MISCELLANEOUS PUBLICATION OCCASIONAL PAPER NO. 95

Transcription

1 MISCELLANEOUS PUBLICATION OCCASIONAL PAPER NO. 95

2 A TAXONOMIC REVIEW OF ORIENTAL PHYTOSEIIDAE WITH KEYS TO GENERA AND SPECIES BY s. K. GUPTA

3 RECORDS OF THE ZOOLOGICAL SURVEY OF INDIA MISCELLANEOUS PUBLICATION OCCASIONAL PAPER NO. 95 A TAXONOMIC REVIEW OF ORIENTAL PHYTOSEIIDAE WITH KEYS TO GENERA AND SPECIES BY s K. GUPTA... at- Edited by the Director, Zoological Survey of India 1987

4 @ Copyright 1987~ Government of India, Published in September, 1987 PRICE: Inland : Rs_ 60'00 Foreign: 6-50, $ 10'00 PRINTED IN INDIA AT SRI AUROBINDO PRESS, 16, HEMENDRA "SEN STREET, CALCUTTA AND PUBLI~HED BY THE DIRECTOR, ZOOLOGICAL SURVEY OF INDIA, CALCUTTA _

5 RECORDS OF THE ZOOLOGICAL SURVEY OF' INDIA MISCELLANEOUS PUBLICATION Occasional Paper No Pages CONTENTS PAGE INTRODUCTION 1 Family Phytoseiidae 3 Key to the subfamilies of Phytoseiidae 3 Subfamily Amblyseiinae 4 Key to the genera of Amblyseiinae 4 GENUS Amblyseius Berlese 4 Key to the subgenera of Amblyseius 5 Subgenus Amblyseius 6 Key to the species of Amblyseiu~ (Amblyseius) 6 1. Amblyseius (Amblyseius) adhatodae Muma 7 2. A. (A.) aerialis (Muma) 7 3. A. (A.) channabasavannai Gupta 8 4. A. (A.) cinctus Corpuz & Rimando 8 5. A. (A.) herbicolus (Chant) 9 6. A. (A.) impactus Chaudhri 9 7. A. (A.) kulini Gupta A. (A.) largoensis (Muma) A. (A.) mcmurtryi Muma A. (A.) neorykei Gupta A. (A.) obtusus (Koch) A. (A.) orientalis Ehara A. (A.) paraaerialis Muma A. (A.) muraleedharani Gupta A. (A.) shoreae Gupta 14

6 [ ii ] 16. A. (A.) bajulus Chaudhri, Akbar & Rasool A. (A.) brevicervix Wei-nan & Zhao-Quan A. (A.) deductus Chaudhri, Akbar & Rasool A. (A.) excelsus Chaudhri, Akbar & Rasool A. (A.) hainanensis Wei-nan & Xing A. (A.) hapoliensis Gupta A. (A.) indirae Gupta A. (A) maai Tseng A. (A. 1) monomacroseta Tseng A. (A.) obtuserellus Wainstein & Beglyarov A. (A.) pubes Tseng A. (A.) spineus Tseng A. (A.) tamtavensis Blommers A. (A.) theae Wei-nan & Xing A. (A.) vestificus Tseng 18 Subgenus Asperoseius Chant 18 Key to the species of AmblYseius (Asper~seius) A. (A.) heveae (Oudemans) A. (A.) nucifera (Gupta) A. (A.) hevearum (Oudemans) A. (A.) hyauliangensis Gupta 19 Subgenus Euseius Wainstein 20 Key to the species of Euseius A. (Euseius) aizawai Ehara A. (E.) alstoniae Gupta A. (E.) bambusae Ghai & Menon A. (E.) coccineae Gupta A. (E.) coccosocius Ghai & Menon A. (E.) delhiensis (Narayanan & Kaur) A. (E.) eucalypti Ghai & Menon A. (E.) finlandicus (Oudemams) A. (E.) irregularis (Evans) A. (E.) neococcineae Gupta A. (E.) nicholsi Ehara & Lee A. (E.) notatus Chaudhri A. (E.) okumae Ehara & Bhandhufalck A. (E.) ovalis (Evans) A. (E.) pruni Gupta A. (E.) rhododendronis Gupta A. (E.) sacchari Ghai & Menon 29

7 [ iii ] 52. A. (E.) alangii Lai-rong & Li-sheng 53. A. (E.) densus Wei-nan 54. A. (E.) guangxiensis Wei-nan 55. A. (E.) macrospatulatus Gupta 56. A. (E.) mediocris (Chaudhri, Akbar & Rasool) 57. A. (E.) ruiliensis Wei-nan & Zhao-Quan 58. A. (E.) septicus (Chaudhri, Akbar & Rasool) 59. A. (E.) womersleyi Schicha 60. A. (E.) yunnanensis Wei-nan 61. A. (E.) relictus (Chaudhri, Akbar & Rasool) Subgenus Neoseiulus Hughes Key to the species of Neoseiulus 62. Amhlyseius (Neoseiulus) aceriae Gupta 63. A. (N.) aequipilus Berlese 64. A. (N.) anuwati Ehara & Bhandhufalck 65. A. (N.) assamensis (Chant) 66. A. (N.) baraki Athias-Henriot 67. A. (N.) cynodonae Gupta 68. A. (N.) calorai Corpuz & Rimando 69. A. (N.) Jallacis (Garman) 70. A. (N.) Jraterculus Berlese 71. A. (N.) imhricatus Corpuz & Rimando 72. A. (N.) indicus (Narayanan & Kaur) 73. A. (N.) longispinosus (Evans) 74. A. (N.) paspalivorus (De Leon) 75. A. (N.) rangatensis Gupta 76. A. (N.) siamensis Ehara & Bhandhufalck 77. A. (N.) stolidus Chaudhri 78. A. (N.) taiwanicus Ehara 79. A. (N.) venustus Chaudhri 80. A. (N.) amantis (Chaudhri, Akbar & Rasool) 81. A. (N.) bheraensis (Chaudhri, Akbar & Rasool) 82. A. (N.) caruncula (Chaudhri, Akbar & Rasool) 83. A. (N.) curvus Wei-nan & Zhao-Quan 84. A. (N:) eremicus (Chaudhri, Akbar & Rasool) 85. A. (N.) myrtia (Chaudhri, Akbar & Rasool) 86. A. (N.) reductus (Chaudhri, Akbar & Rasool) 87. A. (N.) tabularis (Chaudhri, Akbar & Rasool) Subgenus Paraphytoseius Swirski & Shechter Pages

8 [ iv ] Key to the species of Paraphytoseius 88. Amblyseius (Paraphytoseius) multidentatus (Swirski & Shechter) 89. A. (P.) scleroticus Gupta & Ray 90. A. (P.) cracentis (Corpuz & Rimando) Subgenus Phytoscutella Murna Key to the species of P hytoscutella 91. Amb/yseius (Phytoscutella) salebrosus (Chant) 92. A. (P.) wongsirii Ehara & Bhandhufalck Subgenus Proprioseiopsis Murna 93. Amblyseius (Proprioseiopsis) peltatus Van der Merwe 94. A. (P.) arunachalensis Gupta 95. A. (P.) beatus Chaudhri 96. A. (P.) bulga (Chaudhri, Akbar cst. Rasool) 97. A. (P.) dentatus (Chaudhri, Akbar & Rasool) 98. A. (P.) synachattiensis Gupta Subgenus Proprioseius Chant 99. Amblyseius (Proprioseius) kumaonensis (Gupta) Subgenus Typhlodromalus Muma Key to the species of Typhlodromalus 100. Amblyseius (Typhlodromalus) eucalypticus (Gupta) 101. A. (T.) jarooa Gupta 102. A. (T.) kalimpongensis Gupta 103. A. (T.) lablabi Ghai & Menon 104. A. (T.) manipurensis Gupta 105. A. (T.) newsami (Evans) 106. A. (T.) sorghumae Gupta 107. A. (T.) ehikmagalurensis Gupta 108. A. (T.) chitradurgae Gupta 109. A. (T.) ficusi Gupta 110. A. (T.) laaensis Gupta Subgenus Typhlodromips De Leon Key to the species of TYphlodromips 111. Amblyseius (Typhlodromips) arecae Gupta 112. A. (T.) asiatieus (Evans) 113. A. (T.) erotalariae Gupta 114. A. (T.) eujeniae Gupta 115. A. (T.) ghan;; (Murna) Pages

9 [ v 1 Pages 116. A. (T.) guajavae Gupta A. (T.) hidakai Ehara & Bhandhufalck A. (T.) johoreae (Muma) A. (T.) lenis Corpuz & Rimando A. (T.) labis Corpuz & Rimando A. (T.) Iinearis Corpuz & Rimando A. (T.) mangiferae Ghai & Menon A. (T.) meghalayensis Gupta A. (T.) neocrotalariae (Gupta) A. (T.) okinawanus Ehara A. (T.) officinaria Gupta A. (T.) perfectus Chaudhri A. (T.) polyantheae Gupta A. (T.) potentillae (Garman) A. (T.) sapienticola Gupta A. (T.) simi/is (Koch) A. (T.) suknaensis Gupta A. (T.) syzygii Gupta A. (T.) tetranychivorus (Gupta) A. (T.) vignae Lai-rong & Li-sheng A. (T.) baiyunensis Wei-nan A. (T.) draconis (Chaudhri, Akbar & Rasool) A. (T.) neoghanii Gupta A. (T.) oguroi Ehara A. (T.) repletus Wei-nan & Zhao-Quan A. (T.) sijiensis Gupta 71 Genus Amathia Chaudhri, Akbar & Rasool Amathia rancidus Chaudhri, Akbar & Rasool 71 Genus Indoseiulus Ehara lndoseiulus ricini (Ghai & Menon) Indoseiulus eharai Gupta 72 Genus Ipbiseius 72 Key to the species of Iphiseius Jphiseius (/phiseius) andamanicus Gupta Iphiseius (/phiseius) hapoli Gupta Iphiseius (Trochoseius) bakeri Gupta 73 Genus Paraamblyseius) Muma 74 Key to the species of Paraamblyseius Paraamblyseius fl'agariae Gupta... 74

10 [ vi ] 149. P. formosanus (Ehara) 150. P. mumai Gupta Genus Plat),seiella Muma 151. Platyseiella mumai Ray & Gupta 152. P. formosanus Tseng Genus Okiseius Ehara Key to the species of Okiseius 153. Okiseius subtropicus Ehara 154. o. chinensis Wei-nan 155. o. himalayana Gupta 156. o. sikkimensis Gupta 157. O. yazuliensis Gupta Subfamily Phytoseiinae Key to the genera of Phytoseiinae Genus Indodromus Ghai & Menon 158. Indodromus meerutensis Ghai & Menon Genus Pbytoseius Ribaga Key to the subgenera of Phytoseius Subgenus Pennaseius Pritchard & Baker Key to the species of Pennaseius Pritchard & Baker 159. Phytoseius (Pennaseius) curvatus Chaudhri 160. P. (P.) diutius Corpuz 161. P. (P.) hongkongensis Swirski & Shechter 162. P. (P.) improcerus Corpuz 163. P. (P.) kapuri Gupta 164. P. (P.) minutus Narayanan, Kaur & Ghai 165. P. (P.) incisus Wei-nan & Zhao-Quan 166. P. (P.) amba Pritchard & Baker 167. P. (P.) l1amdaphaensis Gupta 168. P. (P.) taianensis Lai-rong & Li sheng Subgenus Phytoseius Ribaga Key to the species of P hytoseius 169. Phytoseius (Phytoseius) bandipurensis Gupta 170. P. (P.) brevicrinis Swirski & Shechter 171. P. (P.) coheni Swirski & Shechter 172. P. (P.) corniger Wainstein 173. P. (P.) crinitus Swirski & Shechter 174. P. (P.) glareosus Corpuz 175. P. (P.) hawaiiensis Prasad... Pages

11 [ vii ] Pages 176. P. (P.) huangi EJJara P. (P.) intermedius Evans & Macfar!ane P. (P.) jujuba Gupta P. (P.) macropilis (Banks) P. (P.) macrosetosus Gupta P. (P.) mansehraensis Chaudhri P. (P.) meyerae Gupta P. (P.) mixtus Chaudhri P. (P.) indicus Bhattacharyya P. (P.) neocorniger Gupta P. fp.) neoferox Ehara & Bhandhufalck P. (P.) punjabensis Gupta P. (P.) rachelae Swirski & Shechter P. (P.) ras;lis Corpuz P. (P.) rimandoi Corpuz P. (P.) rugosus Denmark P. (P.) roseus Gupta P. (P.) spathulatus Chaudhri P. (P.) swirskii Gupta P. (P.) wainsteini Gupta P. (P.) domesticus Rather P. (P.) longus Wei-nan & Zhao-Quan P. (P.) nipponicus Ehara P. (P.) petentis Chaudhri, Akbar & Rasool P. (P.) qianshanensis Lai-rong & Li.. sheng P. (P.) rugatus 'f seng P. (P.) ruidus Wei-nan & Zhao-Quan P. (P.) silvaticus Wei-nan & Zhao-Quan 99 Genus Typhlodromus Scheuten 100 Key to the subgenera of Typhlodromus Subgenus Amblydromella Muma 101 Key to the species of Amblydromella Typhlodromus (Amblydromella) acacia lie-liu, Lai-rong & Li-sheng T. (A.) balakotiensis (Chaudhri, Akbar & Rasool) T. (A.) bambusicolus Gupta T. (A.) cordiae (Muma) T. (A.) chrysanthemi Gupta T. (A.) dactylifera (Chaudhri, Akbar & Rasool) 104

