Pantala flavescens at the coast of the Baltic Sea (Odonata: Libellulidae)

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1 Pantala flavescens at the Baltic Sea st June 043 Pantala flavescens at the coast of the Baltic Sea (Odonata: Libellulidae) Paweł Buczyński, Anatoly P. Shapoval & Edyta Buczyńska 3 Department of Zoology, Maria Curie-Skłodowska University, Akademicka Str. 9, Lublin, Poland; <pawbucz@gmail.com> Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia; <apshap@mail.ru> 3 Department of Zoology, Animal Ecology and Wildlife Management, University of Life Sciences, Akademicka Str. 3, Lublin, Poland; <edyta.buczynska@gmail.com> Received 0 th September 03; revised and accepted st December 03 Abstract. A male Pantala flavescens was recorded in a bird net trap on the Courish Spit, Kaliningrad Oblast, western Russia (55 05 N, 0 44 E) on 9-v-03. This is the northernmost record of this species in Europe and in the whole northern hemisphere. The record is discussed against the background of European records of P. flavescens. Key words. Dragonfly, Anisoptera, migration, Courish Spit, Kaliningrad, Russia, Europe Odonatologica 43(/) 04: 3-

2 Odonata from the Tibesti Mountains st June 04 3 Odonata from the Tibesti Mountains and the Ounianga Lakes in Chad, with notes on Hemianax ephippiger accumulating in the desert Henri J. Dumont Dept. of Hydrobiology and Ecology, Jinan University, Guangzhou, China, and Department of Biology, Gent University, Belgium; <hjdumont@ugent.be> Received 30 th March 04; revised and accepted 4 th April 04 Abstract. Fourteen species of Odonata were collected in Ounianga and Tibesti (Chad, Africa) in March 04. Among them, only one zygopteran (Ischnura saharensis, with a Saharan distribution), one gomphid (the eremian Paragomphus sinaiticus), and two aeshnids (Anax imperator and Hemianax ephippiger) were present. The latter species was not only the most common dragonfly, but also the most abundant insect seen in the desert. It is likely that it was in a phase of accumulating individuals, possibly as a prelude to another massive trans-sahara and even trans-mediterranean migration, for which the species is well-known. The 0 libellulids recorded were almost all Afrotropical species, but several expand to the Maghreb and even Mediterranean Europe. Only Orthetrum cf. hintzi is a tropical African species that had never been recorded from the desert before. Key words. Dragonfly, Africa, migration, Orthetrum cf. hintzi, Sahara Odonatologica 43(/) 04: 3-4

3 Rediscovery of Cordulegaster vanbrinkae in Iran st June 04 5 Rediscovery of Cordulegaster vanbrinkae in Iran (Odonata: Cordulegastridae) Thomas Schneider, Elias Schneider, Jacob Schneider & Ole Müller Arnold-Knoblauch-Ring 76, 409 Berlin, Germany; <thomas.rs@gmx.de> Birkenweg 6d, 5306 Libbenichen, Germany; <olemueller@bioscience-art.de> Received 30 th September 03; revised and accepted 3 th March 04 Abstract. In July 03, a total of 4 males of Cordulegaster vanbrinkae was observed in the Hyrcanian Forest, Alborz Mountains, north-western Iran, not far from the type locality. This is only the second record of this poorly known species from Iran. Seven male specimens were collected. The variation of abdominal colour patterns and other morphological characters are shown. Notes on the biology of this species and a description of the biotope of a recently discovered population in Armenia are given. Key words. Dragonfly, Anisoptera, morphological variation, male behaviour, Hyrcanian Forest, Alborz Mountains Odonatologica 43(/) 04: 5-34

