Amphibians and Reptiles of Honduras

Transcription

1 Amphibians and Reptiles of Honduras Listas Zoológicas Actualizadas UCR McCranie, James R SW 164 Street / Miami, Florida, / USA (Originally Published in 2009, Last Actualization on November 12, 2009) INDEX INTRODUCTION HERPETOFAUNA AMPHIBIA GYMNOPHIONA Caeciliidae CAUDATA Plethodontidae ANURA Bufonidae Centrolenidae Centroleninae Hyalinobatrachinae Hylidae Hylinae Phyllomedusinae Craugastoridae Eleutherodactylidae Strabomantidae Leiuperidae Leptodactylidae Microhylidae Ranidae Rhinophrynidae REPTILIA CROCODYLIA Alligatoridae Crocodylidae SQUAMATA SAURIA Anguidae Corytophanidae Eublepharidae Gekkonidae Gymnophthalmidae Iguanidae Leiocephalidae Phrynosomatidae Polychrotidae Scincidae Teiidae Xantusiidae SERPENTES Anomalepididae Boidae Colubridae Colubrinae Natricinae Xenodontinae Elapidae Leptotyphlopidae Loxocemidae Typhlopidae Ungaliophiidae Viperidae TESTUDINES Cheloniidae Chelydridae Dermochelyidae Emydidae Geoemydidae Kinosternidae BIBLIOGRAPHY How to cite? Amphibians and Reptiles of Honduras 1

2 Museo de Zoología UCR INTRODUCTION McCranie and Castañeda (2007a) and McCranie (2006a) recently reviewed the amphibians of Honduras. Subsequently, numerous nomenclatural changes have been proposed and a few species have been added to the Honduran fauna. The non-avian reptiles of Honduras have not been treated in detail, but such a work is being prepared (McCranie, in prep.). As such, the most recent listing of the reptilian species of the country is that of Wilson and McCranie (2002). The purpose of this list is to update and provide a list of the amphibians and reptiles of Honduras as currently understood. The comments and cross-references included herein are only those pertaining to references published after McCranie and Castañeda (2007a), McCranie (2006a), and Wilson and McCranie (2002). The amphibians are listed in the same sequence by Order and within families as in McCranie and Castañeda (2007a). The reptiles are listed alphabetically by Order and then families within each Order. This listing does not include the taxonomic changes being proposed in McCranie (in prep.). The numbers in parenthesis following each Order or Suborder are the number of families, genera, and species, respectively, known to occur in Honduras. The numbers in parenthesis following each Family and Subfamily are the number of genera and species, respectively, known to occur in Honduras. The number in parenthesis following each genus is the number of species known to occur in Honduras. An asterisk denotes an introduced and established species. HERPETOFAUNA OF HONDURAS (2 classes, 6 orders, 42 families, 150 genera, 373 species) CLASS AMPHIBIA Gray, 1825 (3 orders, 13 families, 42 genera, 129 species) ORDER GYMNOPHIONA Müller, 1832 (1, 2, 2) FAMILY CAECILIIDAE Rafinesque, 1814 (2, 2) Dermophis Peters, 1879 (1) Dermophis mexicanus (A. M. C. Duméril & Bibron, 1841) Gymnopis Peters, 1874 (1) Gymnopis multiplicata Peters, 1874 ORDER CAUDATA Fischer von Waldheim, 1813 (1, 5, 31) FAMILY PLETHODONTIDAE Gray, 1850 (5, 31) Bolitoglossa A. M. C. Duméril, Bibron, & Duméril, 1854 (16) Subgenus Bolitoglossa A. M. C. Duméril, Bibron, & Duméril, 1854 (2) Bolitoglossa mexicana A. M. C. Duméril, Bibron, & Duméril, 1854 Bolitoglossa striatula (Noble, 1918) Subgenus Magnadigita Taylor, 1944 (12) Bolitoglossa carri McCranie & Wilson, 1993 Bolitoglossa cataguana Townsend, Butler, Wilson, & Austin, 2009 Bolitoglossa celaque McCranie & Wilson, 1993 Bolitoglossa conanti McCranie & Wilson, 1993 Bolitoglossa decora McCranie & Wilson, 1997 Bolitoglossa diaphora McCranie & Wilson, 1995 Bolitoglossa dunni (K. Schmidt, 1933) Bolitoglossa heiroreias Greenbaum, McCranie 2009

3 Listas Zoológicas Actualizadas UCR Bolitoglossa longissima McCranie & Cruz, 1996 Bolitoglossa oresbia McCranie, Espinal, & Wilson, 2005 Bolitoglossa porrasorum McCranie & Wilson, 1995 Bolitoglossa synoria McCranie & Köhler, 1999 Subgenus Nanotriton Parra-Olea, García-París, & Wake, 2004 (1) McCranie and Castañeda (2007a) and McCranie (2006a) included Bolitoglossa (Nanotriton) occidentalis Taylor, 1941 in the Honduran herpetofauna based on one poorly preserved specimen that carried erroneous locality data. That specimen is now thought to represent a juvenile B. mexicana (T. Papenfuss, pers. comm.). Bolitoglossa rufescens (Cope, 1869) Subgenus Pachymandra Parra-Olea, García-París, & Wake, 2004 (1) Bolitoglossa dofleini Werner, 1903 Cryptotriton García-París & Wake, 2000 (1) Cryptotriton nasalis (Dunn, 1924) Dendrotriton Wake & Elias, 1983 (1) Dendrotriton sanctibarbarus (McCranie & Wilson, 1997) Nototriton Wake & Elias, 1983 (4) Nototriton barbouri (K. Schmidt, 1936) Nototriton brodiei Campbell and Smith, Kolby et al. (2009) reported this species from Honduras. Nototriton lignicola McCranie & Wilson, 1997 Nototriton limnospectator McCranie, Wilson, & Polisar, 1998 Oedipina Keferstein, 1868 (9) McCranie et al. (2008) demonstrated that the O. cyclocauda of McCranie and Castañeda (2007a) and McCranie (2006a) actually consisted of three undescribed species. Subgenus Oedipina Keferstein, 1868 (4) Oedipina ignea Stuart, 1952 Oedipina leptopoda McCranie, Vieites, & Wake, 2008 Oedipina stuarti Brame, 1968 Oedipina taylori Stuart, 1952 Subgenus Oeditriton McCranie, Vieites, & Wake, 2008 Oedipina kasios McCranie, Vieites, & Wake, 2008 Oedipina quadra McCranie, Vieites, & Wake, 2008 Subgenus Oedopinola Hilton, 1946 (3) Oedipina elongata (K. Schmidt, 1936) Oedipina gephyra McCranie, Wilson, & Williams, 1993 Oedipina tomasi McCranie, 2006 ORDER ANURA Fischer von Waldheim, 1813 (11, 35, 96) FAMILY BUFONIDAE Gray, 1825 (4, 8) Atelophryniscus McCranie, Wilson, & Williams, 1989 (1) Atelophryniscus chrysophorus McCranie, Wilson, & Williams, 1989 Chaunus Wagler, 1828 (1) I follow Savage and Bolaños (2009) in retaining the following species in Chaunus rather than in Rhinella Fitzinger, 1826 as recently proposed by some authors. Chaunus marinus (Linnaeus, 1758) Incilius Cope, 1863 (5) Amphibians and Reptiles of Honduras 3