12 [ viii ] Pages 210. T. (A.) divergentis (Chaud~ri, Akbar & Rasool) T. (A.) darjeelingensis Gupta T. (A.) eremitidis (Chaudhri, Akbar '& Rasool) T. (A.) fleschneri Chant T. (A.) galummatus (Chaudhri, Akbar & Rasool) T. (A.) ghanii (Mum a) T. (A.) gopali Gupta T. (A.) himalayensis Gupta T. (A.) homalii Gupta T. (A.) kodaikanalensis Gupta ' T. (A.) loralaiana (Muma) T. (A.) mori Gupta T. (A.) octavu~ (Chaudhri, Akbar & Rasool) T. (A.) philippinus Corpuz T. (A.) rhenanus (Oudemans) T. (A.) rhododendroni Gupta T. (A.) ryukyuensis Ehara T. (A.) serrulatuj Ehara T. (A.) sica (Chaudhri, Akbar & Rasool) T~ (A.) umbratus (Chaudhri, Akbar & Rasool) T. (A.) zafari Chaudhri T. (A.) arrunachalensis Gupta T. (A.) husaini Khan T. (A.) incasus (Chaudhri) T. (A.) jackmickley; T. (A.) libitus (Chaudhri) T. (A.) limitatus (Chaudhri, Akbar & Rasool) T. (A.) maracus (Chaudhri) r...(a.) nid,bu Pritchard & Baker T. (A.) nilgiriensis Gupta T. (A.) ornatulus (Chaudhri) T. (A.) platycladus Jie-liu, Lai-rong & Li-sheng T. (A.) serratus (Chaudhri) T. (A.) sonprayagensis Gupta 119 Subg~nus Anthoseius De Leon Typhlodromus (Anthoseius) majumderi Gupta 119 Subgenus Brethria Tuttle & Muma 120 Key to the species of Brethria Typhlodromus (Brethria) confusus Narayanan, Kaur &- Ghai 120

13 [ ix ] Pages 246. Typhlodromus (Brethria) roshanlali Narayanan & Ghai 120 Subgenus Clavidromus Muma Typhlodromus (Clavidromus) neotransvaalensis Gupta 121 Subgenus Chiliseius Gonzalez & Schuster Typhlodromus (Chiliseius) lieni (Tseng) T. (C.) separatus T. (C.) tengi Wei-nan & Zhao-Quan 122 Subgenus Metaseiulus Muma 251. Typhlodromus (Metaseiulus) contiguus Chant Subgenus Orientiseius Muma & Denmark 122 Key to the species of Orientiseius Typhlodromus (Orientiseius) agilis (Chaudhri) T. (0.) channabasavannai Gupta T. (0.) hadii Chaudhri T. (0.) lalazariensis (Chaudhri) T. (0.) manipurensis Gupta T. (0.) orissaensis Gupta T (0.) pruni Gupta T. (0.) rickeri Chant T (0.) samliensis (Chaudhri) T. (0.) thailandicus Ehara & Bhandhufalck 127 Subgenus Paraseiulus Muma 127 Key to the species of P araseiulus Typhlodromus (Paraseiulus) comptus Corpuz T. (P.) ignavus (Chaudhri, Akbar & Rasool) T (P.) neosoleiger Gupta T (P.) operantis (Chaudhri, Akbar & Rasool) T (P.) trimediosetus lie-liu, Lai-rong & Li-sheng T. (P.) vitreus (Chaudhri, Akbar & Rasool) 130 Subgenus Typhloctonus Muma 130 Key to the species of Typhloctonus Typhlodromus (Typhloctonus) nesbitti Womersley T (T.) transitans Gupta T (T.) vol/sella (Chaudhri, Akbar & Rasool) T. (T.) prunus (Denmark & Rather) 132 Subgenus Typhlodromus Scheuten Key to the species of Typhlodromus 272. T. (T.) communis Gupta

14 r x ] 273. T. (T.) garhwalicus Gupta 274. T. (T.) neorhenan21s Gupta 275. T. (T.) disparis (Chaudhri, Akbar & Rasool) 276. T. (T.) barker; (Hughes) 277. T. (T.) sijiensis Gupta Subfamily Gnoriminae Chaudhri Key to the genera of Gnoriminae Genus Garhwalicus Gupta & Ray 278. Garhwalicus himalayensis Gupta & Ray Genus Gnorimus Chaudhri 279. Gnorimus tabella Chaudhri SUMMARY REFERENCES ApPENDlX-I ApPENDIX-II ADDENDA INDEX.., List of phytoseiid 'species known from Nepal, Bhutan, Burma and Bangladesh.... Distribution of Phytoseiidae in different countries of the Oriental region Pages 13~ ~: > 1: t t 133'

15 A TAXONOMIC REVIEW OF ORIENTAL PHYTOSEIIDAE WITH KEYS TO GENERA AND SPECIES S. K. GUPTA Zoological Survey of India Calcutta INTRODUCTION The mites of the family Phytoseiidae have been recognised for over seventy five years as potential predators of phytophagous mites (Parrott et al., 1906; Gilliatt, 1935; Huffaker & Spitzer, 1951; Fleschner & Ricker, 1954 ; Chant, 1959a; McMurtry & Scriven, 1964, 1964a; McMurtry et al., 1970; Huffaker et al., 1970; Van de Vrie et al., 1972 ; Kamath, 1968 ; Mallik, 1974 ; Puttaswamy 1978 ; Jagadish & Nagesha Chandra, 1979; Somchoudhury, 1979, 1981; Channa Basavanna, 1981, 1981a ; Mallik & ChannaBasavanna, 1983, etc.) and it has been proved beyond doubt that these mites, if properly and judiciously utilised, can bring effective and profitable biological control of phytophagous mites. In view of this importance, the phytoseiid mites have drawn worldwide interest and considerable amount of work has been done all over the world on the exploration of fauna, resulting in the discovery of over 800 species. So far as the Oriental region is concerned, 279 species in 11 genera and 3 subfamilies are known, represnting 142 species from India (Narayanan & Khot, 1959; Narayanan & Kaur, 1960; Narayanan et al., 1960; Chant, 1960; Narayanan & Ghai, 1964; Ghai, 1964 ; Rao & Rao, 1964; Ghai & Menon, 1967, 1969; Muma,-1967; Bhattacharyya, 1969 ; Gupta ; Ray & Gupta, 1981, 83 ; Gupta & Dhooria, 1972, 1974; Gupta et al., 1971; Gupta & Nahar, 1981; Gupta & Ray, 1981 ; 1981a; Ghai & Gupta, 1984 ; Rather, 1985 ; Denmark & Rather, 1984), 65 species from Pakistan (Chaudhri, 1965, 1967, 1968, 1973, 1975, 1975a, 1975b ; Chaudhri et al., 1974, 1979; Denmark, 1966 ; Muma, 1967 and Kazimi & Ghani, 1971), 29 species from Thailand (Ehara & Bhandhufalck, 1977; Ehara, 1977), 27 species from Taiwan (Ehara, 1970; Lo, 1970; Tseng, 1976, 1977, 1984; Tjying, 1977; Chang & Tseng, 1978; Shih, 1984; Ehara, 1977), 21 speciea from Philippines (Chant, 1959; Corpuz, 1966; Corpuz & Rimando, 1966 ; Swirski & Golan, 1969 ; Schicha & Corpuz-Raros, 1985), 15 species

16 2 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 from Indonesia (Chant, 1957; Chant, 1959a; Evans, 1952, 1953; Oomen, 1982), 6 species from Malayasia (Evans, 1952, 1953; Muma, 1967 ; Ehara, 1977) and 40 species from South China (Jie-Liu et. al., 1980; Lai-rong & Li-sheng, 1981; Lai-rong & Li-sheng, 1981a; Wei-nan, 1984, 1984a ; Wei-nan & Zhao-Quan, 1984, 1985, 1985a; Wei-nan & Xing, 1983, 1983a). So far the information is available to the author, nothing is known from other parts of the Oriental region. Ehara (1966) catalogued the Asian phytoseiid mites and Prasad (1974) catalogued the Indian mites. The former included 36 species from Oriental region and the latter included 58 species of Phytoseiidae from India. Gupta (1974) published a brief review of Indian Phytoseiidae which also included 58 species. Since the publication of Ehara's catalogue in 1966 and till the end of 1985 several species were added from the Oriental region and it was felt necessary to have a consolidated account of this important group of mite from this region. With this view in mind, all the 279 species are reviewed here giving their relevant synonymy, diagnosis, habitat records, type material, type repositories, distribution, etc. and also giving the keys to the subfamilies, genera, subgenera and species, so as to give the taxonomists and workers in the applied field a ready reference to Oriental Phytoseiidae. The measurements given in the text are in microns. The abbreviations used for different Museums while giving type repositories are as below :- ZSI = Zoological Survey of India, Calcutta. BM(NH)::: British Museum Natural History, London. USNM UAL United States National Museum, Washington D. C. Dept. of Entomology, University of Agriculture, Lyallpur, Pakistan. NPC - National Pusa Collection, IARI, New Delhi. Dept. Ento. Univ. Philip = Dept. of Entomology, University of Philippines. Univ. Inst. Agr. Israel - Dept. of Entomology, National and University Institute of Agriculture, Rehovot, Israel. BioI. Inst. Fac. Ed. T ott. U niv, = Biological Institute, Faculty of Education, Tottori University, Japan. Zoo!. Inst. Fac. Sci. Hokk. Univ. = Zoological Institute, Faculty of Science, Hokkaido University, Japan.

17 GUPTA: ORIENTAL PHYTOSEIIDAE 3 Rijksmus. Van. Nat. Hist. = Rijksmuseum Van Natuurlijke Historie, Leiden. UAF Dept. of Entomology, University of Agriculture, Faisalabad, Pakistan. Though sincere efforts were made to make this review a most up-todate one but yet there may be some lacunae and it would be highly appreciated if those are brought to the notice of the author for rectification in future. The author is highly grateful to Dr. B. K. Tikader, Director, Zoological Survey of India, Calcutta, for providing the facilities and encouragements and to the numerous fellow scientists allover the world who helped the author with reprints of their works and also with many valuable suggestions. FAMILY PHYTOSEIIDAE Diagnosis: Dorsal shield entire or divided into 2 subequal shields with 20 or fewer pairs of setae (except in Gnoriminae which has 25 pairs of setae); never more than 2 pairs of sublateral setae (except in Gnoriminae which has 3 pairs); all setae on dorsum either simple, serrate or club-shaped or otherwise shaped. Peritreme extends anteriorly from mesolateral stigmata. Sternal shield well to poorly sclerotized with 2-4 pairs of setae, some may not be on shield, but present either on integument or on platelets. Genital shield truncate with a pair of setae. Ventrianal shield always present (except in Indoseiulus) with 2-4 pairs of preanal setae (Gnoriminae has 6-7 pairs in addition to para and postanal setae). Fixed digit of chelicera well developed, movable digit with or without teeth. Males have spermatophoral process on the movable digit of chelicera. Type genus: Phytoseius Ribaga, 1904 Out of 4 subfamilies, viz. Amblyseiinae Muma, Phytoseiinae Berlese, Gnoriminae Chaudhri and Macroseiinae Chant, Denmark & Baker, known under the family Phytoseiidae, the first three are represented from the Oriental region and those can be separated with the help of the following key. Key to the subfamilies of Phytoseiidae 1. Dorsal shield with 12 pairs of lateral setae; sublateral setae 3 pairs, preanal setae on ventrianal shield 6 pairs Gnoriminae

18 4 REe. ZOOL. SURV. INDIA, OCC. PAPER No. 9S Dorsal shield with 8-11 pairs of lateral setae; sublateral setae 2 pairs, preanal setae 2-4 pairs 2. Four pairs of prolateral setae Five or six pairs of prolateral setae 2 Amblyseiinae Phytoseiinae Diagnosis: Subfamily AMBLYSBIINAE Muma Muma & Denmark (1970) diagonised this subfamily as :-dorsal shield undivided with 4-6 pairs of dorsocentral setae, 1-2 pairs of median setae, 1-4 pairs of prolateral setae out of total of 6-9 pairs. Macrosetae 1-3 on leg. IV. Males with fragmented or entire ventrianal shield with 3-4 pairs of preanal setae and usually 2 pairs of sublateral setae. Type genus Amh/yseius Berlese, Key to the genera of Amblyseiinae 1. Ventrianal shield absent, only preanal setae present Ventrianal shield present 2. R 1 on dorsal shield Rl on lateral integument 3. Lateral integument sclerotized so that ra and Rl though on usual lateral position are on lateroventral extension of dorsal shield Lateroventral extension not sclerotized, as above 4. Metapodal plates large, triangular, genital shield very broad and punctate, ventriana I shield massive, genu III with six setae Metapodal plates normally paired, slender, elongate, genital and ventrianal shields usually narrow, gen u III with 7 setae Indoseiulus 2 Okiseius 3 Iphiseius 4 Paraamblyseius Amblyseius Genus Amblyseius Berlese Amblyseius Berlese, Redia, 10: Amblyseius (Seiopsis) Berlese, Redia, 15 : Amblyseius (Amblyseiopsis) Garman, Bull. Conn. agrie. Exp. SIn., 520: Kampimodromus Nesbitt, Zoo/. Verk., 12 : Anlblyseiopsis Muma, Ann. ent. Soe. Am., 48 : Amblyseiella Muma, ibid, 48: Proprioseius Chant, Can. Ent., 89 : Asperoseius Chant, ibid, 89 : Typhlodromus (Amb/yseius) Chant, Call. Ent., 89 : Phyllodromus De Leon, Ent. News., 70 : Typhlodromus (Typhlodromopsis) De Leon, Fla. Ent., 42: Phytoseutus Muma, Bull. Fla St. Mus., 5 : Phytoseutella Muma, ibid,s: Amblyseiulella Muma, ibid, 5 ~ 276.

