Family SUPERSTITIONIIDAE Stahnke, 1940

Transcription

1 Family SUPERSTITIONIIDAE Stahnke, 1940 W. DAVID SISSOM This taxon, which now includes four genera and nine species, was first recognized by Stahnke ( 1940) as a monotypic subfamily of the Chactidae containing his new genus Superstitionia. Stahnke used the spelling Superstitioninae which is incorrect; the correct latinized subfamily name derived from the type genus is Superstitioniinae (H. D. Cameron, pers. comm), and the spelling is corrected here. Later, Francke ( l 982a) demonstrated a phylogenetic relationship of Superstitionia with members of another chactid subfamily, the Typhlochactinae Mitchell, and merged them into one subfamily, the Superstitioninae. Reluctant to recognize a family Chactidae in the broad sense that it had been previously defined, he placed this subfamily "incertae sedis" in the "Chactoidea". Francke (l 982a) proposed that Superstitionia be placed in a tribe Superstitionini and the members of the former Typhlochactinae (then including Typhlochactas and Alacran) in a tribe Typhlochactini. Subsequently, Typhochactas elliotti Mitchell was placed into its own genus, Sotanochactas (Francke, 1986b). Stockwell (1992) then elevated the Superstitioninae to family level and included two additional eyeless forms: Troglotayosicus from Ecuador and Belisarius Simon from the Pyrenees of France and Spain. The inclusion of these two genera was based on very weak evidence (Stockwell, 1992: ). Recently, (1998d) separated Belisarius and Troglotayosicus in a family Troglotayosicidae. We follow this placement but suggest that the issue requires further study. With removal of these two genera, the geographic range of Superstitioniidae is limited to the southern part of North America. Francke had recognized the two tribes in the subfamily Superstitioninae, and with the elevation of the subfamily to the family level it seems appropriate to regard his tribes as subfamilies. The relationships of the Superstitioniidae (with or without Belisarius and Troglotayosicus) to other families remain obscure. Stockwell (1992) suggested a close relationship to vaejovids and iurids. This placement is only weakly supported by cladistic evidence at this time. The members of the Superstitioniidae are interesting scorpions. Alacran, Sotanochactas, and most Typhlochactas are troglobites in wet subtropical caves of eastern and southern Mexico. Two Oaxacan species of Typhlochactas, however, are epigean and live in leaf litter; like their cave relatives, they are small, eyeless, and lack pigment. Alacran is represented by a remarkable eyeless form living in very deep caves in the Sistema Huautla of Oaxaca. It has been found at depths of about 800 m below the surface, and interestingly, it is a large (ca. 70 mm in length), wellpigmented scorpion. Superstitioninae (IOS) Stahnke, 1940: 102; type genus Superstitionia Stahnke, Superstitioninae (IOS): Millot & Vachon, 1949: 429; Soleglad, 1976b: 254; Francke, 1982a: 60-61; Sissom, 1990a: 113. Superstitionidae (IOS): Stockwell, 1992: 410, 412, 419 (part); Kovaffk, 1998c: 142. DISTRIBUT!ON: NORTH AMERICA (Mexico, southwestern USA). Subfamily SUPERSTITIONIINAE Stahnke, 1940 Superstitioninae (IOS) Stahnke, 1940: 102; type genus Superstitionia Stahnke, Superstitionini (IOS): Francke, 1982a: (tribe). 496