4 Museo de Zoología UCR The following five species were included in Ollotis Cope, 1875 by McCranie and Castañeda (2007a) and McCranie (2006a). Also, I do not recognize I. ibarrai (Stuart, 1954) or I. porteri (Mendelson, Williams, Sheil, & Mulcahy, 2005) as valid species occurring in Honduras for the reasons discussed by McCranie and Castañeda (2007a). Despite the recent review of the I. coccifer complex by Mendelson et al. (2005), a thorough review of this complex is needed that includes all known populations, especially those of the large specimens occurring in the lowlands of southern Honduras. Incilius campbelli (Mendelson, 1994) Incilius coccifer (Cope, 1866) Incilius leucomyos (McCranie & Wilson, 2000) Incilius luetkenii (Boulenger, 1891) Incilius valliceps (Wiegmann, 1833) Rhaebo Cope, 1862 (1) Rhaebo haematiticus (Cope, 1862) FAMILY CENTROLENIDAE Taylor, 1951 (5, 8) The classification of this Family follows Guayasamin et al. (2009). SUBFAMILY CENTROLENINAE Taylor, 1951 (4, 5) Cochranella Taylor, 1951 (1) Cochranella granulosa (Taylor, 1949) Espadarana Guayasamin, Castroviejo Fisher, Trueb, Ayarzagüena, Rada, & Vilà, 2009 (1) The following species was included in Centrolene Jiménez de la Espada, 1872 by McCranie and Castañeda (2007a) and McCranie (2006a). Espadarana prosoblepon (Boettger, 1892) Sachatamia Guayasamin, Castroviejo Fisher, Trueb, Ayarzagüena, Rada, & Vilà, 2009 (1) The following species was included in Cochranella by McCranie and Castañeda (2007a) and McCranie (2006a). Sachatamia albomaculata (Taylor, 1949) Teratohyla Taylor, 1951 (2) The following two species were included in Hyalinobatrachium and Cochranella, respectively, by McCranie and Castañeda (2007a) and McCranie (2006a). Teratohyla pulverata (Peters, 1873) Teratohyla spinosa (Taylor, 1949) SUBFAMILY HYALINOBATRACHINAE Guayasamin, Castroviejo Fisher, Trueb, Ayarzagüena, Rada, & Vilà, 2009 (1, 3) Hyalinobatrachium Ruiz-Carranza & Lynch, 1991 (3) The following two species were placed in the synonymy of H. chirripoi (Taylor, 1958) and H. colymbiphyllum (Taylor, 1949), respectively, by Cisneros-Heredia and McDiarmid (2007). However, the phylogeny based on nuclear genes recovered by Guayasamin et al. (2008, 2009) showed H. cardiacalyptum and H. chirripoi to be paraphyletic with respect to H. colymbiphyllum. Thus, I continue to recognize H. cardiacalyptum as a valid species. I also doubt the validity of placing H. crybetes in the synonymy of H. colymbiphyllum and continue to recognize the former as a distinct species pending rediscovery of the species and study of its molecular data and advertisement call. Hyalinobatrachium cardiacalyptum McCranie & Wilson, 1997 Hyalinobatrachium crybetes McCranie & Wilson, 1997 Hyalinobatrachium fleischmanni (Boettger, 1893) 4 McCranie 2009

5 FAMILY HYLIDAE Rafinesque, 1815 (16, 34) SUBFAMILY HYLINAE Rafinesque, 1815 (14, 30) Anotheca H. Smith, 1939 (1) Anotheca spinosa (Steindachner, 1864) Bromeliohyla Faivovich, Haddad, Garcia, Frost, Campbell, & Wheeler, 2005 (1) Bromeliohyla bromeliacia (K. Schmidt, 1933) Dendropsophus Fitzinger, 1843 (2) Dendropsophus ebraccatus (Cope, 1874) Dendropsophus microcephalus (Cope, 1886) Duellmanohyla Campbell & Smith, 1992 (2) Duellmanohyla salvavida (McCranie & Wilson, 1986) Duellmanohyla soralia (Wilson & McCranie, 1985) Ecnomiohyla Faivovich, Haddad, Garcia, Frost, Campbell, & Wheeler, 2005 (2) Ecnomiohyla miliaria (Cope, 1886) Ecnomiohyla salvaje (Wilson, McCranie, & Williams, 1985) Exerodonta Brocchi, 1879 (1) Exerodonta catracha (Porras & Wilson, 1987) Isthmohyla Faivovich, Haddad, Garcia, Frost, Campbell, & Wheeler, 2005 (2) Isthmohyla insolita (McCranie, Wilson, & Williams, 1993) Isthmohyla melacaena (McCranie & Castañeda, 2006) Plectrohyla Brocchi, 1877 (7) Plectrohyla chrysopleura Wilson, McCranie, & Cruz, 1994 Plectrohyla dasypus McCranie & Wilson, 1981 Plectrohyla exquisita McCranie & Wilson, 1998 Plectrohyla guatemalensis Brocchi, 1877 Plectrohyla hartwegi Duellman, 1968 Plectrohyla matudai Hartweg, 1941 Plectrohyla psiloderma McCranie & Wilson, 1999 Ptychohyla Taylor, 1944 (3) Ptychohyla hypomykter McCranie & Wilson, 1993 Ptychohyla salvadorensis (Mertens, 1952) Ptychohyla spinipollex (K. Schmidt, 1936) Scinax Wagler, 1830 (2) Scinax boulengeri (Cope, 1887) Scinax staufferi (Cope, 1865) Smilisca Cope, 1865 (3) Smilisca baudinii (A. M. C. Duméril & Bibron, 1841) Smilisca phaeota (Cope, 1862) Smilisca sordida (Peters, 1863) Tlalocohyla Faivovich, Haddad, Garcia, Frost, Campbell, & Wheeler, 2005 (2) Tlalocohyla loquax (Gaige & Stuart, 1934) Tlalocohyla picta (Günther, 1901) Trachycephalus Tschudi, 1838 (1) Trachycephalus venulosus (Laurenti, 1768) Triprion Cope, 1866 (1) Triprion petasatus (Cope, 1865) SUBFAMILY PHYLLOMEDUSINAE Günther, 1858 (2, 4) Agalychnis Cope, 1864 (3) Listas Zoológicas Actualizadas UCR Amphibians and Reptiles of Honduras 5