19 GUPTA: ORIENTAL PHYTOSEIIDAE Proprioseiopsis Muma, ibid,s: Amblyseiulus Muma, ibid,s: Amblyscutus Muma, 5 : Paradromus Muma, ibid,s: Cydnodromus Muma, ibid,s: Phytodromus Muma, ibid,s: Typhloseiella Murna, idid, 5 : Typhloseius Muma, ibid,s: Athiaseius Wainstein, Acarologia, 4: Paraphytoseius Swirski & Shechter, Israel J. agric. Res., 11: Amblyseius (Ptenoseius) Pritchard & Baker, Hi/gardia, 33 : Ansaria Chaudhri, Akbar & Rasool, Rept. PL 480 Project, Univ. Agril. Faisalabad, Pakistan, p. 63 (New synonymy) Kashmiricus Chaudhri, Akbar & Rasool, ibid., p. 66 (New synonymy) Amblyseius, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 33. Diagnosis: Chant (1965) characterized the genus as: dorsal shield with pairs of setae, of those, never more than 4 pairs present on prolateral series. Setae of diverse length, sublateral setae (r 3, R 1 ) on lateral integument (except Okiseius where R1 on dorsal shield). Sternal shield with 3 pairs of setae; 4th pair on metasternal plate or on integument. Genital shield broad or narrow, truncate with a pair of genital setae. Ventrianal shield of varied shape, with 3 pairs of preanal setae; 4 pairs of setae present on membrane around ventrianal shield; sometimes may be 3 pairs. Metapodal plates present or absent. Anal opening ventral in position. Tritosternum present. Fixed digit of chelicera normally with teeth and pilus dentilis; movable digit with or without teeth. Peritrematal shield fused anteriorly with dorsal shield; length variable. Legs often with macrosetae on leg IV ; genu II 6-7 setae; genu III 6-7 setae. Tibia II 6 7 setae; tibia III 6-7 setae ; tibia IV 6 setae. Type: Zercon obtusus Koch, 1839 Key to the subgenera of Amhlyseius 1. Dorsal shield with 5 pairs of postlateral setae Dorsal shield wi th less than 5 pairs of postlateral setae 2. Seta Z 1 presen t Seta Z, absent. 3. Seta S; present Seta S~ absent 4. Ventrianal shield massive covering major portion of posteroventral portion Ventrianal shield not like above Proprioseius Asperoseius Paraphytoseius 5 6

20 6 REC. ZOOL. SURV. INDIA, Occ. PAPER No Seta j.5 present Seta j r; absent 6. Setae Zr; and S4 and or Z" long and whup-like, longer than distance between their bases; leg IV with macroseta on genu and erect seta on tarsus I Seta Zr; shorter, at most as long as distance between their bases; leg I with no macroseta or only one on genu, no erect seta on tarsus I 7. Sternal shield distinct and straight or concave posteriorly, ventrianal shield approximately shieldshaped or pentagonal Sternal shield indistinct, may be trilobate posteriorly, ventrianal shield elongate, vase-shaped or concave laterally 8. Zr; and Z" distinctly serrate, sternal shield as wide as or wider than long, macroseta may be present on genu I and present on genu II and III Zr; and Z4 indistinctly serrate, sternal shield longer than wide; genu I, II and III without macroseta 9. Peritreme extends upto j s ; anterior. pair of preanal setae adjacent to anterior margin of ventrianal shield Peritreme not extends upto j,; anterior pair of preanal setae removed from anterior margin of ventrianal shield Propl'ioseiopsis Phytoscutella Amblyseius Typhlodromips Neoseiulus Typhlodromalus Euseius Subgenus Amblyseius Berlese Key to the species of Amblyseius (Amblyseius) 1. Ventrianal shield vase-shaped, with lateral margins concave Ventrianal shield triangular, pentagonal or squarish 2. Spermatheca with cervix straight or of uniform length Spermatheca fundibuliform 3. S4 and Z" of same length Sol shorter than Z4 4. Zr; nearly double the length of Z" Z6 much less than double the length of Z" S. Ventrianal shield triangular Ventrianal shield pentagonal or squarish 6. Z 6 very long, more than 1/2 length of dorsal shield Zr; not so long 7. Cervix of spermatheca elongated or slender Cervix of spermatheca otherwise shaped 8. s" shorter than Z" S4 equal to or longer than Z" 9. S4 longer than Z4 Sol equal to Z4 2 S 3 4 largoensis adhatodae herbicolus imp actus obtusus mcmurtryi 8 cinctus 9 neorykei shoreae

21 GUPTA: ORIENTAL PHYTOSEIIDAE 10. Sol slightly shorter than Z 4 s" noticably shorter than Z" 11. Spermatheca with cervix long, broad, dorsal shield faintly reticulate Spermatheca with cervix narrow, tubular dorsal shield smooth 12. Cervix of spermatheca looped Cervix of spermatheca almost straight 13. Z:a longer than z" Z:a as long as z" 14. Cervix long, tubular Cervix wide, funnel-shaped challnabasavannai kulini paraaerialis 13 aerialis 14 orientalis muraleedharani 1. Amblyseius (Amblyseius) adhatodae Muma Amblyseius adhatodae Murna, Fla.Ent., SO : (Holotype ~, Paratypes 10, 1,larve, Pakistan, Karachi, on Adhatoda vasica, deposited in USNM) ; another ~, India : Bombay, Ipomoea sp Amblyseius ipomoeae Ghai & Menon, Oriental Ins., 1 : (Bombay: Ipomoea sp., infested with tetranychids) Amblyseius (Amblyseius) adhatodae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 37. Diagnosis: Dorsal shield 370 long, 275 wide, lightly sclerotized with 17 pairs of setae; j1' js, S4' Z5' Z4' long; Z4>S4' jl greater than 2 times of Za and Z4. Fixed digit of chelicera with teeth, movable digit with 1 tooth, leg IV with macroseta on genu, tibia and basitarsus Cervix of spermatheca slender and elongate. Spermatophoral process a short shank, short foot and anteriorly bent toe. India (Bombay), Pakistan. Remarks: On examination of the type of A.(A.) ipomoeae it appeared that ipomoeae and adhatodae are conspecific. 2. Amblyseios (Amblyseius) aerialis (Muma) Amblyseiopsis aerialis Muma, Ann. ent. Soc. Am., 48 : (Holotype ~, USA: Florida, Lucern Park, on citrus, in USNM) Amblyseius aerialis, Rao & Rao, Comm. Inst. Bioi. Contra Tech. Bul/., 4 : (Karnataka: citrus) Amblyseius aerialis, Gupta, Internat. J. Acaro/., 1 (2): 31. (Karnataka: citrus) Amblyseius aerialis, Gupta & Nahar. In Contrib. to A car. in India, p. 9., (Bihar: citrus) A. (A.) aerialis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 39.

22 8 REC. ZOOL. SURV. INDIA, Occ. PAPER No. 95 Diagnosis: Dorsal shield with 17 pairs of setae of those is, S4' Z5' Z4' being long measuring, respectively, 55, 120, 288 and 144. Ventrianal shield rectangular, 115 long, 75 wide, with 3 pairs of preanal setae and a pair of elliptical preanal porse. Cervix of spermatheca tubular. Macrosetae on leg IV : genu-132, tibia-100, basitarsus-76. Spermatophoral process with small lateral process, lobate heel and club-shaped foot. USA, India (Karnataka, Bihar). Remarks: It has been reported to feed on citrus mite (P.eitri) (Rao & Rao, 1964). 3. Amblyseios (Amblyseius) channabasavannai Gupta Amblyseius channabasavanni Gupta, Oriental Ins., 12: (Holotype ~,India : Kerala, Trivendrum, on Chrysanthemum; Paratypes 1 ~, 1 ~, Tamil Nadu, Oothu, on dahlia, all in ZSI) A. (A.) channabasavannai, Gupta, Fauna of India, (Acari: Mesostigmata) Family Phytoseiidae, p. 41. Diagnosis: Dorsal shield smooth, with 17 pairs of setae, Z5-212, S4-72, js-40, jl-28, other setae minute. Sternal shield weakly sclerotized with 3 pairs of sternal setae, metasternal plates present. Ventrianal shield pentagonal, much longer (108) than broad (80), with 3 pairs of preanal setae: 2 pairs or" metapodal plates present. Fixed digit of chelicera multidentate, 4 teeth on movable digit. Spermatheca, with tubular cervix. Macrosetae on leg IV : genu-80, tibia-60, basitarsus-52. shorter. Distribution t Chaetotaxy on dorsum similar as in female except being India (Kerala, Tamil Nadu). Remarks: Its biology and predatory importance were studied bv Daniel (1979, 1981). 4. Amblyseios (Amblyseios) cinctos Corpuz & Rimando Alnblyseius cinctus Corpuz & Rimando, Philipp. Agric., SO: (Holotype ~, Philippines: Gamu, Isabela, on Panicum pi/ipes; Paratypes 2 ~ ~, Echague, Isabela, on Streb/us asper, all deposited in Dept. Ento. Univ. Philip.) A. (A.) cinctus, Ebara & Bhandhufalck, J. Fac. Ed. Tottor; Univ. Nat. Sci 27 (2) : 70-71, (Redescribed). (Thailand: on Acaripha sp., grape, mulberry, juniper, citrus, bamboo. longan, Sida rhombi/olia, litchi, papaya, Casuarina, Hibiscus).

23 OUPTA: ORIENTAL PHYTOSEnOAE 9 Diagnosis; Dorsal shield smooth, with 17 pairs of setae and 7 pairs of pores; ii' is, S4' Z5' Z4 long, other setae small. Measurements of setae: j1-19, i 4 -i s l , 4-5 each, js-35, Za, Z46-8 each, s4-51, Z1 1 Sa-SIS 6-7 each, Z5-109, z5-5, Z4-57. Ventrianal shield pentagonal, longer than wide, lateral margins concave, with 3 pairs of preanal setae and a pair of pores. Chelicera with fixed digit multidentate, movable digit with 4 teeth. Macrosetae on leg IV: genu-79, tibia-47, basitarsus-60 ; cervix of spermatheca tubular. Chaetotaxy of male similar to that of female, setae smaller. Thailand, Philippines. 5. Amblyseins (Amblyseius) herbiocolus (Chant) Typhlodromus (Amblyseius) herbicolus Chant, Can. Ent., 91 : Amblyseius (Amblyseiulus) largoensis, Murna, Bull. Fla. St. Mus., 5 (7) : 287 (not largoensis Murna, 1955) Amblyse;us deleon; Muma & Denmark, Arthropods of Florida, 6: (Holotype ~,U.S.A. : Florida, Citrus, in USNM) A. (A.) deleon; Ehara & Bhandhufalck, J. Fac. Ed. Tottori Unrv. Nat. Sci., 27 (2) : (Thailand: citrus; Preu, on rambutan, santol, coffee, bamboo, Gendarussa vulgaris, pine, tea) A. (A.) herbicolus, Gupta, auna of India (Acari: Mesostigrnata) Family Phytoseiidae, p. 45. Diagnosis: Dorsal shield smooth, 357 long, 255 wide, with 17 pairs of setae, jl-35, ja-33, s4-107, Z5-205, Z4-125 ; other setae small. Ventrianal shield 116 long, 67 wide, with 3 pairs of preanal setae. Fixed ;digit of chelicera with 5 teeth movable digit with 3 teeth. Macrosetae on leg IV: genu-129, tibia-90, basitarsus-76. Spermatheca with fundibuliforn cervix and greatly swollen atrium. Spermatophoral process L-shaped, with a weakly bifurcate toe on the foot. U.S.A., Thailand, India (Tripura). 6. Amblyseius (Amblyseins) impactus Chaudhri Amblyseius impactus Chaudhri, Acar%gia, 10 : (Holotype ~, Pakistan: North Sheikhupura, Eriobotrya japonica, in UAL.). Diagnosis: Dorsal shield with 17 pairs of setae, smooth. Measurements of setae: jl-34, j4' i 6 J 2 -minute, J 5-10, ja 51, za-z4 minute, 7 -each, s4-107, Zl-10, Sa-14, S4-14, S5-10, Z5-239, z5-5, Z4-120, ra-10, R Ventrianal shield 131 long, 78 wide, with deep notch just below 2

24 10 REC. ZOOL. SURV. INDIA, Oec. PAPER No. 9S preanal setae, with 3 pairs of preanal setae ; 2 pairs of metapodal plates present. Chelicera with 9 subapical teeth on the fixed digit and pilus dentilis; one tooth on the movable digit. Macrosetae on leg IV ; genu-119, tibia-7s, basitarsus-56. Pakistan. 7. Amblyseius (Amblyseius) kulini Gupta Amblyseius kulini Gupta, Indian J. Acar., 2 (2) : (Holotype ~,India : Assam, Oossaigaon, on Bambusa sp., in ZSI) A. (A.) kulinl, Gupta, Fauna of India (Acari: Mesostigmata) Family Phyta. seiidae, p. SO. Diagnosis: Dorsal shield 300 long, 220 wide, with 17 pairs ofsetae. Excepting setae jl' js, 84, Z5 and Z4 which measure respectively 28, 44, 80, 200 and 84, others are small. Sternal, metastemal and genital shields normal. Ventrianal shield pentagonal, nearly Ii times as long as wide with 3 pairs of preanal setae. Cervix of spermatheca, tubular. Macrosetae on leg IV : genu-80, tibia-64, basitarsus-62. India (Assam). 8. Amblyseius (Amhlyseius) largoensis (Muma) Amblyseiopsis largoensis Muma, Ann. ent. Soc. Am., 48 : 266. (Holotype ~, USA : Florida, Largo, on key lime, in USNM) Amblyseius largoensis, Rao & Rao, Comm. Insf. Bioi. Contr. Tech. Bull., 4,J (Karnataka: citrus) A. largoensis, Corpuz & Rimando, Philipp. Agric., 50 : (Philippines: Casuarina equiseti/o/ia, Thuja orientalis, fern, Dieffenbachia piela, Cinnamomum camphora, Schizostachyum lima, Citrus nobilis, Cord),line frulicosa, Rhaphidophora aurea, Premna odorata, Cocos nuci/era, Ipomoea triioba) A. largoensis, Gupta, Sci. & Cult., 36 : 98. (West Bengal: Caiophyllum inophyl/um) A. (A.) largoensis, Ebara, Mushi, 43 (6) : 58. (Taiwan: on bamboo, mango, papaya, guava, avocado, Codiaeum variegalum, Capparis formosana, Paederia chinensis, Myristica cag2yaneas;s, Caryola milis, Reevesia /ormosana) Amblyseius iargoensis, Gupta et. al., Sci & Cult., 37 : 298. (Himachal Pradesh: on an undet. plant infested with T. cinnabarinus) A. largoensis, Gupta, Ent. Rec., 86: A. largoensis, Gupta, Internat. J. Acarol., 1 (2) : 38. (West Bengal: Mangifera indica, Ficus sp., Caiophyllum inophyl/um, Musancka corymbosa, 1abernaemontana coronaria).

