. Catalogue of the Chiroptera in the collection of the British Museum. Vol. 1 (2nd Edition). Megachiroptera. London: British Museum (N.H.) - Cl+854 pp. ANDERSEN, K. 1912b. Brief diagnoses of eight new Petalia, with a list of the known forms of the genus. Ann. Mag. nat. Hist. (8) 10: ANDERSON, J. & W. E. DE WINTON Zoology of Egypt: Mammalia. Revised and completed by W. E. De Winton. London: Hugh Rees, Ltd. XVII+374 pp. ANDRAL, L., P. BRES, C. SERrE, J. CASALS & R. PANTHIER Etudes sur la fievre jaune en Ethiopie. 3. Etude serologique et virologique de la faune sylvatique. Bull. Wid. Hlth. Org. 38: BEADLES, J. K. & R. H. INGERSOL An annotated check-list of the mammal fauna of the Chercher Highlands of Ethiopia present in the museum at the College of Agriculture, HSIU. Experiment. Stat. Bull. No. 51. Haile Selassie I University, College of Agriculture, Dire Dawa. BRAESTRUP, F. W On the taxonomic value of the subgenus Lophomops (nyctinomine bats), with remarks on the breeding times of African bats. Ann. Mag. nat. Hist. (10) 11: CARCASSON, R. H A preliminary survey of the zoogeography of African butterflies. E. Afr. Wildl. J. 2: CARPENTER, G. D. H The Rhopalocera of Abyssinia; a faunistic study. Trans. R. ent. Soc. Lond. 83:

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78 MAMMALS OF ETHIOPIA. I. CHIROPTERA 297 INGERSOL, R. H The ecological stratification of mammals in the eastern Chercher Highlands of Harar Province, Ethiopia. Ph.D. thesis, Oklahoma State University. JABLONOWSKI, J Die li:iffelartigen Haare der Molossi. Abh. Ber. K. zool. anthrop.-ethn. Mus. Dresden 7 (7): }ENTINK, F. A Catalogue systematique des mammiferes (Rongeurs, insectivores, cheiropteres, edentees et marsupiaux). Vol. 12. Leiden: Mus. Hist. nat. Pays-Bas. 280 pp. }OBLING, B On the African Streblidae (Diptera Acalypterae) including the morphology of the genus Ascodipteron Adens and a description of a new species. Parasitology 31 (2): KERSHAW, P. S On two collections of mammals: (1) from Mongalla and Bahr-el-Ghazal, (2) from Lake Tsana, Abyssinia. Ann. Mag. nat. Hist. (9) 13: KLATT, B Bericht iiber eine Reise nach Eritrea im Friihjahr Sber. Ges. naturf. Freunde Berl KocK, D Die Fledermaus-Fauna des Sudan. Abh. senckenb. naturforsch. Ges. 521: KocK, D Merkmalsdifferenzen tropischer Fledermause. Z. Siiugetierk. 38 (3 ): LANZA, B Parco Nazionale dell'auasc (Scioa, Etiopia). L'Universo, Firenze 52 (5): MAGRETTI, P Nel Sudan orientale. Atti Soc. ital. Sci. nat. 27: MoHR, P. A The Geology of Ethiopia. University College of Addis Ababa Press. 268 pp. MoNTICELLI, F. S Note chirotterologiche. Annali Mus. civ. Stor. nat. Giacomo Doria (2) 5: MoNOD, TH. & W. H. MoRTON Contribution a l'etude de la grotte de So Omar (Ethiopie meridionale). 1. Introduction. Annls Speleol. 27 (1): OusTALET, E Les mammiferes et les oiseaux d'obock et du pays <;:omalis. Mem. Soc. zool. Fr. 7 (1): PETERS, W. C. H Ober einige neue oder weniger bekannte Flederthiere. Mber. K. preuss. Akad. Wiss. Berlin: PETERS, W. C. H Eine monographische Obersicht der Chiropterengattungen Nycteris und Atalapha. Mber. K. preuss. Akad. Wiss. Berlin (for 1870): PETTER, F Deux rongeurs nouveaux d'ethiopie: Stenocephalemys griseicauda sp. nov. et Lophuromys melanonyx sp. nov. Mammalia, 36 (2): POscHER, H Ober die Schultertaschen von Epomophorus (Epomophorini, Pteropodidae, Megachiroptera, Mammalia). Z. Siiugetierk. 37 (3): RADFORD, C. D A new larval mite from Eritrea (Acarina: Trombiculidae). Fieldiana, Zool. 34 (29): RoDE, P Les mammiferes de la mission Dakar-Djibouti (Mission Griaule). Bull. Mus. natn. Hist. nat. Paris (2) 5 (5): ROPPELL, E Verzeichnis der in dem Museum der Senckenbergischen naturforschenden Gesellschaft aufgestellten Sammlungen. Erste Abtheilung: Siiugethiere und deren Skelette. Mus. Senckenberg. 3: SANBORN, C. C Descriptions and records of African bats. Zool. Ser. Field Mus. nat. Hist. 20 (17): SENNA, A Contributo alla conoscenza dei chirotteri Eritrei. Archo zool. ital. 2 (3 ): SMrT, F. G. A. M The African species of the bat-flea Lagaropsylla. Revue Zool. Bot. afr. 55 (1-2):