6 Museo de Zoología UCR Agalychnis callidryas (Cope, 1862) Agalychnis moreletii (A. H. A. Duméril, 1853) Agalychnis saltator Taylor, 1955 Cruziohyla Faivovich, Haddad, Garcia, Frost, Campbell, & Wheeler, 2005 (1) Cruziohyla calcarifer (Boulenger, 1902) FAMILY CRAUGASTORIDAE Hedges, Duellman, & Heinicke, 2008 (1, 28) This Family was included in the Brachycephalidae Günther, 1858 by McCranie and Castañeda (2007a) and McCranie (2006a). Craugastor Cope, 1862 (28) Subgenus Campbellius Hedges, Duellman, & Heinicke, 2008 (8) Craugastor chrysozetetes (McCranie, Savage, & Wilson, 1989) Craugastor cruzi (McCranie, Savage, & Wilson, 1989) Craugastor epochthidius (McCranie & Wilson, 1997) Craugastor fecundus (McCranie & Wilson, 1997) Craugastor milesi (K. Schmidt, 1933) Craugastor omoaensis (McCranie & Wilson, 1997) Craugastor saltuarius (McCranie & Wilson, 1997) Craugastor stadelmani (K. Schmidt, 1936) Subgenus Craugastor Cope, 1862 (19) Craugastor anciano (Savage, McCranie, and Wilson, 1988) Craugastor aurilegulus (Savage, McCranie, and Wilson, 1988) Craugastor chac (Savage, 1987) Craugastor charadra (Campbell & Savage, 2000) Craugastor coffeus (McCranie & Köhler, 1999) Craugastor emleni (Dunn, 1932) Craugastor fitzingeri (O. Schmidt, 1857) Craugastor laevissimus (Werner, 1896) Craugastor laticeps (A. H. A. Duméril, 1853) Craugastor lauraster (Savage, McCranie, & Espinal, 1996) Craugastor megacephalus (Cope, 1875) Craugastor merendonensis (K. Schmidt, 1933) Craugastor mimus (Taylor, 1955) Craugastor noblei (Barbour & Dunn, 1921) Craugastor olanchano (McCranie & Wilson, 1999) Craugastor pechorum (McCranie & Wilson, 1999) Craugastor rhodopis (Cope, 1867) Craugastor rostralis (Werner, 1896) Craugastor sp. This recently collected and apparently undescribed species from Intibucá will key out to C. emleni in the key in McCranie and Castañeda (2007a), but appears to be a bromeliad specialist, whereas C. emleni is a streamside inhabitant. Subgenus Hylactophryne Lynch, 1968 (1) Craugastor cyanochthebius McCranie & Smith, 2006 FAMILY ELEUTHERODACTYLIDAE Lutz, 1954 (2, 2) This Family was included in the Brachycephalidae and genus Eleutherodactylus by McCranie and Castañeda (2007a) and McCranie (2006a). Diasporus Hedges, Duellman, & Heinicke, 2008 (1) 6 McCranie 2009

7 Listas Zoológicas Actualizadas UCR Diasporus diastema (Cope, 1875) Eleutherodactylus A. M. C. Duméril & Bibron, 1841 (1) The following introduced and established species was recently reported from the country by McCranie et al. (2008). *Eleutherodactylus planirostris (Cope, 1862) FAMILY STRABOMANTIDAE Hedges, Duellman, & Heinicke, 2008 (1, 2) This Family was included in the Brachycephalidae and genus Eleutherodactylus by McCranie and Castañeda (2007a) and McCranie (2006a). Pristimantis Jiménez de la Espada, 1871 (2) Pristimantis cerasinus (Cope, 1875) Pristimantis ridens (Cope, 1866) FAMILY LEIUPERIDAE Bonaparte, 1850 (1, 1) This Family was included in the Leptodactylidae by McCranie and Castañeda (2007a) and McCranie (2006a). Engystomops Jiménez de la Espada, 1872 (1) Engystomops pustulosus (Cope, 1864) FAMILY LEPTODACTYLIDAE Werner, 1896 (1, 4) Leptodactylus Fitzinger, 1826 (4) Leptodactylus fragilis (Brocchi, 1877) Leptodactylus melanonotus (Hallowell, 1861) Leptodactylus savagei Heyer, 2005 Leptodactylus silvanimbus McCranie, Wilson, & Porras, 1980 FAMILY MICROHYLIDAE Günther, 1858 (2, 3) Gastrophryne Fitzinger, 1843 (1) Gastrophryne elegans (Boulenger, 1882) Hypopachus Keferstein, 1867 (2) Hypopachus barberi K. Schmidt, 1939 Hypopachus variolosus (Cope, 1866) FAMILY RANIDAE Rafinesque, 1814 (1, 5) Lithobates Fitzinger, 1843 (5) The next two species are members of the L. pipiens complex. That complex is in need of serious study in Honduras. Such a study using molecular and morphological data would likely indicate that at least three species are represented in the country, none of which are conspecific with L. brownorum or L. forreri. Lithobates brownorum (Sanders, 1973) Lithobates forreri (Boulenger, 1883) Lithobates maculatus (Brocchi, 1877) Lithobates vaillanti (Brocchi, 1877) Lithobates warszewitschii (O. Schmidt, 1857) FAMILY RHINOPHRYNIDAE Günther, 1859 (1, 1) Rhinophrynus A. M. C. Duméril & Bibron, 1841 (1) Rhinophrynus dorsalis A. M. C. Duméril & Bibron, 1841 Amphibians and Reptiles of Honduras 7