2 SCORPIONES: SUPERSTITIONIIDAE 497 DISTRIBUTION: NORTH AMERICA (western Mexico, southwestern USA). NOTES: The correct latinized subfamily name derived from the type genus is Superstitioniinae (H. D. Cameron, pers. comm), and the spelling is corrected here. Genus SUPERSTITIONIA Stahnke, 1940 Superstitionia Stahnke, 1940: 102; type species Superstitionia donensis Stahnke, SYNONYMS Diplops Mulaik & Higgins, 1944: ; type species Diplops desertorum Mulaik & Higgins, 1944 (= Superstitionia donensis Stahnke, 1940) (synonymized by Stahnke, 1949: 243). Superstitionia: Millot & Vachon, 1949: 429; Vachon, 1974: 914, 923; Williams, 1980: 7-8, fig. 5, 6; Francke, 1982a: 60-61, fig. 26; Francke, 1985: 13, 16, 21; Sissom, 1990a: 110, 113; Nenilin & Pet, 1992: 9; Stockwell, 1992: 410, 412, 419, fig. 22, 25, 27, 29; Kovarik, 1998c: 142; Lourern;o, 1998d: 136; Sissom & Cokendolpher, 1998: 285. DISTRIBUTION: NORTH AMERICA. Southwestern USA, western Mexico. Superstitionia donensis Stahnke, 1940 Superstitionia donensis Stahnke, 1940: I 02. SYNTYPES: 4 specimens (CAS, Type No ), Superstition Mountains, Arizona, USA. SYNONYMS Diplops desertorum Mulaik & Higgins, 1944: , fig. 1-9 (synonymized by Stahnke, 1949: 243). HOLOTYPE: Q (ANSP?), 16 mi E Tucson, Arizona, USA. PARATYPES: 6 specimens (see Notes), same locality as holotype; 1 Q (depository unknown), 6 mi W Bishop, California, USA; 1 (depository unknown), Mount Springs, San Diego County, California, USA. Broteochactas alleni (MIS; part): Banks, 1910: 185, 188. Superstitionia donensis: Stahnke, 1949: 243; Gertsch &Allred, 1965: 2, 7, 13, 14, fig. 20; Johnson & Allred, 1972: 156; Hjelle, 1972: 5-7; Vachon, 1974: 931, fig. 182; Frohlich & Clark, 1976: 91; Williams, 1976: 2; Williams, 1980: 8-9, fig. 5, 6; Francke, 1982a: 59-60, fig ; Sissom, 1997: 13; Kovarfk, l 998c: 142; Lourenc;o, 1998d: 142, fig. 11. DISTRIBUTION: Mexico (Baja California Norte, Baja California Sur, Sonora), USA (Arizona, California, New Mexico, Nevada). NOTES: Mulaik & Higgins (1944) indicated that the types of Diplops desertorum were split between the Zoological Museum of the University of Utah and the Academy of Natural Sciences in Philadelphia. There was no indication where specific material was deposited. Subfamily TYPHLOCHACTINAE Mitchell, 1971 Typhlochactinae Mitchell, 1971 (see Notes); type genus Typhlochactas Mitchell, Typhlochactinae: Mitchell, 1968: 774, nomen nudum (based on unavailable generic name); Soleglad, 1976b: 299, 301. Typhlochactini: Francke, 1982a: 60, 61 (as tribe). DISTRIBUTION: NORTH AMERICA (Mexico). NOTES: The genus Typhlochactas was first described by Mitchell (1968: ) without type species designation. It becomes first available with the type species designation by Mitchell (1971), which is also a date when the subfamily name based on this genus becomes available. Genus ALACRAN Francke, 1982 Alacran Francke, 1982a: 51-53; type species Alacran tartarus Francke, 1982.