25 GUPTA: ORIENTAL PHYTOSEllDAE A. largoensis, Lo, Bull. Sun- Yat Sen Cult. Found." 5: (Taiwan: Chayi : on bean with Tetranychus sp; Wulai, on citrus with P. citri ; F uoshan on chinese hibiscus with Tetranyehus sp. ; on purple grandilla with Tetranyehus sp. ; peach with Tetranychus sp. and P. citri ; an undet. plant with Eotetranychus sp.) A. (A.) largoensis Ehara & Bhandhufalck, J. Fac. Ed. Tottori Univ. Nat. Sci., 27 (2) : 67. (Thailand: on Citrus, pomela, casuarina, Lagerstroemia, coconut) A. largoensis, Gupta, Oriental Ins., 11 : (Andaman & Nicobar IsIs: Eucalyptus, bamboo, coconut, Tabernaemontana, arecanut, mango, Eugenia, Tectona, fig, guava, cashewnut, pepper, citrus) A. amtalaensis Gupta, Entomdogists' mon. Mag., 112: 53. (Holotype ~, West Bengal, on Citrus medica, in ZSI) A. largoensis, Gupta, Indian J. Acar., 1 : 33. (Gujarat : china rose, Nerium) A. largoensis, Gupta, Indian J. Acar., 2 (2) : (Orissa: cashew, litchi; Meghalaya: banana, plum, Manglietia insignis, chilli; Assam: undet., Shorea sp.; Tripura: papaya, Musa, Bauhinia acuminata; Manipur : poppy, Nagaland: undet.) A. largoensis, Gupta, Oriental Ins., 12: (Tamil Nadu: Castor, banana, citrus, coconut, pomegranate, Do/berg;a sp., eucalyptus, Teetona grandis; Pondicherry: black berry, sugarcane; Kerala : coconut, rose; Karnataka: Cassia, sugarcane; Andhra Pradesh: cashewnut) A. largoensis, Gupta & Nahar, In COlltrib. to A car. in Indio, p. 9 (Bihar: Mango) A. (A.) largoensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 51. Diagnosis: Dorsal shield smooth with 17 pairs of setae. All being small except i 1, js, S4' Z5 and Z4 which measure respectively 34, 48, 93, 270 and 95. Sternal, metasternal and genital shields normal. Ventrianal shield vase-shaped, widest at preanal level, with 3 pairs of preanal setae and a pair of pores. Macrosetae on leg IV ; genu-11s, tibia-90, basitarsus-62. Spermatheca with parallel sided cervix. Chaetotaxy of dorsum similar as in female. India (West Bengal, Assam, Orissa, Meghalaya, Manipur, Tripura, Nagaland, Andaman & Nicobar Isis., Himachal Pradesh, Punjab, Tamil Nadu, Pondicherry, Kerala, Karnataka, Andhra Pradesh, Bihar, Gujarat), U.S.A, Taiwan, Thailand, Philippines. Remarks: Predatory importance of this species was reported by Rao & Rao (1964), Gupta (1977) and Gupta (1981). 9. AmbJyseius (AmbJyseius) mcmurtryi Muma f167. Amblyseius mcmurtryi Muma, Fla. Ent., SO : 270. (Holotype ~, Paratype 1 ~, India : Assam, citrus, in USNM).

26 12 REC. ZOOL. SURV. INOlA, OCC. PAPER No A. mcmurtryi, Gupta, Tnternat. J. Acaro/., 1(2) : A. mcmurtryi, Gupta, Internal. J. Acaro/., 2(2) : 67. (Meghalaya: guava; Assam: citrus) A. (A.) mcmurtryi, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 55. Diagnosis: Dorsal shield 360 long, 208 wide, with 17 pairs of setae, all being minute except jl' j3' S4' Z5 and Z4 which measure: 25, 36, 58, 140 and 60, respectively. Sternal, metasternal and genital shields norma1. Ventrianal shield 108 long 80 wide, with 3 pairs of preanal setae and a pair of pores. Macrosetae on leg IV : genu-72; tibia-44 and basitarsus-56. Spermatheca with tubular cervix. India (Assam, Meghalaya). 10. Amblyseius (Amblyseius) neorykei Gupta Amblyse~us neorykei Gupta, Entomologists' mon. Mag., 112: (Holotype ~, India : Darjeeling, Sukna on Chrysanthemum sp. in ZSI) A. (A.) neorykei. Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 59. Diagnosis: Dorsal shield 325 long, 203 wide, with 17 pairs of setae and 6 pairs of pores. Measurements of setae jl-27, is-49, za, z4' ZI' Sa, S4-5 each; Z5-135, z5-4, Z4-85. Srernal shield with 3 pairs of setae and with posterolateral projections. Ventrianal shield 100 long, 81 wide, with 3 pairs cf preanal setae and a pair of preanal pores. Macrosetae on leg IV : genu-7 0, tibia-50, basitarsus-72. India (West Bengal). 11. Amblyseius (Amblyseius) obtusus (Koch) Zercon obtusus Koch, Fase. 27. (Holotype Italy, in moss and soil in Berlese's Collection, Florence) Amblyseius (Alnblyseius) obtllslls, Ehara, Mushi, 39 (2) : 22. Diagnosis: Dorsal shield with 17 pairs of setae; 54' Z5' Z4 long. Z4 = Za; Z4 == Sa) both minute. Ventrianal shield triangular with rounded converging lateral margins, with 3 pairs of preanal setae; leg IV with 3 macrosetae, one each on genu, tibia and basitarsus. Indonesia (according to Ehara, 1966), Europe, AIg~riat Australia, Canada.

27 GUPTA : ORIENTAL PHYTOSEDDAE Amhlyseius (Amhlyseius) orientalis Ehara Amblyseius sp. Ehara, Annotes Zool. Jap., 31 (1) : 55. '1959. Amblyseius orientalis Ehara, Acarologia, 1 : (Holotype ~, Paratypes 2 00, 15 ~ '.i, Japan: Sapporo, Hokkaido, on Quercus crispula, in Zool. Inst. Fac. Sci. Hokk. Univ.) A. orientalis, Gupta, Indialt J. Acar., 2(2) : (Assam: undet. plant) A. (A.) orientalis, Gupta, Fauna of India (Acari: Mesosti~mata) Family PhytoseH.da~, p. 61. Diagnosis: Dorsal shield 340 long, 245 wide. Measurements of setae: j1-28, ja-56, s4-95, Z5-280, Z4-112 ; other setae tiny. Ventrianal shield 124 long, 80 wide. Spermatheca with tubular cervix. Macroseteae on leg IV : genu-112, tibia-64, basitarsus-72. Dorsal chaetotaxy similar to that of female except being shorter. Spermatophoral process shallowly bifurcate at the distal end. India (Assam), Japan. 13. Amhlyseius (Amhlyseius) paraaerialis Murna Amblyseills paraaerialis Muma, Flat Ent., 50 : (Holotype ~, Paratypes 1 ~, India: Kerala, on citrus; 3 ~ ~, Assam, on citrus, all in USNM) Amblyseius paraaerialis, Gupta, Internat. J. Acarol., 1(2) : Amblyseiusparaaerialis Ehara & Bhandhufalck, J. Fac. Ed. Tottori Univ. Nat. Sci., 27 (2) : (Re-described). (Thailand: litchi, citrus, Pinus kesiya, Bridelia sp., longan, bamboo, Urena lobata, sugarcane, mango, guava, conifer, Eupatorium odoratum, tea, peach, persimmon) Amblyseius paraaerialis, Gupta, Indian J. Acar., 2 (2): (Meghalaya : undet. plant; Assam: undet. plant) A. (A.) paraaerialis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 63. Diagnosis: Dorsal shield 350 long, 210 wide, with 17 pairs of setae. Measurenlents of setae: i 1-32, j4' is, J 2' J 5' each, J 3-28, zs' z4t-6 each, s4-68, Z1-9, S2-12, S4,-16, S5-8, Z5-192, z5-6, Z4-84. Sternal, metasternal and genital shields normal. Ventrianal shield l0410ng, 84 wide, lateral margins slightly concave, with 3 pairs of preanal setae. Macrosetae on leg IV : genu-84, tibia-68, basitarsus-80. India (Assam, Meghalaya), Thailand. 14. Amblyseius (Amblyseius) muraleedharani Gupta Amblyseius rha!jdus, Muraleedharan & Chandrasekharan, Pestology, 5 : 13 (Tamil Nadu : Tea)-Misidentification A. (A.) muraleedharani Gupta, Env. & Ecol., 3(3) : 434.

28 14 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 95 Diagnosis: Dorsal shield 403 long, 247 wide, with 17 pairs of setae, jl' ja, S4' Z4' Z~ long measuring 27, 47, 103, 156 and 319, respectively, other setae small (5-6). Ventrianal shield pentagon~lj reticulate anteriorly, with 3 pairs of preanal setae and a pair of preanal pores. Fixed digit of chelicera multidentate, movable digit 2-dentate. Not known. India (Tamil Nadu). 15. Amblyseius (Amblyseius) shoreae Gupta Amblyseius shoreae, Gupta, Entomologists' mon. Mag., 112: 57. (Holotype ~,India : West Bengal, Midnapur, Jhargram, on Shorea robusta, in ZSI) A. (A.) shoreae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 65. Diagnosis: Dorsal shield 335 long, 205 wide, with 17 pairs of setae. Measurements of setae: jl-27, js-50, s4-60, Z~-98, Z4-60, other setae minute. Sternal shield as long as broad. Ventrianal shield 100 long, 84 wide, with 3 pairs of preanal setae and a pair of preanal pores. Macrosetae on leg IV : genu-54, tibia-44, basitarsus-37. India (West Bengal). In addition to the above species of Amblyseius (Amblyseius) the following species are also known from the Oriental region but those could not be included in the key either for want of adequate information or due to the late availability of information i.e. when the entire manuscript including the key was finalised and the manuscript was sent to the press. 16. Amblyseius (Amblyseius) bajulus Chaudhri, Akbar & Rasool Amblyseius bajulus Chaudhri, Akbar & Rasool, Rept. PI 480 Project, Univ. Agril. f aisalabad, Pakistan, p. 70. (Holotype ~, Pakistan: Kalan (swat), on undet. plant, in UAF; Para type 1 ~, Abbotabad l on Cardulls mutanus). Diagnosis: Dorsal shield 374 long, 302 wide, notched near R 1 Measurements of setae: j 1-23, s4t -88, Z~-260, Z4:-143, rs-lo, R 1-8 ; other setae minute. Macrosetae on leg IV: genu-114, tibia-94, basitarsus-70; other macrosetae: genu 1-36, genu 11-38, genu III-57, tibia

29 GUPTA: ORIBNTAL PHYTOSEUDAE 15 19l5S. Pakistan. 17. Amblyseius (Amblyseius) brevicervix: Wei-nan & Zhao-Quan Amb/yseius (Amh/yseius) hrevicervix Wei-nan & Zhao-Quan, Acta Zootaxonomica Sinica, 10(3) : (Holotype ~, Paratypes 3 ~ ~, South China: Mt. Emei, Sichuan Province, on Tanacelum vulgare). Diagnosis: Dorsal shield long, wide. Measurements of setae: i l -26, ;4-6, i 5-6, j6-7-s, Ja-S, Jfj-7, js-47, za-10, Z4-7, s , Zl-7-10, Sa-7-8, S4-7, S5-7, Z , z5-7, Z , ra-15, R 1-10, JV South China. 18. Amblyseius (Amblyseius) deductus Chaudhri, Akbar & Rasool Amblyseius deductus Chaudhri, Akbar & Rasool, Rept. PI. 480 Project, Univ. Agril. Faisalabad, Pakistan, p. 72. (Holotype ~, Pakistan: Charrapani, on Peristrophe, in UAF). Diagnosis: Dorsal shield 354 long, 270 wide. Measurements of setae: il-26, i 8-34, s4-86, Z5-270, Z Ventrianal shield 99 long, 103 wide. Macrosetae on leg IV: genu-120, tibia-62, basitarsus-68, other macro setae : genu 1-36, genu 11-29, genu III-55, tibia Pakistan. 19. Amblyseius (Amblyseius) excelsus Chaudhri, Akbar & Rosool Amblyseius excelsus Chaudhri, Akbar & Rasool, Rept. PI 480 Project, Univ. Agril. Faisalabad, Pakistan, p. 68. (Holotype ~, Pakistan: Abbotabad, on Ricinus communis, in UAF). Diagnosis: Dorsal shield 343 long, 239 wide, notched at the level of R 1, Measurements of setae: j1-31, js-49, za-18, Z4-21, 54-81, Zl-10, Sa' S4-10 each, ZI5-174, Z4-96, rs-21, R1-10. Ventrianal shield pentagonal, Macrosetae on leg IV : genu-75, tibia-63, basitarsus-75; other macrosetae: genu 1-29, genu 11-29, genu Spermatheca sacular. Uuknown. Pakistan.