8 Museo de Zoología UCR CLASS REPTILIA Laurenti, 1768 (3 orders, 29 families, 108 genera, 244 species) ORDER CROCODYLIA Gmelin, 1789 (2, 2, 2) FAMILY ALLIGATORIDAE Cuvier, 1807 (1, 1) Caiman Spix, 1825 (1) Caiman crocodilus (Linnaeus, 1758) FAMILY CROCODYLIDAE Cuvier, 1807 (1, 1) Crocodylus Laurenti, 1768 (1) Crocodylus acutus (Cuvier, 1807) ORDER SQUAMATA Oppel, 1811 (21, 96, 227) SUBORDER SAURIA Macartney, 1802 (12, 29, 93) FAMILY ANGUIDAE Gray, 1825 (3, 6) Abronia Gray, 1838 (2) Abronia montecristoi Hidalgo, 1983 Abronia salvadorensis Hidalgo, 1983 Celestus Gray, 1839 (3) Celestus bivittatus (Boulenger, 1895) Celestus montanus K. Schmidt, 1933 Celestus scansorius McCranie & Wilson, 1996 Mesaspis Cope, 1877 (1) Mesaspis moreletii (Bocourt, 1871) FAMILY CORYTOPHANIDAE Fitzinger, 1843 (3, 7) Basiliscus Laurenti, 1768 (2) Basiliscus plumifrons Cope, 1875 Basiliscus vittatus Wiegmann, 1828 Corytophanes H. Boie, 1827 (3) Corytophanes cristatus (Merrem, 1820) Corytophanes hernandesii (Wiegmann, 1831) Corytophanes percarinatus A. H. A. Duméril, 1856 Laemanctus Wiegmann, 1834 (2) Laemanctus longipes Wiegmann, 1834 Laemanctus serratus Cope, 1864 FAMILY EUBLEPHARIDAE Boulenger, 1883 (1, 1) Coleonyx Gray, 1845 Coleonyx mitratus Peters, 1863 FAMILY GEKKONIDAE Gray, 1825 (6, 14) Aristelliger Cope, 1862 (2) Aristelliger georgeensis (Bocourt, 1873) Aristelliger praesignis (Hallowell, 1856) 8 McCranie 2009

9 Listas Zoológicas Actualizadas UCR Gonatodes Fitzinger, 1843 (1) Gonatodes albogularis (A. M. C. Duméril & Bibron, 1836) Hemidactylus Cuvier, 1820 (3) The next species was listed as H. brooki1 Gray, 1845 by Wilson and McCranie (2002). However, the correct name for the Honduran populations is more likely to be H. angulatus based on the molecular studies of Greater Antillean populations by Weiss and Hedges (2007). *Hemidactylus angulatus Hallowell, 1854 *Hemidactylus frenatus Schlegel, 1836 *Hemidactylus mabouia (Moreau de Jonnès, 1818) Phyllodactylus Gray, 1828 (2) Phyllodactylus palmeus Dixon, 1968 Phyllodactylus tuberculosus Wiegmann, 1834 Sphaerodactylus Wagler, 1830 (5) Sphaerodactylus dunni K. Schmidt, 1936 Sphaerodactylus glaucus Cope, 1865 Sphaerodactylus millepunctatus Hallowell, 1861 Sphaerodactylus notatus Baird, 1858 Sphaerodactylus rosaurae Parker, 1940 Thecadactylus Cuvier, 1820 (1) Thecadactylus rapicaudus (Houttuyn, 1782) FAMILY GYMNOPHTHALMIDAE Mac Lean, 1974 (1, 1) Gymnophthalmus Merrem, 1820 (1) Gymnophthalmus speciosus (Hallowell, 1861) FAMILY IGUANIDAE Oppel, 1811 (2, 7) Ctenosaura Wiegmann, 1828 (6) Ctenosaura bakeri Stejneger, 1901 Ctenosaura flavidorsalis Köhler & Klemmer, 1994 Ctenosaura melanosterna Buckley & Axtell, 1997 Ctenosaura oedirhina De Queiroz, 1987 Ctenosaura praeocularis Hasbún & Köhler, 2009 Ctenosaura similis (Gray, 1831) Iguana Laurenti, 1768 (1) Iguana iguana (Linnaeus, 1758) FAMILY LEIOCEPHALIDAE Frost & Etheridge, 1989 (1, 1) Leiocephalus Gray, 1827 *Leiocephalus carinatus Gray, 1827 FAMILY PHRYNOSOMATIDAE Fitzinger, 1843 (1, 4) Sceloporus Wiegmann, 1828 (4) The next named species is a complex that appears to represent at least three species in the country. Sceloporus schmidti is available for the population in western Copán and Cortés (E. Smith, pers. comm.), but no names are available for the others. Eric Smith and myself plan to study the Honduran populations in the near future. Sceloporus malachiticus Cope, 1864 Amphibians and Reptiles of Honduras 9

10 Museo de Zoología UCR Sceloporus olloporus H. Smith, Listed as S. variabilis Wiegmann, 1834 by Wilson and McCranie (2002). Sceloporus schmidti Jones, See comment preceding S. malachiticus. Sceloporus squamosus Bocourt, 1874 FAMILY POLYCHROTIDAE Fitzinger, 1843 (3, 38) Anolis Daudin, 1802 (1) Anolis allisoni Barbour, 1928 Norops Wagler, 1830 (36) Norops amplisquamosus McCranie, Wilson, & Williams, 1992 Norops bicaorum Köhler, 1996 Norops biporcatus (Wiegmann, 1834) Norops capito (Peters, 1863) Norops crassulus (Cope, 1864) Norops cupreus (Hallowell, 1861) Norops cusuco McCranie, Köhler, & Wilson, 2000 Norops dariense (Fitch & Seigel, 1984). Included in N. cupreus by Wilson and McCranie (2002). Norops heteropholidotus (Mertens, 1952) Norops johnmeyeri (Wilson & McCranie, 1982) Norops kreutzi McCranie, Köhler, & Wilson, 2000 Norops laeviventris (Wiegmann, 1834) Norops lemurinus (Cope, 1861) Norops limifrons (Cope, 1862) Norops loveridgei (K. Schmidt, 1936) Norops morazani (Townsend & Wilson, 2009) Norops muralla Köhler, McCranie, & Wilson, 1999 Norops ocelloscapularis Köhler, McCranie, & Wilson, 2001 Norops oxylophus (Cope, 1875). Listed as N. lionotus (Cope, 1861) by Wilson and McCranie (2002). Norops pentaprion (Cope, 1862) Norops petersii (Bocourt, 1873) Norops pijolensis McCranie, Wilson, & Williams, 1993 Norops purpurgularis McCranie, Cruz, & Holm, 1993 Norops quaggulus Cope, Listed as N. humilis Peters, 1863 by Wilson and McCranie (2002). Norops roatanensis Köhler & McCranie, 2001 Norops rodriguezii (Bocourt, 1873) Norops rubribarbaris Köhler, McCranie, & Wilson, 1999 *Norops sagrei (A. M. C. Duméril & Bibron, 1837). The mainland and Bay Island populations are introduced by man, whereas those on the Islas del Cisne appear to be natural. Norops sericeus Hallowell, 1857 Norops sminthus (Dunn & Emlen, 1932) Norops tropidonotus (Peters, 1863) Norops uniformis (Cope, 1885) Norops utilensis Köhler, 1996 Norops wampuensis McCranie & Köhler, 2001 Norops yoroensis McCranie, Nicholson, & Köhler, 2002 Norops zeus Köhler & McCranie, McCranie 2009