3 498 CATALOG OF THE SCORPIONS OF THE WORLD Alacran: Francke, 1985: 5, 16, 19; Lourenc;:o & Francke, 1985: 4; Francke, 1986b: 8; Sissom, 1990a: 109, 113; Nenilin & Fet, 1992: 10; Stockwell, 1992: 410, 412, 419, fig. 24, 31, 32; Armas, 1994: 19; Kovarfk, 1998c: 142; Sissom & Cokendolpher, 1998: 285. DISTRIBUTION: NORTH AMERICA (southern Mexico) Alacran tartarus Francke, 1982 Alacran tartarus Francke, 1982a: 53-57, fig. 1-17, 27, 32. HoLOTYPE: (AMNH), S6tano de San Agustin (720m below the surface), San Agustin (5 km SE Huautla de Jimenez), Oaxaca, Mexico. PARATYPES: 1 juv (AMNH), same locality as holotype; 1 o, 1 1 juv (AMNH), adjoining caves in vicinity of San Agustin, Oaxaca, Mexico. Alacran tartarus: Lourenc;:o & Francke, 1985: 5 (tab. l); Nenilin & Fet, 1992: 28; Armas, 1994: 19, 21; Lourenc;:o, 1994b: 182, 183; Kovarik, 1998c: 142. D1STRIBUTION: Mexico (Oaxaca: caves of the Huautla Plateau). Genus SOTANOCHACTAS Francke, 1986 Sotanochactas Francke, 1986b: 8, fig ; type species Typhlochactas elliotti Mitchell, 1971 [= Sotanochactas elliotti (Mitchell, 1971)]. Sotanochactas: Lourenc;:o & Francke, 1985: 4 (nomen nudum); Sissom, 1990a: 109, 113; Stockwell, 1992: 410, 412, 419, fig. 10, 34; Kovarfk, 1998c: 142; Sissom & Cokendolpher, 1998: 285. DISTRIBUTION: NORTH AMERICA. Central Mexico. Sotanochactas elliotti (Mitchell, 1971) Typhlochactas elliotti Mitchell, 1971: , fig. 1-17, 26. HOLOTYPE: o (AMNH), El S6tano de Yerbaniz, approx. 21 km N Ciudad Valles, San Luis Potosf, Mexico. PARATYPES: 1 juv (MNHN RS 5376), 1 specimen (depository unknown), same locality as holotype. Typhlochactas elliotti: Reddell & Mitchell, 197la: 144, fig. 1; Reddell, 1973: 33, 37; Reddell & Elliott, 1973a: 171; Gonzalez-Sponga, 1974c: 56; Soleglad, 1976b: 253; Mitchell & Peck, 1977: ; Lourenc;:o, 198lb: 660; Reddell, 1981: 115, fig. 17; Francke, 1982a: 59-60, fig , 26, 28, 33. Typlochactas (ISS) elliotti: Dfaz Najera, 1975: 3, 30. Sotanochactas ellioti (ISS): Lourenc;:o & Francke, 1985: 5 (Table l); Armas, 1994: 21; Kovarik, 1998c: 142. Sotanochactas elliotti: Francke, 1986b: 8, fig ; Lourenc;:o, 1994b: 182, 183. DISTRIBUTION: Mexico (San Luis Potosi: El S6tano de Yerbaniz). Genus TYPHLOCHACTAS Mitchell, 1971 Typhlochactas Mitchell, 1971: 238; type species by subsequent designation (Mitchell, 1971: 238) Typhlochactas rhodesi Mitchell, Typhlochactas: Mitchell, 1968: ; Mitchell, 1971: 238 (part); Vachon, 1974: 914, 923; Soleglad, 1976b: ; Mitchell & Peck, 1977: (part); Francke, 1985: 14, 16, 20; Francke, 1986b: 8; Sissom, 1990a: 109, 114; Nenilin & Fet, 1992: 10, 28; Stockwell, 1992: 410, 412, 419, fig. 3, 28, 30, 33, 35; Armas, 1994: 18-19; Kovarfk, 1998c: 142; Sissom & Cokendolpher, 1998: Typlochactas (ISS): Dfaz Najera, 1975: 3. DISTRIBUTION: NORTH AMERICA. Eastern & southern Mexico. NOTES: The genus was first described by Mitchell (1968: ) without type species designation. The name becomes first available with the type species designation by Mitchell (1971).