30 16 REC. ZOOL. SURV. INDIA, Oce. PAPER No Amblyseius (Amblyseius) hainanensis Wei-nan & Xing Amblyseius (Amblyseius) hainanensis Wei-nan & Xing Entofl'Jotaxonomia, 5(3) : (Holotype ~,Paratypes 4 ~ ~, South China : Guangdong, Hainan, on citrus). Diagnosis: Dorsal shield 320 long, 212 wide. Measurements ~f setae: j , js-50-52, s4-75-~2, Z , Z4-100-l07, JV,, South China. 21. Amblyseius (Amblyseius) hapoliensis Gupta Amblyse;us (Amb/yseius) hapoliensis Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p (Holotype ~,Paratypes 3 ~ ~, India : Arunachal Pradesh, Hapoli, on apple, in ZSI). Diagnosis: Dorsal shield 342 long, 268 wide with 17 pairs of setae. Measurements of setae: j1-34, j4-43, s4-44, Z5-112, Z4-60, rs-16, R Macrosetae on leg IV : genu-80, tibia-62, basitarsus-63. India: Arunachal Pradesh. 22. Amhlyseius (Amblyseios) indirae Gupta Amblyseius (Amblyseius) indirae Gupta, Entomon, 10(3) : (Holotype ~, Paratypes 2 ~ ~, 1 <1, India: Karnataka, Chikmagalur, Mudigere, on undet. plant, in ZSJ). Diagnosis: Dorsal shield 358 long, 224 wide, with 17 pairs of setae. Measurements of setae: i 1-33, i4 -is' J s-j 5' extremely small, ja-36, Za, z4-6 each, s4-105, Z,,-235, Z4-100, ra, R1-IO each. Ventrianal shield 100 long, 67 wide, with 3 pairs of preanal setae. - Mac~osetae on leg IV: genu-117, tibia-94, basitarsus-71. Male; Gupta (1986) described male. India: Karnataka. 23. Amhlyseius (Amblyseius) maai Tseng Amblyseius (Amblyseius) maai Tseng, J. Agr. Ass. China, 94 : Remarks: Neither the material nor the original literature was

31 GUPTA: ORIENTAL PHYTOSEIlDAB 17 available to the author and, hence, it was not possible to provide diagnosis, type data, etc., for this species. Taiwan. 24. Amblyseius (Amblyseius?) modomacroseta Tseng Amb/yseius (Amblyseius) monomacroseta Tseng, J. Agr. Ass. China, 94: Remarks: Same as for species No. 23. Taiwan. 25. Amblyseios (Amblyseius) obtuserellus Wainstein & Beglyarov ~971. Amblyseius obtuserellus Wainstein & Beglyarov, Zool. Zh., 50 l (Holotype ~ (1), USSR : Primorsky Territory) Amblyseius obtuserellus, Wei-nan, Acarology, 6(1) : 222. Diagnosis: Measurements of setae: is-40, S4, -60" ZIS-175, Z4r -85, ii-20, JV Macrosetae on leg IV : genu-55, tibia-20, basitarsus-40. Sparmatophoral process T-shaped. South China, USSR, Japan. 26. Amblyseius (Amblyseius) pubes Tseng Amblyseius (Amblyseius) pubes Tseng, J. Agr. Ass. China, 94 : Remarks: Same as for species No. 23. Taiwan. 27. Amblyseius (Amblyseius?) spenceus Tseng Amb/yseius (Amb/yseius) spenceus Tseng, J. Agr. Ass. China,94: 8S 128. Remarks: Same as for species No. 23. Taiwan. 28. Amblyseius (Amlyseios) tamtavensis Blommers Alllblyseius (Amh/yseius) tamtavensis Blommers, Bull. zool. Mus. Univ. Amsterdam, 3(19): (Holotype ~, Paratypes 2 ~ ~, Madagascar: Tamtave, on Citrus hyslrix, in Institute of Taxonomic Zoology, (Zool. Mus), Univ. Amsterdam). l1j82. Amblyseius tamtavelisis, Oomen, In Veenonan & Zonen, Wagen~ngen, p. 89. Diagnosis: Dorsal shield 340 long, 250 wide, with 17 pairs of setae. Measurements of setae: ;1-32, ja-54, ;,-4, ;5-3, je-3, Ja-5, Jo 6, z4a- Q, 3

32 18 REC. ZOOL. SURV. INDIA, Oce. PAPER. No. 95 Z5-3, Zl-3, Z4-115, Z5-250, s4-85, Ss-5, S4:-6, S5-6. IV : genu-120, tibia-75, basitarsus-73. Unknown (1) Madagascar, Indonesia. Macrosetae on leg 29. Amblyseius (Amblyseius) theae Wei-nan & Xing Amblyseius (Amhlyseius) theae Wei-nan & Xing, Entomotaxonomia, 5(3) (Holotype ~, Paratype 10', South China: Jianxi Jiulian Shan Mountain, on Thea sinensis). Diagnosis: Dorsal shield 335 long, 205 wide. Measurements of setae: jl-22, z~-5, is-27, j4:' je' J a, J 5-5 each, 84:-62, Sa-II, , Za' Z4-13 each, Zl-8, ZI5-96, Z4: DistributiDn: South China. 30. Amblyseius (Amblyseius?) vestificus Tseng Amblyseius (Amblyseius) vestificus Tseng, J. Agr. Ass. China, 94 : Remarks: Same as for species No. 23. Subgenus Asperoseius Chant Key to the species of Amblyseius (Asperoseius) 1. Macrosetae on genu and tibia IV equal, all macrosetae subcapitate Macrosetae on genu IV longer than that on tibia IV ; macrosetae knobbed heveae nuci/era 31. Amblyseius (Asperoseius) heveae (Oudemans) Typhlodromus heveae Oudemans, Ent. Ber., 8: 97. (Holotype ~, Indonesia: Sumatra, on Hevea sp., in Rijksmus. Van. Nat. Hist.) Amh/yseius (Asperoseius) heveae, Ehara & Bhandhufalck, J. Fac. Ed. Totto,i Univ. Nat. Sci., 27 (2) : (Thailand: on Solallum/erox)-Redescribed A. (A.) heveae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 67. Diagnosis: Dorsal shield incised lateral to S4' smooth, with 16 pairs of setae. jl' is, Za' Z~, S4' Sa, Z5' Z4 4long, finely serrate. Setae ra and Rl serrate and on interscutal membrane. Measurements of.

33 GUPTA: ORIENTAL PHYTOSBIIDAE 19 setae: i l -38, i4-7, i5-7, i a -14, 15-4, rs-59, R 1.32, i 3-73, za-20, z4,-37, 84-97, ZI-11, Sa-44, Z5-71, zes-8, Z4-60. Ventrianal shield much longer than wide, lateral margins concave with 3 pairs of preanal setae and a pair of pores. Fixed digit of chelicera multidentate, movable digit 3-dentate. Macrosetae on leg IV: genu-52, tibia-50, basitarsus-76, distitarsus-50. Indonesia, Thailand. 32. Amhlyseios (Asperoseios) nucifera (Gupta) Paraphytoseius (Tropicoseius) nucifera Gupta, Bull. zool. ~urv. India, 2. (1): (Holotype ~,India: Kerala, on Cocos nucifera, in ZSI) AmblyseiuJ (Asperoseius) nuclfera, Gupta & Ray, ibid, 4 (1) : A. (A.) nucifera, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 71. Diagnosis: Body elongated oval. Dorsal shield 305 long, 165 wide, notched at the level of S4. Measurements of setae: jl-28, i4,-i6 4 each, ja-68, za-16, z4-32, 54-80, ZI-4, Sg-40, Zr.,... 76, z5-5, Z4,-68, r a -56, R l -28. Ventrianal shield 80 long, 60 wide, lateral margins concave, with 3 pairs of preanal setae. Fixed digit of chelicera multidentate. Spermatheca with bell-shaped cervix. India (Kerala). In addition to the above, the undermentioned species of this,subgenus are also known from the Oriental region but could not be included in the key because of the reasons as mentioned for A. (A.) spp. 33. Amhlyseios (Asperoseius) hevearom (Oudemans) Typhlodromus hevearum Oudemans, Ent. Ber., 8 : (Holotype 0, Sumatra, on 1) Diagnosis: Female: Z4 shorter than Za ; R1 minute; lateral setae serrate. Indonesia. 34. Amblyseius (Asperoseius) hyauliangensis Gupta Amhlyseius (Asperoseius) hyauliangensis Gupta, Oriental Ins., (In press). (Holotype ~, Paratype 1 ~, India : Arunachal Pradesh, Hyauliang. on an undet. plant, in ZSI).

34 20 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 Diagnosis: Dorsal shield 336 long, 150 wide, with 15 pairs of setae and 6 pairs of fairly large pores. Measurements of setae: jl 36, j4:-j~-minute, is-16, J 5 -minute, js-84-88, za-24, z4-40, s.1-133, Zl-13, Sa-80, Sa-30, ZD-I05, z5-minute, Z4-100, rs-56, R}-33, JV Ventrianal shield 100 long, 56 wide, lateral margins straight, with 3 pairs of preanal setae. Macrosetae on leg IV: genu-26, tibia-49" basitarsus-89, all being knobbed. DistributiDn: India: Arunachal Pradesh. Subgenus Euseius Wainstein Key to the species of Euseius 1. Ventrianal shield with 2 pairs of setae irregularis Ventrianal shield with 3 pairs of setae 2 2. Ventrianal shield wider than long notatus Ventrianal shield longer than wide 3 3. All setae minute excepting ZIS and jl 4 Besides Z 6 and j l' some other setae also long (; longer than 8~ and 8~ and the latter two being equal rhododendronis 84- to S 6 of same length ovulis 84- and S~ longer than S~ S s. Duct of spermatheca long, tubular sacchari Spermatheca duct short, cervix funnel-shaped aizawai 6. Dorsal shield smooth 7 Dorsal shield reticulate 9 7. jl longer than j. neococcineae jl shorter than j ~ noticably longer than Z 1 al810niae S~ only slightly longer than Zl delhiensis 9. Z a gently serrate pruni Za smooth z~ and z, almost of same length 11 z~ noticably shorter than Z4- cocci1leae 11. Metapodal plate hook-shaped 12 Metapodal plate not as above Z, much shorter than S4-... nicholsi Z, longer than 84- okumae 13. Spermatheca sacular finlandicus Spermatheca not sacular Macrosetae on leg IV acuminate eucalypti Macrosetae with broadened tip Sa smaller than S, bambusae S 6 lonler than S4- coccosocius

35 GUPTA : ORIBNTAL PHYTOSEUDAE Amblyseius (Euseius) aizawai Ehara Amblyseius (Amblyseius) aizawai Ebara & Bbandhufalck, J. Fac. Ed. Tottori Univ. Nat. Sci., 27(2) : (Holotype ~,Paratypes 5 ~ ~, 10, Thailand: on papaya, in Bioi. Inst. Fac. Ed. Jrotf. Univ.). Diagnosis: Dorsal shield reticulate along lateral margins, with 17 'pairs of setae; Z5 short, gently serrate. Measurements of setae: i l -31, i.,-7, i 5-8, i 6-9, Ja-10, J 5-5, rs-13, R l -12, js-24, za-14, z4:,13, s4-21, Zt-10, S~-13, S4-15, S5-15, Z5-53. Ventrianal shield oval, widest at anal region with 3 pairs of preanal setae and a pair of conspicuous pores. Fixed digit of chelicera with 1 subapical tooth and 5 adjacent teeth; movable digit with a single tooth. Macrosetae on leg ~V : genu-49, tibia-38, basitarsus-65. Chaetotaxy of dorsal shield in male similar as in female. Thailand. 36. Amhlyseius (Euseius) alstoniae Gupta Ambly.ieills alstoniae Gupta, Internat. J. Acarol., 1 (2) : (Holotype ~, Parat}pes 1 ~, 10, India: West Bengal, Howrah Botanical Garden, on Alstonia scholaris ; 2 ~ ~, Hooghly, undet. plant; all in ZSI) A. a/stoniae, Gupta, Indian J. Acar., 1: 29. (Rajasthan: undet. plant; Gujarat : sapota, chille, cotton, Acacia, date plam, Nerium indicum, cucurbits, pomegranate. Tabernaemontana coronaria, palm; Punjab: cotton, maize, pear, sugarcane, Dalbergia sissol?) A. alstoniae, Gupta, Indian J. A car., 2(2) : 61. (Bihar : undet. plant; Orissa : Butea monosperma, mango) Euseius alsloniae, Gupta, Oriental Ins., 12(3) : 327 (Tamil Nadu : Zinia sp.) A, (E) a/stoniae, Gupta, Fauna of India (Acari Mesostigmata) Family Phytoseiidae, p. 74. Diagnosis: Dorsal shield smooth with 17 pairs of setae and 6 pairs of pores. Measurements of setae: jl-27, J 4, j5-13, each j8' ia-26 each, J 5-6, js-31, za-27, z4-36, s4-56, Z1-19, Sa-27, S4-18, S[,-27, Z5-56 (weakly serrate), z5-12, Z&-28. Ventrianal shield vase-shaped, longer than wide, constricted at preanal region, with 3 pairs of preanal setae. Macrosetae on leg IV : genu-49, tibia-36, basitarsus-70. Spermatheca with elongate cervix and rounded vesicle. Dorsal chaetotaxy similar as in female. India (Rajasthan, Gujarat, West Bengal, Bihar, Orissa, Tamil Nadu, Punjab). Remarks: Its importance in biological control was mentioned by Gupta (1977) and Dhooria (1980).