4 Two new Heteropodagrion species from Ecuador st June Two new Heteropodagrion species from Ecuador (Odonata: Megapodagrionidae) Jerrell J. Daigle 067 Little River Lane, Tallahassee, Florida 33, U.S.A.; Received th February 04; revised and accepted 7 th April 04 Abstract. Heteropodagrion nigripes sp. nov. is described and illustrated (holotype : Ecuador, Morona Santiago Province, 07-xi-997). It can be distinguished by the black legs and stout white cercus. Heteropodagrion varipes sp. nov. is described and illustrated (holotype : Ecuador, Morona Santiago Province, 8-ix-005). The bicolored legs and middorsal cercus tooth separates it from other Heteropodagrion species. Both holotypes are deposited in the Florida State Collection of Arthropods, Gainesville, Florida, USA. Key words. Dragonfly, damselfly, Zygoptera, new species, Heteropodagrion nigripes sp. nov., Heteropodagrion varipes sp. nov., South America Odonatologica 43(/) 04: 35-4

5 Anisopleura pelecyphora sp. nov. from Yunnan, China st June Anisopleura pelecyphora sp. nov. from south-western Yunnan, China (Odonata: Euphaeidae) Hao-miao Zhang, Matti Hämäläinen & Qing-hua Cai State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 43007, China; Naturalis Biodiversity Center, P.O. Box 957, 300 RA, Leiden, The Netherlands; Received nd February 04; revised and accepted 4 th February 04 Abstract. A new euphaeid damselfly species, Anisopleura pelecyphora sp. nov. (holotype from Ximeng Wa, Yunnan, China) is described and illustrated for the male sex. A comparison with its congeners is provided. Key words. Dragonfly, damselfly, Zygoptera, new species Odonatologica 43(/) 04: 43-50

6 The status of Psolodesmus mandarinus st June 04 5 A revision of the status of Psolodesmus mandarinus based on molecular and morphological evidence (Odonata: Calopterygidae) Sue-cheng Lin, Young-fa Chen, Sen-her Shieh 3 & Ping-shih Yang 4 Division of Zoology, Endemic Species Research Institute, Chichi, Nantou 55, Taiwan Experimental Forest, National Taiwan University, Taipei 06, Taiwan 3 Department of Ecological Humanities, Providence University, Salu, Taichung 433, Taiwan 4 Department of Entomology, National Taiwan University, Taipei 06, Taiwan; <psyang@ntu.edu.tw> Received nd November 03; revised and accepted 3 th March 04 Abstract. To investigate the relationships between the three recognized taxa in the genus Psolodesmus, traditionally ranked as subspecies of Psolodesmus mandarinus: mandarinus, dorothea and kuroiwae, the nuclear internal transcribed spacers and ribosomal 5.8S gene, mitochondrial cytochrome c oxidase subunit I gene, and wing pterostigma data were analyzed. Both molecular and morphological evidence suggest the presence of two distinct species, viz. P. mandarinus in Taiwan and P. kuroiwae in the Japanese Yaeyama Islands. Based on our results we continue the traditional practice of dividing the Taiwanese populations of P. mandarinus into two geographical subspecies, P. m. mandarinus in northern Taiwan and P. m. dorothea in central and southern Taiwan. Key words. Dragonfly, damselfly, Zygoptera, Psolodesmus kuroiwae, P. mandarinus dorothea, nuclear DNA, mitochondrial DNA, pterostigma, taxonomy Odonatologica 43(/) 04: 5-66

7 Amphicnemis triplex sp. nov. from Indonesia st June Amphicnemis triplex sp. nov. from Central Kalimantan, Indonesia (Odonata: Coenagrionidae) Rory A. Dow Naturalis Biodiversity Centre, P.O. Box 957, 300 RA Leiden, The Netherlands; Received 7 th September 03; revised and accepted 0 th February 04 Abstract. Amphicnemis triplex sp. nov. is described from locations in Central Kalimantan in Indonesian Borneo. Holotype Indonesia, Kalimantan, Kalimantan Tengah, between Buntok and Ampah, black water stream in shallow peat over sand, 9-vi-0; to be deposited in RMNH. The related species A. erminea is discussed. Key words. Dragonfly, damselfly, Zygoptera, new species, Borneo Odonatologica 43(/) 04: 67-77