11 Listas Zoológicas Actualizadas UCR Polychrus Cuvier, 1816 (1) Polychrus gutturosus Berthold, 1845 FAMILY SCINCIDAE Gray, 1825 (4, 6) Mabuya Fitzinger, 1826 (1) Mabuya unimarginata Cope, 1862 Mesoscincus Griffith, Ngo, & Murphy, 2000 (1) Mesoscincus managuae (Dunn, 1933). Listed as Eumeces Wiegmann, 1834 by Wilson and McCranie (2002). Plestiodon A. M. C. Duméril & Bibron, 1839 (1) Plestiodon sumichrasti (Cope, 1867). Listed as Eumeces Wiegmann, 1834 by Wilson and McCranie (2002). Sphenomorphus Fitzinger, 1843 (3) Sphenomorphus assatus (Cope, 1864) Sphenomorphus cherriei (Cope, 1893) Sphenomorphus incertus (Stuart, 1940) FAMILY TEIIDAE Gray, 1827 (3, 6) Ameiva Meyer, 1795 (3) Ameiva ameiva (Linnaeus, 1758) Ameiva festiva (Lichtenstein & von Martens, 1856) Ameiva undulata (Wiegmann, 1834) Aspidoscelis Fitzinger, 1843 (2) Aspidoscelis deppii (Wiegmann, 1834) Aspidoscelis motaguae (Sackett, 1941) Cnemidophorus Wagler, 1830 (1) Cnemidophorus lemniscatus (Linnaeus, 1758) FAMILY XANTUSIIDAE Baird, 1859 (1, 2) Lepidophyma A. H. A. Duméril, 1851 (2) Lepidophyma flavimaculatum A. H. A. Duméril, 1851 Lepidophyma mayae Bezy, McCranie (2005) reported this species from Honduras. SUBORDER SERPENTES Linnaeus, 1758 (9, 67, 134) FAMILY ANOMALEPIDIDAE Taylor, 1939 (1, 1) Anomalepis Jan, 1860 (1) Anomalepis mexicanus Jan, McCranie (2004b) reported this species from Honduras. FAMILY BOIDAE Gray, 1825 (2, 2) Boa Linnaeus, 1758 (1) Boa constrictor Linnaeus, 1758 Corallus Daudin, 1803 (1) Corallus annulatus (Cope, 1875) FAMILY COLUBRIDAE Oppel, 1811 (50, 106) I am following the Subfamily arrangement (Colubrinae, Natricinae, and Xenodontinae) proposed by Lawson et al. (2005), although other workers would elevate those to Family level (see Zaher et al. Amphibians and Reptiles of Honduras 11

12 Museo de Zoología UCR 2009; their Dipsadidae corresponds to the Xenodontinae of Lawson et al.). Resolution of the paraphyletic Colubridae is in its early stages; therefore I take the conservative approach for the time being. SUBFAMILY COLUBRINAE Oppel, 1811 (22, 41) Chironius Fitzinger, 1826 (1) Chironius grandisquamis (Peters, 1868) Coluber Linnaeus, 1758 (1) Coluber mentovarius (A. M. C. Duméril, Bibron, & Duméril, 1854). Included in Masticophis Baird & Girard, 1853 by Wilson and McCranie (2002). Dendrophidion Fitzinger, 1843 (3) Dendrophidion nuchale (Peters, 1863) Dendrophidion percarinatum (Cope, 1893) Dendrophidion vinitor H. Smith, Wilson et al. (2003) reported this species from the country. Drymarchon Fitzinger, 1843 (1) Drymarchon melanurus (A. M. C. Duméril, Bibron, & Duméril, 1854). Listed as D. corais F. Boie, 1827 by Wilson and McCranie (2002). Drymobius Fitzinger, 1843 (3) Drymobius chloroticus (Cope, 1886) Drymobius margaritiferus (Schlegel, 1837) Drymobius melanotropis (Cope, 1875) Ficimia Gray, 1849 (1) Ficimia publia Cope, 1866 Lampropeltis Fitzinger, 1843 (1) Lampropeltis triangulum (Lacepède, 1789) Leptodrymus Amaral, 1927 (1) Leptodrymus pulcherrimus (Cope, 1874) Leptophis Bell, 1825 (5) Leptophis ahaetulla (Linnaeus, 1758) Leptophis depressirostris (Cope, 1861). Köhler and Ferrari (2009) reported this species from the country. Leptophis mexicanus (A. M. C. Duméril, Bibron, & Duméril, 1854) Leptophis modestus (Günther, 1872) Leptophis nebulosus Oliver, 1942 Mastigodryas Amaral, 1935 (2) The following two species were included in Dryadophis Stuart, 1939 by Wilson and McCranie (2002). Mastigodryas dorsalis (Bocourt, 1890) Mastigodryas melanolomus (Cope, 1868) Oxybelis Wagler, 1830 (4) Oxybelis aeneus (Wagler, 1824) Oxybelis brevirostris (Cope, 1861) Oxybelis fulgidus (Daudin, 1803) Oxybelis wilsoni Villa & McCranie, 1995 Pseudelaphe Mertens & Rosenberg, 1943 (1) Pseudelaphe flavirufa (Cope, 1867). Included in Elaphe Fitzinger, 1843 by Wilson and McCranie (2002). Pseustes Fitzinger, 1843 (1) 12 McCranie 2009