36 22 REC. ZOOL. SURV. INDIA, OCC. PAPER No Amblyseius (Euseius) bambusae Ghai & Menon Amblyseius bambusae Ghai & Menon, Oriental Ins., 1 : (Holotype ~, Paratypes I&, and 5 others (sex not specified), India: Bangalore, bamboo, infested with eriophyids, deposited in NPe) A. bambusae, Gupta, Internat. J. Acarol., 1(2): Euseius bambusae, Gupta, Oriental Ins., 12(3): (Tamil Nadu : Pyrus communis, Butea monosperma, Dalbergia sp., Ficus religiosa, Coffea arabica, pear; Kerala : unde'. plant; Karnataka: \,jndet. plant; Andhra Pradesh: undet. plant) A. (E.) bambusae, Gupta.. Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 76. Diagnosis: Dorsal shield reticulate with 17 pairs of setae, measurements of Eetae; j1-32, j4' j5 4-6 each, js' j2-11 each, J 5-4, js-24 za, z4-20 each, 54-28, , S2-16, S4-20, S5-17, Z5-50, Z5-4, Z4-12 Sternal, metasternal and genital shields normal. Ventrianal shield vase-shaped with 3 pairs of preanal setae and a pair of pores; fixed digit of chelicera with 4 teeth; movable digit with one tooth, spermatheca with elongate cervix. Macrosetae on leg IV : genu-52, tibia-45 ; basitarsus-62, all with pointed teeth. Gupta (1978) figured its spermatophoral process. India (Karnataka, Tanlil Nadu, Kerala, Andhra Pradesh). 38. Amblyseius (Euseius) coccineae Gupta Amblyseius coccineae, Gupta, Internat J. Acarol., 1(2) : 33. (Holotype ~, Paratypes 3 ~ ~, India: West Bengal, Darjeeling Dist., on Coccinea indica, in 2SI ; 2 ~ ~, Kalimpong, citrus, all in ZSI) Amblyseius coccineae, Gupta, Indian J. Acar., 1 : 30. (Gujarat : Mango, castor) Amb/yseius coccineae, Gupta, Indian J. Acar., 2(2) : 61~ (Orissa: jackfruit, cashewnut, bamboo, Terminalia atjuna, Eugenia sp; Meghalaya: arum) Euseius coccineae, Gupta, Oriental Ins., 12 : 329. (Pondicherry: undet. plant; Tamil Nadu: marigold, Tab ernaemon tana coronar;a ; Andhra Pradesh: on cashewnut) A. (E.) co ccineae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 78. Diagnosis: Dorsal shield reticulate, with 17 pairs of setae. Measurements of setae: ;1-25, ;4' ;5' J s ' z each; J G -6, i 8-27, Za, Z4: each, 54-32, ZI-11, Sa, S4' S5-21 each, Z5-58, Z4t-13. Ventrianal shield vase-shaped, with 3 pairs of preanal setae. Chelicera with Z teeth on fixed digit and none on movable digit. Leg IV with mactosetae on genu-47, tibia-37, basitarsus-55 ; all with flattened tip.

37 GUPTA: ORIENTAL PHYTOSEIIDAE 23 Gupta (1977) figured its spermatophoral process. India (West Bengal, Gujarat, Orissa, Meghalaya, Pondicherry, Tamil Nadu, Andhra Pradesh). 39. Amblyseius (Euseius) coccosocius Ghai & Menon, Amblyseius coccosocius Ghai & Menon, Oriental Ins., 1 : (Holotype ~ J Paratypes 8 ~ ~, Bangalore, on mango, infested with scale insects, in NPC) A. coccosocius, Gupta, Sci. & Cult., 36 : 98. (West Bengal: Alstonia scholaris) Amblyseius coccosocius, Gupta, Internat. J. Acarol., 1 (2) : (West Bengal: on Alstonia scholaris, Clerodendron macrosiphon) A. coccosocius, Gupta, Indian J. Acar., 1: (Punjab: guava) Euseius coccosocius, Gupta, Oriental Ins., 12 : 329. (Pondicherry : undet. plant; Tamil Nadu : papaya; Kerala: undet. plant; Andhra Pradesh : mango) A. (E.) coccosocius, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 80. Diagnosis: Dorsal shield reticulate with 17 pairs of setae. Measurements of setae: i l 28, i4r -ifs' J a each, js-21, za-16, z4-16, , Z1-9, Sa-16, S4-25, S~-29, Z~-57, zf)-8" Z4-16. Sternal shield as long as broad. Ventrianal shield vase-shaped, slightly broadest at preanal region" with 3 pairs of preanal setae and a pair of semilunar pores. Macrosetae on leg IV : genu-36, tibia-32, basitarsus-45. Gupta (1977) figured its spermatophoral process. India (Karnataka, West Bengal, Punjab, Pondicherry, Tamil Nadu, Kerala, Andhra Pradesh). 40. Amblyseius (Euseius) delbiensis (Narayanan & Kaur) Typhlodromus (Amblyseius) delhiensis Narayanan & Kaur, Proc. Indian A cad. Sci., SIB: 5-7. (Holotype ~, (1) described from 10 ~ ~, 5 00' New Delhi, on Hibi3cUS esculentus and Gossypium sp., infested with tetranychid mites, in NPC) Amblyseius delhiensis, Gupta, Sci. & Cult., 36 : 98 (West Bengal: Syzygium javanica) A. delhiensis, Gupta, Sandhu, Dhooria & Singh, Sci. & Cult., 37 : 298. (Punjab: maize, feeding on O. indicus) A. delhiensis, Gupta, Dhooria & Sidhu, Sci. & Cult., 37 : 484. (punjab: citrus, feeding on E. orientalis) A. delhiensis, Gupta & Dhooria, Curro Sci., 41 : (Punjab: grape vines) A. delhiensis, Sandhu, Kaushal & Gupta, Sci. & Cult., 39 : (Punjab; maize, feedins on phytophagous mites).

38 24 REC. ZOOL. SURV. INDlA, OCC. PAP!R No A. delhiensis, Gupta, Internat. J. Acaro/., 1(2) : 36. (West Bengal: Hibiscus rosa-sinensis, litchi, feeding on Oligonychus sp. and Tetranychus sp.) Amblyseius de/hiensis, Gupta, Indian J. Acar., 1 : 30. (punjab: grapevines) A. delhiensis, Gupta, Indian J. Acar., 2(2) : 62 (Orissa: guava) A. (E.) delhiensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytose idae, p. 82. Diagnosis: Dorsal shield smooth with 17 pairs of setae. Za = S4' j6' J a' Zl' S4' S5 and Z4: all of same length, Z5 slightly serrate; z4>z9' Z1 = Sa. Ventrianal shield vase-shaped, Ii times as long as. broad with 3 pairs of preanal setae. Leg IV with macrosetae on genu, tibia, basitarsus, that on tibia being smaller. Narayanan & Kaur (1960) described male. India (New Delhi, West Bengal, Punjab, Orissa). Remarks: Its predatory importance was reported by Gupta et ale (1971), Gupta & Dhooria (1972) and Somchoudhury (1979, 1981.). 41. Amblyseins (Enseius) eucalypti Ghai & Menon Amblyseius eucalypti Ghai & Menon, Orienta/Ins., 1 : (Holotype ~, Paratypes 8 ~ ~, India: Bangalore, on Eucalyptus, in NPC) A. eucalypti, Gupta, Internat. J. Acarol., 1(2) : A. (E.) eucalypti, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 84. Diagnosis: S4>jl = js = za = Z4; j4' Dorsal shield faintly reticulate, with 17 pairs of setae, J 5' Z5' Z4 short, ZI = Sa = S4:; Z5 longest. Ventrianal shield longer than broad, with 3 pairs of preanal setae. Macrosetae of l~g IV present on genu, tibia and basitarsus. India (Karnataka). 42. Amblyseius (Euseius) finlandicus (Oudemans) Seiulus finlandicus Oudemans, Ent. Ber., 4: 183. (Holotype ~, Finland, on Salix caprea, in Rijksmus Van Nat. Hijt. Leiden) Amblyseius finlandicus, Athias-Henriot, Bull. Soc. Hist. nat. Air. N., 49 :" Typhlodronlus finlandicus, Evans, Ann. Mag. nat. Hi~t., (12)6: Amblyseius finlandicus, Ghai & Menon, Orienta/Ins., 1 : 70. (Bangalore : hedge plant) A. hibisci, Gupta, Sci. & Cult., 36 : 98. (West Bengal: arum)-misidentification A. finlandicus, Gupta et al, Sci. & Cult., 37 : 298. (Punjab: Pear, feeding on E. orienta/is).

39 GUPTA: ORIENTAL PHYTOSEIIDAE A. hibisei, Gupta et al., Sci. & Cult., 37: 298. (punjab: cotton)-misidentification A. hibisci, Dhooria & Sidhu, ibid., 37 : 4 ~4. (punjab: feeding on E. orientalis) A. fin/adieus, Sandhu, Kaushal & Gupta, Sci. & Cult., 39 : (Punjab: on maize feeding on all stages of phytophagous mites) A. finlandieus, Gupta, Ent. Rec., 86: A.. finlandicus, Gupta, Internal. J. Acarol., 1(2) : 36. (West Bengal: guava, pear) A finlandieus, Gupta, Indian J. Acar., 1 : 32. (Punjab: undet. plant) A. (E.) finlandicus, Gupta, r auna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 86. Diagnosis: Dorsal shield reticulate with 17 pairs of setae. Measurements of setae: j1-28, j4' j5-10 each, js' 1a-12 each, J 5-6, i a -24, z.-z4-16 each s4-28, Z1-12, Sa-S4-20 each, S5-25, Z5-56, z5-10, Z4-16. Ventrianal shield vase-shaped, with 3 pairs of preanal setae; leg IV with macrose~ae on genu-40, tibia-24, basitarsus-48. Fixed digit of chelicera with 2 teeth; movable digit toothless. Gupta (1977) figured its spermatophoral process. India (Karnataka, West Bengal, Punjab); outside India : Indonesia. Remarks: Gupta et ale (1971) and Sandhu et ale (1973) reported the predatory nature of this species. 43. Amhlyseius (Euseius) irregularis (Evans) 19S3. Typhlodromus irregularis Evans, Ann. Mag. nat. Hisl., (12) 6 : (Holotype ~, Paratypes 2 ~ ~, Malaya, on leaves infested with tetranychids in BM (NH). Diagnosis: Dorsal shield with 16 pairs of setae; 8 in lateral rows; 54: absent. Ventrianal shield oval, lateral margins concave with 2 pairs of preanal setae and a pair of pores. Leg IV with 3 macrosetae one each on genu, tibia and basitarsus.? Malaya. 44. Amhlyseius (Euseius) neococcineae Gupta Amblyseius neocoeeineae Gupta, Indian J. Acar., 2 (2) : 69. (Holotype ~, Paratypes 4 ~ ~, Meghalaya, Tura, on Syzygium eumini; all in ZSI, Calcutta) A. (E.) neococcineae Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p

40 26 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 9-5 Diagnosis: Dorsal shield 265 long, 145 wide, smooth with 17 pairsof setae. Measurements of setae: jl-26, j4' ja-8 each, je, 1 a-12 each, J 5-5, js, za-12 each, z4.. 16, s4-24, 21-12, Sa-16, S4-16, Sa-18, Za-45, z5-12, 2d, -16. Sternal and genital shields normal; ventrianal shield with lateral margins concave at preanal region with 3 pairs of preanal setae and a pair of crescent-shaped pores. Macrosetae on leg IV : genu-20, tibia-24, basitarsus-32 ; all with flattened tip. India (Meghalaya). 45. Amblyseius (Euseius) nicholsi Ehara & Lee Amblyseius (Amblyseius) nicholsi Ehara & Lee, J. Fac. Ed. Tottor; Univ. Nat. Sci., 22 : 67. (Holotype ~,Hong Kong: Chai Wan, on grass, in BioI. Inst. Fac. Ed. Tolt. Univ.) Amblyseius (Amblyse;us) nicholsi Ehara & Bhandhufalck, J. Fac. Ed. Tottor; Univ. Nat. Sc;., 27(2) : 61. (Thailand: on Combretum quadrangulare, Cassava, Leauminous tree). Diagnosis: Dorsal shield reticulate, with 17 pairs of setae and 6 pairs of pores. Za faintly barbed; jl also stout. Measurements of setae: j1-35, j4-11, ja-ll, 1 a -14, J 5-7, rs-15, R l -14, js-28, za-18, S4-30, Z1-14, Sa-18, S4-26, SI)-26, 2a-62. Ventrianal shield nearly oval, widest at anal region, with 3 pairs of preanal setae. Metapodal plates hook-shaped. Macrosetae on leg IV : genu-52, tibia-41, basitarsus-66, all with blunt tip. Hong Kong, Thailand. 46. Amblyseius (Euseius) notatus Chaudhri Amblyseius notatus Chaudhri, Acarologia, 10 : 555. (Holotype ~, Pakistan: 4 miles south of Murree Hills, on Fragaria vesca, in UAL). Diagnosis: Dorsal shield 347 long, 219 wide, reticulate, simple, with 17 pairs of setae. jl-34, j4-16, ja-18. js-21, J a -23, js-39, Za-z4:-34 each, 54-47, ZI' Sa-S4-26 each, Sa-31, Z~-60 (weakly serrate), z5-18, , ra-18, R1-17. Ventrianal shield oval or round, with 3 pairs of preanal setae. Metapodal plates absent. Spermatheca with narrow tube, atrium bell-shaped. Macrosetae on leg IV : genu-39, tibia-36 and basitarsus-54. Dis tribution: Pakistan.