8 Indocypha neglecta sp. nov. from Vietnam st June Indocypha neglecta sp. nov. from northern Vietnam (Odonata: Chlorocyphidae) Matti Hämäläinen Naturalis Biodiversity Center, P.O. Box 957, 300 RA Leiden, The Netherlands; Received 8 th January 04; revised and accepted 0 th February 04 Abstract. Indocypha neglecta sp. nov. is described and illustrated from both sexes and compared with its congeners. The holotype was collected at»tonkin, Montagnes du Haut Song-Chai«in northern Vietnam in 895 and is deposited at MNHN, Paris. Key words. Dragonfly, damselfly, Zygoptera, new species, René Martin Odonatologica 43(/) 04: 79-90

9 Drepanosticta elaphos sp. nov. and D. pterophora sp. nov. from Papua New Guinea st June 04 9 Drepanosticta elaphos sp. nov. and Drepanosticta pterophora sp. nov. from Papua New Guinea (Odonata: Platystictidae) Günther THeischinger & Stephen J. Richards Office of Environment and Heritage New South Wales, Sydney, NSW, Australia, and Australian Museum, Entomology, 6 College Street, Sydney, NSW, 00, Australia; <gunther.theischinger@environment.nsw.gov.au> Museum and Art Gallery of the Northern Territory, current address: Herpetology Department, South Australian Museum, North Terrace, Adelaide, S.A Australia; <steve.richards@samuseum.sa.gov.au> Received 9 th May 03; Reviewed and Accepted 30 th January 04 Abstract. Two species of Drepanosticta Laidlaw from Papua New Guinea are described as new. They are: Drepanosticta elaphos sp. nov. (holotype : Papua New Guinea) and D. pterophora sp. nov. (holotype : Papua New Guinea), both from the upper Sepik Basin. Characters of the adults are illustrated, habitat notes are provided and their affinities are discussed. Key words. Dragonfly, damselfly, Zygoptera, new species, Sepik Basin Odonatologica 43(/) 04: 9-03

10 Colour variation of female Antipodochlora braueri st June An undescribed colour variation of female Antipodochlora braueri (Odonata: Corduliidae) Milen Marinov Plant Health & Environment Laboratory, Investigation and Diagnostic Centres and Response, Ministry for Primary Industries, 4 Sir William Pickering Drive, Burnside, PO Box 408, Christchurch, New Zealand; <milen.marinov@mpi.govt.nz> Received st September 03; revised and accepted 5 th February 04 Abstract. Four Antipodochlora braueri (Selys, 87) specimens (, ) were collected in the North Island of New Zealand. One female from the Taranaki region had a wing colouration previously unknown. The new colour variation is described here and is compared to what had been so far reported of female A. braueri. Important diagnostic features, such as appendages and vulvar scales, are illustrated. The colour variant female is also compared to the female of Procordulia smithii because of the similarity of the wing pattern, which had caused confusion in the past. A male collected together with the female had wings more tinged with yellow than other known males. Key words. Dragonfly, Anisoptera, wing colouration, New Zealand, Procordulia smithii Odonatologica 43(/) 04: 05-4