4 SCORPIONES: SUPERSTITIONIIDAE 499 Typhlochactas cavicola Francke, 1986 Typhlochactas cavicola Francke, 1986b: 5-8, fig. 1-10, HOLOTYPE: o (AMNH), Cueva del Vandalismo, entrance about 2600 m elev., 1 km SE Rancho Nuevo, Tamaulipas, Mexico. Typhlochactas cavicola: Armas, 1994: 21; Lourencto, 1994b: 182, 183; Kovarik, 1998c: 142; Sissom & Cokendolpher, 1998: 285, 287. DISTRIBUTION: Mexico (Tamaulipas: Cueva del Vandalismo). Typhlochactas granulosus Sissom & Cokendolpher, 1998 Typhlochactas granulosus Sissom & Cokendolpher, 1998: , fig HoLOTYPE: o (AMNH), S6tano de Poncho, Municipio Tlaquilpa, Veracruz, Mexico. DISTRIBUTION: Mexico (Veracruz). Typhlochactas mitchelli Sissom, 1988 Typhlochactas mitchelli Sissom, 1988b: , fig HoLOTYPE: o (AMNH), Cerro Ocote, 5 mi S Tenango, Oaxaca, Mexico. PARATYPES: 1 o, 1 sub (AMNH), same locality as holotype. Typhlochactas mitchelli: Kovarik, 1998c: 142; Sissom & Cokendolpher, 1998: 285, 287. DISTRIBUTION: Mexico (Oaxaca) Typhlochactas reddelli Mitchell, 1968 Typhlochactas reddelli Mitchell, 1968: , , fig HOLOTYPE: <.? (AMNH), La Cueva del Ojo de Agua de Tlilapan (1400 m), in the village oftlilapan, Municipio de Tlilapan, (ca. 5 km S Orizaba), Veracruz, Mexico. Typhlochactas reddelli: Reddell, 197la: 26; Reddell, 197lb: 217; Mitchell, 1971: 135, 138, 143, , fig , 25, 27; Reddell, 1973: 32, 37; Vachon, 1974: fig. 167, ; Soleglad, 1976b: 254; Mitchell & Peck, 1977: ; Reddell, 1981: 115, fig. 17; Francke, 1982a: 59-60, fig. 26, 30, 35; Lourencto & Francke, 1985: 3, 5 (tab. l); Francke, 1986b: 8, fig ; Armas, 1994: 18, 21; Lourencto. 1994b: 182, 183; Sissom & Cokendolpher, 1998: 285, 286, 287. Typhlochactas reddeli (ISS): GonzaJ.ez-Sponga, 1974c: 56. Typlochactas (ISS) redelli (ISS): Diaz Najera, 1975: 4, 33. Typhlochactas redelli (ISS): Lourencto. 1981b: 660; Kovarfk., 1998c: 142. DISTRIBUTION: Mexico (Veracruz: La Cueva del Ojo de Agua de Tlilapan). Typhlochactas rhodesi Mitchell, 1968 Typhlochactas rhodesi Mitchell, 1968: , , fig HoLOTYPE: '? (AMNH), La Cueva de la Mina ( 1600 m), Sierra de Guatemala, within the Municipio de Gomez Farias, Tamaulipas, Mexico. PARATYPES: 1 '? (MNHN), I'? (depository unknown), same locality as holotype. Typhlochactas rhodesi: Mitchell, 1971: 135, 138, 143, , fig , 24, 27; Delamare Deboutteville, 1971: 49; Reddell, 197la: 3, 27; Reddell & Mitchell, 197lb: 181, 184, fig. 9; Reddell, 1973: 32, 37; Reddell & Elliott, 1973b: 181; Gonzalez-Sponga, 1974c: 56; Soleglad, 1976b: 253; Mitchell & Peck, 1977: ; Reddell, 1981: 115, fig. 17; Lourencto, 198lb: 660; Francke, 1982a: 59-60, fig. 26, 29, 34; Lourencto & Francke, 1985: 3, 5 (Table l); Francke, 1986b: 8, fig ; Armas, 1994: 18, 21; Louren<to, 1994b: 182, 183; Kovarik, 1998c: 142; Sissom & Cokendolpher, 1998: 285, 286, 287. Typlochactas (ISS) rhodesi: Diaz Najera, 1975: 4, 32. DISTRIBUTION: Mexico (Tarnaulipas: La Cueva de la Mina).

5 500 CATALOG OF THE SCORPIONS OF THE WORLD Typhlochactas sylvestris Mitchell & Peck, 1977 Typhlochactas sylvestris Mitchell & Peck, 1977: , fig HoLOTYPE: <j? (AMNH), 25 km S village of Valle Nacional at 1200 m along Highway 175 (highway connecting Oaxaca and Tuxtepec), Oaxaca, Mexico. Typhlochactas sylvestris: Lourem;o, 1981b: 659; Reddell, 1981: 115; Francke, 1982a: 59-60, fig. 26, 31, 36; Francke, 1986b: 8, fig ; Sissom, 1988b: ; Kovai'fk, 1998c: 142; Sissom & Cokendolpher, 1998: 285, 287. DISTRIBUTION: Mexico (Oaxaca). NOTES: Mitchell & Peck (1977) listed Texas Tech University (TTU) as the depository, but the specimen has since been transferred to the AMNH.