41 GUPTA: ORIENTAL PHYTOSEllDAE Amblyseius (Euseius) okumae Ehara &. Bhandhufalck Amblyseius (Amblyseius) okumae Ehara & Bhandhufalck, J. Fac. Ed. Tottori Univ. Nat. Sci., 27(2): (Holotype ~, ~aratype) 9 ~ ~, 1 d", Thailand: Doi CQiang Oao, on Tea; 1 ~, 10' same locality, on Michelia champaca, in BioI. Inst. Fac. Ed. Tolt. Univ.). Diagnosis: Dorsal shield indented near R1.' reticulate with 6 pairs of pores and 17 pairs of setae, Z5 stout with minute barbs. Peritreme ends between js and Za. Measurements of setae: j1.-27, j4-14, j5-15, J s -18, J&-8, rs-16, R I -14, js-25, zs-21, z4-23, s4-31, ZI-18, Sa-22, S.,-22, S5-27, Z5-53, z5-15, Z4-21. Ventrianal shield oval, widest at anal region, with 3 pairs of preanal setae. Fixed digit of chelicera with a few teeth distally, movable digit with 1 tooth. Macrosetae on leg IV ; genu-33, tibia-28, basitarsus-47, all with blunt tip. Measurements of setae smaller than those of females. Spermatophoral process was figured by Ehara & Bhandhufalck (1977). Thailand. 48,. Amblyseius (Euseius) ovalis (Evans) Typhlodromus ovalis Evans, Ann. Mag. nat. His!., 6: (Holotype ~, Malaya: Kuala Lumpur, on rubber, in BM (NH) Typhlodromus (Amblyseius) ovalis, Narayanan, Kaur & Ghai, Proc. natn. Inst. Sci. India, 26(B)(6) : (Bombay: Ficus sp., Terminalia arjuna, both infested with tetranychid mites, banana, Karnataka : undet. plant) A. o),'alis, Corpuz & Rimando, Philipp. Agrie., 50 : (Philippines: Averrhoa carambola, Breynia, rhamnoides, Gliricidia sepium, Pandanus sp., Allamanda hendersonii, Swietenia macrophy1ia) A. ovalis, Gupta, Sci. & Cult., 36 : 98. (West Bengal: Bougainvillea spectabilis) A. (A) ovalis, Ebara, Mushi, 43(6) : (Taiwan: Spathodea campanulata) A. ovalis, Gupta, Ent. Rer., 86: A. ovalis, Gupta, Internat. J. Acaroi., 1(2) : (West Bengal: Vitex pubescens, Ficus sp.) A. o valis, Lo, Bull. Sun- Yat. Sen. Cult. Found., 5 : (Taiwan: grape, Brassica with Tetranychus sp.; pear with Oligonychus sp. ; Bryophyl/um calycimum with Brevipalpus ~p. ; Lycium chinense with eriopbyid mites) A. ovalis, Gupta, Orienta/Ins., 11 : 632. (Andaman Nicobar IsIs. : Tabernaemontana, Ficus, mango, banana).,1977. A. ovalis, Gupta, Indian J. Acar., 1 : 34 (Gujarat : beans)..

42 28 REC. ZOOL. SURV. INDIA, Oce. PAPER No Euseius ovalis, Gupta, Oriental Ins., 12: (Tamil Nadu: Napier grass, lucern, banana, castor, Nerium sp., mango, coconut, cashewnut, oilseed plant, Bauhinia sp., papaya, pomegranate, rose, chrysanthemum, coffee, Pondicherry: Bauhinia purpuria, Kerala: paddy, ornamental plant, Karnataka: undet. plant, sugarcane; Andhra Pradesh 2 cashewnut) A. ovalis, Gupta, Indian J. Acar., 2(2) : 71. (Meghalaya: undet. plant; Tripura : Bougainvillea, banana; Manipur : grass) A. (E.) ovalis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 92. Diagnosis: Dorsal shield 345 long, 244 wide, with 17 pairs of setae. Measurements of setae: jl-31, j4-i6' J S -J each, is-12, Za, Z4:-7 each, s4-12, ZI' Sa-S each, Z5-55, z5-5, Z4-8. Ventrianal shield oval, 102 long, 81 wide, lateral margins convex to make widest at anal level, with 3 pairs of preanal setae and a pair of crescent-shaped pores. Leg IV with macroseta on genu-20, tibia-24, basitarsus-32, all smooth with flattened tip. Spermatheca with bell-shaped cervix. Gupta (1977) figured its spermatophoral process. Malaya, Philippines, Taiwan, Indonesia; India (Maharashtra, Karnataka, West Bengal, Andaman & Nicobar IsIs., Tamil Nadu, Pondicherry, Kerala, Meghalaya, Tripura, Manipur t Gujarat). Remarks: Predatory behaviour of this species was reported by Rao et ale (1970) and Gupta (1977). 49. Amhlyseius (Euseius) pruni Gupta Amblyseius sojaensis Ebara, Sci. & Cult., 36: 98 (West Bengal: Anthocephalus indicus)-misidentification A. sojaensis, Gupta, Internal. J. Acarol., 1(2) : (West Bengal: Bignonia mangifera, Mangifera indica) - Misidentification Amblyseius pruni Gupta, Internal. J. Acarol., 1(2) : (Holotype ~. Paratypes 2 ~ ~, 1& India: West Bengal, Kalimpons. Prunus persica, in ZSI, ) A. pruni, Gupta, Indian J. Acar., 2(2) : 73. (Meghalaya : pear; Assam: undet. plant; Tripura : Bauhinia acuminata) A. (E.) pruni, Gupta, i auna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 94. Diagnosis: Dorsal shield 315 long, 230 wide, reticulate, with 17 pairs of setae and 4 pairs of pores. Measuremets of setae: jl-3!, j4-j5 8 each, j6-ja each, J 5-5, js-27, za-z4: 19 each, s4-33, Zl-13, Sa-18, S4-18, S5-22, Z5-58, z5-12, Z4-13. Sternal, metasternal and' genital shields normal. Ventrianal shield vase-shaped, slightly longer'

43 GUPTA: ORIENTAL PHYTOSEllDAE 29 than wide, with 3 pairs of preanal setae and a pair of crescent-shaped pores. Macrosetae on leg IV : genu-40, tibia-36, basitarsus-58. India ( West Bengal, Meghalaya l Assam, Tripura). 50. Amblyseius (Euseius) rhododendronis Gupta Amblyseius (Euseius) rhododendronis Gupta, Oriental Ins., 4 : (Holotype ~, Paratypes 2 ~ ~, India: West Bengal, Darjeeling, on Rhododendron sp. ; another ~, Lataguri, on undet. plant; 1 ~, Sevak, on Shorea robusta, all in ZSI) A. rhododendronis, Gupta, Internal. J. Acaro/., 1(2) : Euseius kodaikanalensis Gupta, Oriental Ins., 12: (Tamil Nadu : Pear) A. (E.) rhododendronis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p. 96. Diagnosis: Dorsal shield 315 long, 220 wide with 17 pairs of setae and 5 pairs of pores. Measurements of setae: i l -27, j4 -ie' J a 8-9 each, J~-4, ja-16, Za-Z each, S4-20, Z1-8, Sa-II, S4,-17, S5-9, Z~rot54, z~-5, Z4-9. Sternal, matastemal and genital shields normal. Ventrianal shield 94 long l 72 wide, with 3 pairs of preanal setae and a pair of preanal pores. Spermatheca filamentous. Macrosetae on leg IV : genu-31, tibia-22, basitarsus-42, all with broadened tip. India (West Bengal). Remarks: A. (E.) kodaikanalensis Gupta is a synonym for A. (E.) rhododendronis Gupta. 51. Amblyseius (Euseius) sacchari Ghai & Menon Amblyseius sacchari Ghai & Menon, Oriental Ins., 1 : (Holotype i!, Paratypes ~ and 7 others (sex not specified), India: Bangalore, sugarcane, infested with eriophyids, in NPC.) A. sacchari, Sci & Cult., (West Bengal: sugarcane) A. sacchari, Gupta & Dhooria, Abstr. Indian 'Sci. Congr., 1974: 69. (Punjab & Himachal Pradesh) A. sacchari, Gupta, Internal J. Acaro/., 1(2) : 43. (West Bengal: sugarcane) A. sacchari, Gupta, Indian J. Acar., 1 : (Redescribed) (Gujarat : sugarcane) Euseius sacchari, Gupta, Oriental Ins., 12: 337. (Tamil Nadu : Sugarcane) A. (E.) sacchari, Gupta, Fauna ot" India (Acari: Mesostign1ata) Family Phytoseiidae, p. 98.

44 30 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 95 Diagnosis: Dorsal shield 337 long, 211 wide, with 17 pairs of setae and a few pairs of pores. Measurements of setae: jl-27, j4 -j6 J a-j each, ja-12, Za' Z4' ZI' Sa-S eaeh, S4-17, (serrate), Z5-5, Ventrianal shield elongate, 114 long, 72 wide, with 3 pairs of preanal setae and a pair of preanal pores. Macrosetae on leg IV : genu-36, tibia-40, basitarsus-60. Gupta (1977) fi~ured spermatophoral process. India (Karnataka, West Bengal, Punjab, Himachal Pradesh, Gujarat, Tamil Nadu). Besides the above species, the following are also known from the Oriental region but could not be included in key for the reasons as mentioned for A. (A.) spp. 52. Amblyseius (Euseius) alangii Lai-rong & Li-sheng AlnblYseius alangii Lai-rong & Li-sheng, Fudan J. (Nat. Sci.), 20(2) : 222. (Holotype ~, Paratype 1 ~, South China: Jiangsu, Yixing, on Alangium chinense). Diagnosis: Dorsal shield 330 long, 206 wide. Measurements of setae: jl-33, j4-13, j5-13, j6-17, la-is, 15-4, ja-26, za-17, z4,-15, , , S2-22, S4-19, S5-28, Z5-59, fa-17, R 1 -l3, z5-13, Z4-19, JV 5-44 South China. 53. Amblyseius (Euseius) densus Wei-nan Amblyseius (Amblyseius) densus Wei-nan, Acta. Zootaxonomica Sinica, 9(2): (Holotype ~, Paratypes 10 ~ ~, South China: Longchuan & Lanceng Counties, on Palonia sp. and Prunus persica). Diagnosis: Dorsal shield long, wide. Measurements of setae: j~-27, j4-17, j5-20, j6-22, J a -20, 1 5-5, js-25-28, z4-28, za-27, 54-30, , S2-22, S4-21, S5-21, Z5-45, z5-15, Z4-21, ra-20, R l -13, JV South China. 54. Amblyseius (Euseius) guangxiensis Wei-nan Amblyseius (A111h/yseius) guql1gxiensis Wei-nan, Acarology, 6(1) : 223. (Holotype ~, Paratypes 3 ~ ~, South China : Gulien, Gnangxi).

45 GUPTA: ORIBNTAL PHYTOSEIIDAE 31 Diagnosis: Dorsal shield long, wide. Measurements of setae: i l -32, i 4-10, j5-12, i s -13, J 2-17, J 5-15, i s -32, za-18, z4,-18, s4-35, Zl-15, Sa-22, S4-30, S5-31, Z5-67, z5-10, , ra-17, R l -15, JV 5-37 South China. 55. Amblyseius (Eoseius) macrospatulatos Gupta Amblyseius (Euseius) macrospatulatus Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p (Holotype ~, India: Arunachal Pradesh, Likabali, on an undet. plant, in ZSI). Diagnosis: Dorsal shield 414 long, 291 wide, with 17 pairs of setae. Measurements of setae: i I -51, i4-7, h~-9, js-11, J a -15, J 5-5, ja-12, za-12, z4-17, s4-29, Zl-15, SJ-17, S4-17, S5-17, Z5-68, z5-9, , r s ' R l -12 each. Ventrianal shield 123 long, 100 wide with 3 pairs of preanal setae. Macrosetae on leg IV : genu-47, tibia-45, basitarsus-69, all with spatulate tip. India: Arunachal Pradesh. 56. Amblyseios (Euseius) mediocris (Chaudhri, Akbar & Rasool) Euseius mediocris Chaudhri, Akbar & Rasool, Rept. PL 480 Project, Univ. Agril. Faisalabad, Pakistan, p. 60. (Holotype ~, Pakistan: Karachi, on Ricinu ~ communis, in UAF). Diagnosis: Dorsal shield 357 long, 224 wide. Measurements of setae: jl-31, j4-13, j5-13, js-23, J 5-5, ja-34, za-29, z4-29, , Zl-23, Sa-18, S4-18, S5-27, Z5-60, z5-13, , rs-18, R l -14. Macrosetae on leg IV : genu-55, tibia-29, basitarsus-70. Pakistan. 57. Amblyseius (Euseios) ruiliensis Wei-nan & Zhao-Quan Amh/yseius (Amh/yseius) ruiliensis Wei-nan & Zhao-Quan Acta Zootaxollolnica Sinica, 10(3) : (Holotype C, Paratypes 6 ~ ~, South China, Ruili. County, Yunnan Province, on undet. plant). Diagnosis: Dorsal shield 250 long, 187 wide. Measurements of setae : j , j , j rs -1 7, j , J 2-1 7, J 5-5, j , z ,

46 32 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 95 Z , , ZI-17-20, S , S , S5-25, Z , z5-20, Z , rs-23-26, R I , JV ~ South China. 58. Amblyseius (Euseius) septic us (Chaudhri, Akbar & Rasool) Euseius septicus Chaudhri, Akbar & Rasool, Rept. PL 480 Project, Univ. Agril. Faisalabad, Pakistan, p. 58. (Holotype ~,Pakistan: Galwala forest, Faisalabad, on Ricinus communis, in UAF). Diagnosis: Dorsal shield 306 long, 253 wide. Measurements of setae: jl-31, j4-j5-16 each, j6' J a -29 each, J 5-5, js-29, za-29: z4-42, 54-57, ZI-34, S2-26, S4-34, S5-23, Z5-57, z5-16, Z4-29, rs-20, R 1 -lj. Macrosetae on leg IV : genu-47, tibia-36, basitarsus-70. Pakistan. 59. Amblyseius (Euseius) womersleyi Amblyseius womersleyi, Shih, A carologia, 6(1) : 710. Remarks: Same as for species No. 23. Taiwan, 60. Amblyseius (Euseius) Yunnanensis Wei-nan Amblyseius (Amblyseius) yunnanensis Wei-nan, Acta Zootaxonomica Sinica, 9(2) : (Holotype ~, Paratypes 2 ~ ~, South China: Yiangjiang County, on Mangifera indica). Diagnosis: Dorsal shield 350 long, 225 wide. Measurements of setae: ;1-25, ;4r -;6' J 5-6 each, J a-7, js-22, za-12, Z4-6, s4-37, Zl-8, Sa-10, S4-6, S~-6, Z5-65, z~-7, Z4-50, rs-15, R South China. 61. Amblyseius (Euseius) relictus (Chaudhri, Akbar & Rasool) Euseius relictus Chaudhri, Akbar & Rasool, Rept. PL 480 Project, Univ. Agril. Faisalabad, Pakistan, p. 56. (Holotype ~, Pakistan: Madyan, on Canabis sativa, in UAF).