13 Listas Zoológicas Actualizadas UCR Pseustes poecilonotus (Günther, 1858) Rhinobothryum Wagler, 1830 (1) Rhinobothryum bovallii Andersson, 1916 Scaphiodontophis Taylor & Smith, 1943 (2) Scaphiodontophis annulatus (A. M. C. Duméril, Bibron, & Duméril, 1854) Scaphiodontophis venustissimus (Günther, 1894). Resurrected from synonymy of S. annulatus and reported from Honduras by McCranie (2006c). Scolecophis Fitzinger, 1843 (1) Scolecophis atrocinctus (Schlegel, 1837) Senticolis Dowling & Fries, 1987 (1) Senticolis triaspis (Cope, 1866) Spilotes Wagler, 1830 (1) Spilotes pullatus (Linnaeus, 1758) Stenorrhina A. M. C. Duméril, 1853 (2) Stenorrhina degenhardtii (Berthold, 1845) Stenorrhina freminvillii A. M. C. Duméril, Bibron, & Duméril, 1854 Tantilla Baird & Girard, 1853 (6) Tantilla armillata Cope, Listed as T. melanocephala (Linnaeus, 1758) by Wilson and McCranie (2002). Tantilla impensa Campbell, 1998 Tantilla lempira Wilson & Mena, 1980 Tantilla schistosa (Bocourt, 1883) Tantilla taeniata (Bocourt, 1883) Tantilla tritaeniata H. Smith & Williams, 1966 Tantillita H. Smith, 1941 (1) Tantillita lintoni (H. Smith, 1940) Trimorphodon Cope, 1862 (1) Trimorphodon quadruplex H. Smith, Listed as T. biscutatus (A. M. C. Duméril, Bibron, & Duméril, 1854) by Wilson and McCranie (2002). SUBFAMILY NATRICINAE Bonaparte, 1838 (2, 4) Storeria Baird & Girard, 1853 (1) Storeria dekayi (Holbrook, 1839) Thamnophis Fitzinger, 1843 (3) Thamnophis fulvus (Bocourt, 1893) Thamnophis marcianus (Baird & Girard, 1853) Thamnophis proximus (Say, 1823) SUBFAMILY XENODONTINAE Bonaparte, 1845 (26, 61) Adelphicos Jan, 1862 (1) Adelphicos quadrivirgatum Jan, 1862 Amastridium Cope, 1861 (1) Amastridium sapperi (Werner, 1903). Listed as A. veliferum Cope, 1861 by Wilson and McCranie (2002). Clelia Fitzinger, 1826 (1) Clelia clelia (Daudin, 1803) Coniophanes Hallowell, 1860 (4) Coniophanes bipunctatus (Günther, 1858) Coniophanes fissidens (Günther, 1858) Coniophanes imperialis (Baird & Girard, 1859) Amphibians and Reptiles of Honduras 13

14 Museo de Zoología UCR Coniophanes piceivittis Cope, 1870 Conophis Peters, 1860 (1) Conophis lineatus (A. M. C. Duméril, Bibron, & Duméril, 1854) Crisantophis Villa, 1971 (1) Crisantophis nevermanni (Dunn, 1937) Cubophis Hedges & Vidal, 2009 in Hedges et al (1) Cubophis cantherigerus (Bibron, 1840). Included in Alsophis Fitzinger, 1843 by Wilson and McCranie (2002). Dipsas Laurenti, 1768 (1) Dipsas bicolor (Günther, 1895) Enuliophis McCranie & Villa, 1993 (1) Enuliophis sclateri Boulenger, McCranie et al. (2003) reported this species from the country. Enulius Cope, 1871 (3) Enulius bifoveatus McCranie & Köhler, 1999 Enulius flavidorsalis (Cope, 1869) Enulius roatanensis McCranie & Köhler, 1999 Erythrolamprus F. Boie, 1827 (1) Erythrolamprus mimus (Cope, 1869) Geophis Wagler, 1830 (5) Geophis damiani Wilson, McCranie, & Williams, 1998 Geophis fulvoguttatus Mertens, 1952 Geophis hoffmanni (Peters, 1859) Geophis nephodrymus Townsend & Wilson, 2006 Geophis rhodogaster (Cope, 1868). McCranie & Castañeda (2007b) reported this species from the country. Hydromorphus Peters, 1859 (1) Hydromorphus concolor Peters, 1859 Imantodes A. M. C. Duméril, 1853 (3) Imantodes cenchoa (Linnaeus, 1758) Imantodes gemmistratus Cope, 1862 Imantodes inornatus Boulenger, 1896 Leptodeira Fitzinger, 1843 (3) Leptodeira annulata (Linnaeus, 1758) Leptodeira nigrofasciata Günther, 1868 Leptodeira septentrionalis (Kennicott, 1859) Ninia Baird & Girard, 1853 (5) Ninia diademata Baird & Girard, 1853 Ninia espinali McCranie & Wilson, 1995 Ninia maculata (Peters, 1861) Ninia pavimentata (Bocourt, 1883). Townsend et al. (2005) reported this species from the country, Ninia sebae (A. M. C. Duméril, Bibron, & Duméril, 1854) Nothopsis Cope, 1871 (1) Nothopsis rugosus Cope, 1871 Omoadiphas Köhler, Wilson, & McCranie, 2001 (2) Omoadiphas aurula Köhler, Wilson, & McCranie, 2001 Omoadiphas texiguatensis McCranie & Castañeda, McCranie 2009

15 Listas Zoológicas Actualizadas UCR Oxyrhopus Wagler, 1830 (1) Oxyrhopus petola (Linnaeus, 1758) Pliocercus Cope, 1860 (2) Pliocercus elapoides Cope, 1860 Pliocercus euryzonus Cope, Wilson et al. (2003) demonstrated the presence of this species in Honduras. Rhadinaea Cope, 1863 (8) Rhadinaea anachoreta E. Smith & Campbell, Townsend et al. (2006) reported this species from the country. Rhadinaea decorata (Günther, 1858). McCranie (2004a) reported this species from the country. Rhadinaea godmani (Günther, 1865) Rhadinaea kinkelini Boettger, 1898 Rhadinaea lachrymans (Cope, 1870) Rhadinaea montecristi Mertens, 1952 Rhadinaea pegosalyta McCranie, 2006d Rhadinaea tolpanorum Holm & Cruz, 1994 Sibon Fitzinger, 1826 (8) Sibon annulatus (Günther, 1872). McCranie et al. (2003) reported this species from the country. Sibon anthracops (Cope, 1868) Sibon carri (Shreve, 1951) Sibon dimidiatus (Günther, 1872) Sibon longifrenis (Stejneger, 1909) Sibon manzanaresi McCranie, 2007 Sibon miskitus McCranie, 2006b Sibon nebulatus (Linnaeus, 1758) Tretanorhinus A. M. C. Duméril, Bibron, & Duméril, 1854 (1) Tretanorhinus nigroluteus Cope, 1862 Tropidodipsas Günther, 1858 (2) Tropidodipsas fischeri Boulenger, 1894 Tropidodipsas sartorii Cope, 1863 Urotheca Bibron, 1843 (2) Urotheca decipiens (Günther, 1893). McCranie et al. (2003) reported this species from the country. Urotheca guentheri (Dunn, 1938) Xenodon H. Boie, 1826 (1) Xenodon rabdocephalus (Wied-Neuwied, 1824) FAMILY ELAPIDAE F. Boie, 1827 (2, 6) Micrurus Wagler, 1824 (5) Micrurus alleni K. Schmidt, 1936 Micrurus browni K. Schmidt & Smith, 1943 Micrurus diastema (A. M. C. Duméril, Bibron, & Duméril, 1854) Micrurus nigrocinctus (Girard, 1854) Micrurus ruatanus (Günther, 1895) Pelamis Daudin, 1803 (1) Pelamis platura (Linnaeus, 1766) FAMILY LEPTOTYPHLOPIDAE Stejneger, 1891 (1, 2) Amphibians and Reptiles of Honduras 15