11 Counter-adaptation in response to sperm removal in Ischnura asiatica st June 04 5 Counter-adaptation in response to sperm removal by stimulating the sensory system in female Ischnura asiatica (Odonata: Coenagrionidae) Yusuke Tajima & Mamoru Watanabe Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki , Japan; corresponding author, <watanabe@kankyo.envr.tsukuba.ac.jp> Received 3 th July 03; revised and accepted st January 04 Abstract. The male Ischnura asiatica (Brauer, 865) has a pair of horns on its penis head. Because each horn is shorter than the spermathecal duct, the spermatheca is inaccessible to the male. Thus, males can not directly displace spermathecal sperm using the horns. Nevertheless during copulation displacement of sperm from the spermatheca does occur. By stimulating vaginal sensilla that communicate the presence of an egg to the muscles surrounding the sperm storage organs, sperm ejection can be induced as fertilization is anticipated by the female. Males with a large penis head might be better adapted to stimulate the sensilla and displace more sperm than those with a small penis head. From the viewpoint of females, on the other hand, there are costs if complete sperm displacement occurs. Decreasing the number of sensilla might make it difficult for the male s stimulation to displace spermathecal sperm, especially where a population includes males with a large penis head. To test this hypothesis, the width of the penis head and the number of sensilla were measured in several local populations, with different body sizes, distributed in mainland Japan. The number of sensilla decreased with the width of the penis head. Therefore, a low number of sensilla in the females might be a counter-adaptation against the male sensory stimulation. Key words. Dragonfly, damselfly, Zygoptera, fertilization mechanism, interpopulation variation, male-female coevolution, penis head, sexual conflict Odonatologica 43(/) 04: 5-4

12 Exotic Odonata in Europe st June 04 5 Exotic Odonata in Europe Gerold Laister, Gerhard Lehmann & Andreas Martens 3 Hans-Hofmann-Ring 3, 4470 Enns, Austria; <glaister@aon.at> Haunfeldstraße 4, 6330 Kufstein, Austria; <gerilehmann@hotmail.com> 3 University of Education Karlsruhe, Bismarckstraße 0, 7633 Karlsruhe, Germany; <martens@ph-karlsruhe.de> Received nd February 04; revised and accepted 4 th March 04 Abstract. Between 99 and 0, more than,000 adults of exotic odonate species were recorded from glasshouses of a wholesaler dealer of aquarium plants near Wels, Austria. Twenty-three species could be identified to species level. All species were accidentally introduced as eggs or larvae. The majority are widely distributed and common southeastern Asian species. About 7 taxa are first recorded from Europe bringing the list of exotic Odonata in Europe to 4 taxa. These odonates are mainly introduced via aquarist trade and many species have emerged from home aquaria and glasshouses. Currently, tropical plants for aquarists are mainly imported to Europe from Singapore, Indonesia and Thailand, which suggests that the exotic Odonata originated in those countries. So far, the introduction of exotic Odonata species into Europe is not ecologically relevant because none of the introduced species have become established in the wild. However, this study will improve understanding of the significance of trading connections in establishing exotic species which could become invasive. Key words. Dragonfly, neobiota, check-list, aquaristic trade, incidental introduction, dispersal, invasive species, long-term study Odonatologica 43(/) 04: 5-35

13 36 st June A. Autor 04 Erratum to Odonatologica 4 (4): The true status of Somatochlora taiwana Inoue & Yokota: A genuine species or a synonym of S. dido Needham? (Anisoptera: Corduliidae) K. Inoue, A. Sasamoto & R. Futahashi 3 5-9, Fuminosato 4-chome, Abeno-ku, Osaka , Japan; <ks-inoue@mx.nisiq.net> 53-3, Oh, Tawaramoto-cho, Shiki-gun, Nara Pref , Japan; <akssmt@sea.plala.or.jp> 3 National Institute of Advanced Industrial Science and Technology (AIST), Central 6, Tsukuba, Ibaraki Japan; <ryo-futahashi@aist.go.jp> Received 3 rd January 04; accepted 3 rd January 04 Odonatologica 43(/) 04: 36

Anisopleura pelecyphora sp. nov. from south-western Yunnan, China (Odonata: Euphaeidae)

Anisopleura pelecyphora sp. nov. from Yunnan, China 1 st June 2014 43 Anisopleura pelecyphora sp. nov. from south-western Yunnan, China (Odonata: Euphaeidae) Hao-miao Zhang 1, Matti Hämäläinen 2 & Qing-hua