16 Museo de Zoología UCR Epictia Gray, 1845 (2) The following two species were listed as Leptotyphlops goudotii A. M. C. Duméril & Bibron, 1844 in Wilson and McCranie (2002). Epictia ater (Taylor, 1940). My reasons for using this name for the Honduran mainland populations will be discussed in my volume on the snakes of Honduras (in prep.). Epictia magnamaculata (Taylor, 1940). Adalsteinsson et al. (2009), based on mtdna sequence data, resurrected this species from the synonymy of E. goudotii. I had earlier reached the same conclusion for the Caribbean Island populations of the E. goudotii complex based on morphological and color pattern differences. A discussion of these differences will be provided in my volume on the snakes of Honduras (in prep.). FAMILY LOXOCEMIDAE Cope, 1861 (1, 1) Loxocemus Cope, 1861 Loxocemus bicolor Cope, 1861 FAMILY TYPHLOPIDAE Fitzinger, 1826 (2, 4) Ramphotyphlops Fitzinger, 1843 (1) Ramphotyphlops braminus (Daudin, 1803). Vesely and Köhler (2009) reported this species from the country. Typhlops Oppel, 1811 (3) Typhlops costaricensis Jiménez & Savage, 1962 Typhlops stadelmani K. Schmidt, 1936 Typhlops tycherus Townsend, Wilson, Ketzler & Luque-Montes, 2008 FAMILY UNGALIOPHIIDAE McDowell, 1987 (1, 1) Ungaliophis Müller, 1880 (1) Ungaliophis continentalis Müller, 1880 FAMILY VIPERIDAE Oppel, 1811 (7, 11) Agkistrodon Palisot de Beauvois, 1799 (1) Agkistrodon bilineatus (Günther, 1863) Atropoides Werman, 1992 (2) Atropoides indomitus E. Smith & Ferrari-Castro, 2008 Atropoides mexicanus (A. M. C. Duméril, Bibron, & Duméril, 1854). Listed as A. nummifer Rüppell, 1845 by Wilson & McCranie (2002). Bothriechis Peters, 1859 (3) Bothriechis marchi (Barbour & Loveridge, 1929) Bothriechis schlegelii (Berthold, 1845) Bothriechis thalassinus Campbell & Smith, Listed as B. bicolor Bocourt, 1868 by Wilson and McCranie (2002). Bothrops Wagler, 1824 (1) Bothrops asper (Garman, 1884) Cerrophidion Campbell & Lamar, 1992 (1) Cerrophidion godmani (Günther, 1863) Crotalus Linnaeus, 1758 (1) Crotalus simus Latreille, Listed as C. durissus Linnaeus, 1758 by Wilson and McCranie (2002). Porthidium Cope, 1871 (2) 16 McCranie 2009

17 Listas Zoológicas Actualizadas UCR Porthidium nasutum (Bocourt, 1868) Porthidium ophryomegas (Bocourt, 1868) ORDER TESTUDINES Batsch, 1788 (6, 10, 15) FAMILY CHELONIIDAE Oppel, 1811 (4, 4) Caretta Rafinesque, 1814 (1) Caretta caretta (Linnaeus, 1758) Chelonia Brongniart, 1800 (1) Chelonia mydas (Linnaeus, 1758) Eretmochelys Fitzinger, 1843 (1) Eretmochelys imbricata (Linnaeus, 1766) Lepidochelys Fitzinger, 1843 (1) Lepidochelys olivacea (Eschscholtz, 1829) FAMILY CHELYDRIDAE Gray, 1831 (1, 2) Chelydra Schweigger, 1812 (1) The following two species were listed as C. serpentina (Linnaeus, 1758) in Wilson and McCranie (2002). Chelydra acutirostris Peters, 1862 Chelydra rossignonii (Bocourt, 1868). FAMILY DERMOCHELYIDAE Fitzinger, 1843 (1, 1) Dermochelys Blainville, 1816 (1) Dermochelys coriacea (Vandelli, 1761) FAMILY EMYDIDAE Rafinesque, 1815 (1, 1) Trachemys Agassiz, 1857 (1) Trachemys venusta (Gray, 1855). Listed as T. scripta (Schoepff, 1792) by Wilson and McCranie (2002). FAMILY GEOEMYDIDAE Theobald, 1868 (1, 4) Rhinoclemmys Fitzinger, 1835 (4) Rhinoclemmys annulata (Gray, 1860) Rhinoclemmys areolata (A. M. C. Duméril & Bibron, 1851) Rhinoclemmys funerea (Cope, 1875) Rhinoclemmys pulcherrima (Gray, 1855) FAMILY KINOSTERNIDAE Agassiz, 1857 (2, 3) Kinosternon Spix, 1824 (2) Kinosternon leucostomum (A. M. C. Duméril & Bibron, 1851) Kinosternon scorpioides (Linnaeus, 1766) Staurotypus Wagler, 1830 (1) Staurotypus triporcatus (Wiegmann, 1828) BIBLIOGRAPHY Amphibians and Reptiles of Honduras 17

18 Museo de Zoología UCR Adalsteinsson, S. A., W. R. Branch, S. Trape, L. J. Vitt & S. B. Hedges Molecular phylogeny, classification, and biogeography of snakes of the family Leptotyphlopidae (Reptilia, Squamata). Zootaxa 2244:1 50. Cisneros-Heredia, D. F. & R. W. McDiarmid Revision of the characters of Centrolenidae (Amphibia: Anura: Athesphatanura), with comments on its taxonomy and the description of new taxa of glassfrogs. Zootaxa 1572:1 82. Available. Guayasamin, J. M., S. Castroviejo-Fisher, J. Ayarzagüena, L. Trueb & C. Vilà Phylogenetic relationships of glassfrogs (Centrolenidae) based on mitochondrial and nuclear genes. Molecular and Phylogentics Evolution 48: Guayasamin, J. M., S. Castroviejo-Fisher, L. Trueb, J. Ayarzagüena, M. Rada & C. Vilà Phylogenetic systematics of glassfrogs (Amphibia: Centrolenidae) and their sister taxon Allophryne ruthveni. Zootaxa 2100:1 97. Available. Hasbún, C. R. & G. Köhler New species of Ctenosaura (Squamata, Iguanidae) from southern Honduras. Journal of Herpetology 43: Hedges, S. B., W. E. Duellman & M. P. Heinicke New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 1737: Hedges, S. B., A. Couloux & N. Vidal Molecular phylogeny, classification, and biogeography of West Indian racer snakes of the Tribe Alsophiini (Squamata, Dipsadidae, Xenodontinae). Zootaxa 2067:1 28. Köhler, G. & J. Ferrari Geographic distribution. Leptophis depressirostris. Herpetological Review (in press) Kolby, J. E., J. R. McCranie & S. M. Rovito Geographic distribution. Nototriton brodiei (NCN). Herpetological Review (in press) Lawson, R., J. B. Slowinski, B. I. Crother & F. T. Burbrink Phylogeny of the Colubroidea (Serpentes); new evidence from mitochondrial and nuclear genes. Molecular and Phylogentics Evolution 37: McCranie, J. R. 2004a. Geographic distribution. Rhadinaea decorata. Herpetological Review 35:294. McCranie, J. R. 2004b. Anomalepis mexicanus Jan (Serpentes, Anomalepididae) in Honduras. Herpetological Bulletin 89:21. McCranie, J. R (dated 2004). The herpetofauna of Parque Nacional Cerro Azul, Honduras (Amphibia, Reptilia). Herpetological Bulletin 90: McCranie, J. R. 2006a. Specimen locality data & museum numbers / Ubicación y números de museo de los especímenes, Información complementaria for / a la "Guía de Campo de los Anfibios de Honduras" by / por James R. McCranie y Franklin E. Castañeda. Smithsonian Herpetological Information Service 137:1-39. Available. McCranie, J. R. 2006b. New species of Sibon (Serpentes: Colubridae) from northeastern Honduras. Journal of Herpetology 40: McCranie, J. R. 2006c. Northeastern Honduras and the systematics of the colubrid snake Scaphiodontophis. Caribbean Journal of Science 42: McCranie, J. R. 2006d. New species of snake of the colubrid genus Rhadinaea (godmani group) from Parque Nacional El Cusuco, Honduras. Proceedings of the Biological Society of Washington 119: McCranie, J. R A second new species of Sibon (Squamata: Colubridae) from La Mosquitia, northeastern Honduras. Herpetologica 63: McCranie, J. R. & F. E. Castañeda A new species of snake of the genus Omoadiphas (Reptilia: Squamata: Colubridae) from the Cordillera Nombre de Dios in northern Honduras. Proceedings of the Biological Society of Washington 117: McCranie 2009

19 Listas Zoológicas Actualizadas UCR McCranie, J. R. & F. E. Castañeda. 2007a. Guía de Campo de los Anfibios de Honduras. Bibliomania!, Salt Lake City, Utah. USA. x pp. McCranie, J. R. & F. E. Castañeda. 2007b. The colubrid snake Geophis rhodogaster (Cope) in Honduras. Herpetological Bulletin 99:19. McCranie, J. R., J. H. Townsend & L. D. Wilson Three snakes new to the herpetofauna of Honduras. Herpetological Review 34: McCranie, J. R., J. R. Collart, F. E. Castañeda & J. M. Solis Geographic distribution. Eleutherodactylus (Euhyas) planirostris (Greenhouse Frog). Herpetological Review 39: McCranie, J. R., D. R. Vieites & D. B. Wake, Description of a new divergent lineage and three new species of Honduran salamanders of the genus Oedipina (Caudata, Plethodontidae). Zootaxa 1930:1 17. Mendelson, J. R., III, B. L. Williams, C. A. Sheil & D. G. Mulcahy Systematics of the Bufo coccifer complex (Anura: Bufonidae) of Mesoamerica. Scientific Papers, Natural History Museum, The University of Kansas 38:1 27. Savage, J. M. & F. Bolaños A checklist of the amphibians and reptiles of Costa Rica: additions and nomenclatural revisions. Zootaxa 2005:1 23. Smith, E. N. & J. A. Ferrari-Castro A new species of jumping pitviper of the genus Atropoides (Serpentes: Viperidae: Crotalinae) from the Sierra de Botaderos and Sierra La Muralla, Honduras. Zootaxa 1948: Townsend, J. H. & L. D. Wilson A new species of snake of the Geophis dubius group (Reptilia: Squamata: Colubridae) from the Sierra de Omoa of northwestern Honduras. Proceedings of the Biological Society of Washington 119: Townsend, J. H. & L. D. Wilson New species of cloud forest Anolis (Squamata: Polychrotidae) in the crassulus group from Parque Nacional Montaña de Yoro, Honduras. Copeia 2009: Townsend, J. H., L. D. Wilson, T. L. Plenderleith, B. L. Talley & J. C. Nifong Ninia pavimentata (Squamata: Colubridae); an addition to the snake fauna of Honduras. Caribbean Journal of Science 41: Townsend, J. H., J. C. Nifong & L. D. Wilson (dated 2005). First record of the colubrid snake Rhadinaea anachoreta Smith and Campbell from Honduras. Herpetological Bulletin 94:2 3. Townsend, J. H., L. D. Wilson, L. P. Ketzler & I. R. Luque-Montes The largest blindsnake in Mesoamerica: a new species of the Typhlops (Squamata: Typhlopidae) from an isolated karstic mountain in Honduras. Zootaxa 1932: Townsend, J. H., J. M. Butler, L. D. Wilson & J. D. Austin A new species of salamander in the Bolitoglossa dunni group (Caudata: Plethodontidae: Bolitoglossinae) from Parque Nacional Montaña de Yoro, Honduras. Salamandra 45: Vesely, M. & G. Köhler Geographic distribution. Ramphotyphlops braminus (Braminy Blindsnake). Herpetological Review 40:116. Weiss, A. J. & S. B. Hedges Molecular phylogeny and biogeography of the Antillean geckos Phyllodactylus wirshingi, Tarentola americana, and Hemidactylus haitianus (Reptilia, Squamata). Molecular and Phylogentics Evolution 45: Wilson, L. D. & J. R. McCranie Update on the list of reptiles known from Honduras. Herpetological Review 33: Wilson, L. D., J. R. McCranie, S. Gotte & J. H. Townsend Distributional comments on some members of the herpetofauna of the Mosquitia, Honduras. Herpetological Bulletin 84: Zaher, H., F. G. Grazziotin, J. E. Cadle, R. W. Murphy, J. Cesar de Moura-Leite & S. L. Bonatto Molecular phylogeny of advanced snakes (Serpentes, Caenophidia) with an emphasis on South American xenodontines: a revised classification and description of new taxa. Papéis Avulsos de Zoologia (São Paulo) 49: Available. Amphibians and Reptiles of Honduras 19

20 Museo de Zoología UCR HOW TO CITE? McCranie, J. R Amphibians and Reptiles of Honduras. Listas Zoológicas Actualizadas UCR: Museo de Zoología UCR. San Pedro, Costa Rica. Last Actualization on November 12, McCranie 2009