47 GUPTA: ORIENTAL PHYTOSElIDAE 33 Diagnosis: Dorsal shield 316 long, 224 wide. Measurements of set'ae: j1-31, j4-10, i 5-13, j6-16, J 2-16, J 5-8, js-29, z2-23, z4-26, 54-30, Macro Z1-21, S2-21, S4-26, 'S5-13, Z6-55, z5-13, Z4-16, ra-16, R l -16. setae on leg IV : genu-43, tibia-34, basitarsus-60. Pakistan. Subgenus Neoseiulus Hughes Key to the species of Neoseiulus 1. Dorsal shield narrow and elongate Dorsal shield round and oval 2. Z1 and S2 very small (8-10 p,) Zl and S2 longer than 8-10p, 3. Sternal shield without p03tlateral angulation Sternal shield with postlateral angulation 4. js=s4 js<s" S. All setae on dorsal shield very small except Zzs Setae on dorsal shield of diverse length 6. Dorsal shield reticulate Dorsal shield smooth 7. Macrosetae on basitarsus IV longer than that on genu IV Macrosetae on genu IV and basitarsus IV equal Setae on dorsal shield of same length or uniformly long, none greatly longer than others Some setae on dorsal shield appreciably longer, others being small or very small 9. All setae on dorsal shield except j l' J!S and S 15 being very long Not as above 10. j 1 longer than j s j 1 shorter than j s 11. No macroseta on tibia IV Macrosetae present on genu, tibia and basitarsu} of leg IV 12. z" shorter than Z2 z" longer than Z2 13. Setae on dorsal shield very long almost touch bases of next setae Setae on dorsal shield shorter 14. Setae on dorsal hexagonal area shorter than the interval between their bases Setae on dorsal hexagonal area longer. than the interval between their bases ~ 2 S aceriae 3 baraki 4 taiwanicus paspalivorlls assamensis siamensis anuwati 13 9 longispinosus 10 cynodonae 11 rangatensis 12 aequipi/us frafercli/us inlbricatlls fallacis

48 34 REC. ZOOL. SURV. INDIA, Oce. PAPER No Zl) and Z" serrate Zrs and Z" smooth 16. Ventrianal shield with sharp inward curve at near anterior lateral side No such character on ventrianal shield 17. Fixed digit of chelicera with 5 teeth... - Fixed digit of chelicera with 4 teeth indicus venustus 17 stolidus colora; 62. Amblyseius (Neoseiulus) aceriae Gupta Amb/yseius aceriae Gupta, Internat. J. Acarol., 1 (2) : (Holotype ~ J India: West Bengal, MaIda, on Mangifera indica, associated with mango bud-mite, in ZSI ) Amh/yseius aceriae, Gupta, Indian J. A car. J 1(1-2): (Gujarat: Chrysanthemum sp.) Amblyseius (Neoseitl/us) aceriae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield reticulate, with 17 pairs of setae, all being small (8-10) except Z5 which is 49 long, thick and serrate. Sternal shield longer than wide, metasternal plate \vith seta. Ventrianal shield quadrangular, longer than broad, with 3 pairs of preanal setae. Spermatheca bell-shaped. Macroseta present only on basitarsus IV. Gupta (1977) figured its spermatophoral process. India (West Bengal, Gujarat). 63. Amhlyseius (Neoseiulus) aequipilos Berlese Amb/yseius aequipi/us Berlese, Redia 10 : 145. (Ho!otype ~, Java, in Berlese's Collections, Florence). Diagnosis: Dorsal shield with 17 pairs of setae, setae S4' Z5' Z4 moderately long; Zl and S!) minute, js =za = 1/2 S4. Ventrianal shield oblong, longer than wide with 3 pairs of preanal setae. Leg IV with macrosetae on genu, tibia and basitarsus.? Indonesia, British Columbia. 64. Amhlyseius (Neoseiulus) anuwati Ehara & Bhandhufalck Amblyseius (All1blyseius) allilwati Ehara & Bhandhufalck, J. Fac. Ed. Tottori. Univ. Nat. Sci., 27(2) : (Holotype ~,Paratypes 2 ~ ~, ThailQ,nd: Prew, on coffee, in Biol. Inst. F410. ~d. Tott. Univ.).

49 GUPTA. : ORIENTAL PHYTOSEIIDA 35 Diagnosis: Dorsal shield reticulate, with 11 pairs of setae and 7 pairs of pores; Z5 stout with minute barbs. Peritreme extends beyon~ jl. Measurements of setae: j1-12, j4-7, j5-7, js-8, J 2-11, J 5-8, rs-lo, R l -9, i 3, z2-z4-11 each, s4-13, , S2-11, S4-10, S5-8, , z&-8, Z4-12. Ventrianal shield near pentagonal, slightly longer than wide, with 3 pairs of preanal setae and a pair of conspicuous crescentic pores. Fixed digit of chelicera multidentate. Leg IV with macrosetae on genu-29, tibia-18, basitarsus Thailand. 65. Amblyseius (Neoseiulus) assamensis (Chant) Typhlodromus ( Amblyseius ) assamensis Chant, Call. Ent., 92 : 60. (Holotype ~,India: Assam, Jorhat, on citrus, in Citrus Expt. Stn., Riverside California. ) Amblyseius assamensis, Gupta, Internat. J. Acaroi., 1 (2): Amblyseius (Neoseiulus) assamensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield smooth with 17 pairs of setae; all being small except s4. (12) and Z& (40, thick). Ventrianal shield pentagonal. Genital shield normal. Fixed digit of chelicera multidentate; movable digit with no tooth. Macrosetae on leg IV : genu-12, tibia.. 20, basitarsus-32. Male; India (Assam). 66. Amblyseius (Neoseiulus) baraki Athias.. Henriot Amblyseius baraki Athias-Henriot, Bull. Scientijique Bourgogne, 25: (Holotype ~,DZ Baraki, on Phalaris sp., respository-unknown.) Amblyseius (Amblyseius) baraki, Ebara & Bhandhufalck. J. Fac. Ed. Tottori Univ. Nat. Sci., 27(2) : (Thailand: on grass; ~.. re-described, a-described for the first time) Amblyseius dhooriai Gupta, Indian J. Acar., 1 : (Holotype ~, India : Punjab, sugarcane, in ZSI) Amb/yseius (Neoseiulus) baraki, Gupta, Fauua of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Body slender, with 17 pairs of setae and 8 pairs of por~s ; setae Z6 barbed, long; other setae small. Measurements of

50 36 REC. ZOOL. SURV. INDIA, Oee. PAPER No. 95 setae: jl-13, j4-11, js-13, 12-12, 15-11, rs-14, R1-16, ja-16, za-13.. z4-15, s4-16, ZI-14, S2-16, S4-23, S5-21, Z5-65, z5-11, Z4-20. Ventrianal shield slightly longer than wide, reticulate with 3 pairs of preanal setae and a pair of crescentic pores. Spermatheca with bowl-shaped cervix. 'Fixed digit of chelicera multidentate, rnovable digit bidentate. Basitarsus IV with macroseta-34 long. Chaetotaxy same as in female. Palaearctic region, Thailand, India (Punjab). 67. Amhlyseius (Neoseiulus) cynodonae Gupta Amhlyseius cynodonae Gupta, Oriental Ins., 11: (Holotype ~,Paratypes 3 ~ ~,Andaman Isis., Port Blair, South Point, on Cynodon dactyloll, in ZSI) Amblyseius (Neoseiulus) cynodonae Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield smooth, Z4' Zo and S4 being long; other setae smooth, short, pointed. Measurements of setae: jl-17, j4-j5' J 5-5 each, js' J a.. 10 each; js-12, Za, z4-9 each, 84-25, Z1.. 10, S2-16, S4-12, SI)-10, ZI)-60, zo-5, Z4-36 Ventrianal plate shield.. shaped, as long as wide. Macrosetae on leg IV : genu-40, tibia-is, basitarsus-55. India (Andaman and Nicobar IsIs). 68. Amblyseius (Neoseiulus) calorai Corpuz & Rimando Amblyseius calorai Corpuz & Rimando, Philipp. Agrie., 50 : (Holotype ~, Philippines: Tuguegarao, Cagayan, on Oryza sativa, in Dept. Ento, Univ. Philip. Paratype 1 ~, same data as for holotype; 10', Caba, Albay, on paddy; 4 ~ ~, 10', Laguna, on Digitaria sanquinalis; 10' on Elusine indica; 10', Laguna, on Blumea balsanlijera, all in Dept. Ento. Univ. Philip.). Diagnosis: Dorsal shield with conspicuous groups of wart-like areas, 4 pairs of pores, 370 long, 204 wide with 17 pairs of setae. All setae minute except Z4 and Z5 measuring respectively 32 and 54 long; both thick, latter serrate. Ventrianal shield pentagonal, slightly longer than wide, reticulate with 3 pairs of preanal setae. Peritreme extends anteriorly upto j1. Fixed digit of chelicera with 2 teeth and pilus dentilis, movable digit unidentate. Macroseta on basitarsus IV-52 long. Spermatheca with cylindrical cervix, slightly narrowed and constricted at the junction of atrium; atrium with anterior portion produced into 2 lateral arms at the bases of the ducts giving a U"shaped appearance.

51 GUPTA: ORIENTAL PHYTOSEllDAE 37 Chaetotaxy of dorsum similar to female. Spermatophoral process T -shaped. DistributiQn: Philippines. 69. Amblyseius (Neoseiulus) fallacis (Garman) Iphidulus /allacis Garman, Bull. Conn. agric. Expt. Stn., 520: 13. (Holotype ~, USA: Connecticut, on apple, in Connecticut Agr. Expt. Stn., New Haven) Amblyseius/allacis, Athias-Henriot, Bull. Soc. Hist. nat. A/r. N., 49 : Typhlodromus (Amblyseius)jallacis, Narayanan, Kaur & Ghai, Proc. /latn. lost. Sci. India, 26 (B) (6) : (India: Haryana: on sowank.) Amblyseius jallacis, Gupta, Sci. & Cult., 36 : 98. (West Bengal: paddy) Amblyseius/allacis, Gupta et al., Sci. & Cull. t 37 : 298. (Punjab: paddy, feeding on O. indicus) Amblyseius fal/acis, Gupta, Ent. Rec., 86: Amblyseius jallacis, Gupta, Internat. J. Acarol., 1(2): 36. (West Bengal: paddy, feeding on O. indicus) Amblyseius ja/lacis, Gupta, Oriental Ins., 11 : 629 (Andamao lsi. : paddy) A. jallacis, Gupta, Indian J. Acar., 1 : (Punjab: carrot, paddy) A. fallacis, Gupta, Indian J. Acar., 2(2): 62. (Meghalaya : paddy; Assam : paddy, arum; Tripura : Mlisa sp.) Neoseiulus fallacis, Gupta, Orienta/Ins., 12 : 331 (Tamil Nadu : grass & paddy) Amblyseiuy (Neoseiulus) fallacis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidao, p Diagnosis: Dorsal shield smooth with 17 pairs of setae measuring : jl-23, j4-20, j5-23, js-37, J 2-43, J~-11, js-36, z2-31, z4-32, S4-40, ZI-40, Ss 41, S4-48, S5-48, Z5-68, z5-28, Z~-S9. Sternal and genital shields normal. Ventrianal shield triangular, as long as broad or slightly longer, with 3 pairs of preanal setae. Macrosetae on leg IV: genu-17, tibia-32, basitarsus-so. The setae on the dorsal hexagonal area longer than the distance between their bases.? India (Haryana, West Bengal, Punjab, Andaman IsIs.,,Meghalaya, Assam, Tripura, Tamil Nadu). 70. Amhlyseius (Neoseiulus) fraterculus Berlese Amb/yseius jraterculus Berlese, Redia, 12: 172. (Holotype ~, Argentina: La Plata, in Berlese's Collections, Florence) Amblyseills fraterell/us, Ghai & Menon, Oriental IllS., I : 72. lcoimbatore-red gram).

52 38 REC. zoot. SURv'. INDIA, Oce. PAPER No Amblyseius /raterculus, Gupta, Internal. J. Acarol., 1(2) : Amblyseius (Neoseiulus) Ira/ercu/us, Gupta, Fauna of India (Acari: Mesostig. mata) Family Phytoseiidae, p Diagnosis: Dorsal shield with 17 pairs of setae; s 4.' Z 5 and.z 4 long; Za, Z4' Z1. and Sa equal to one another; ventrianal shield broad, triangular, with lateral margins concave, with 3 pairs of preanal setae. Four pairs of setae present around ventrianal shield. Leg IV with mocrosetae on genu, tibia and basitarsus.? India (Tamil Nadu), Argentina. 71. Amhlyseius (Neoseiulus) imbricatus Corpuz & Rimando Alnblyseius il1zbricatus Corpuz & Rimando, Philipp. Agric., 50: 127. (Holotype ~ J Paratypes 5 ~ ~, Philippines: Laguna, Los Banos, on Oryza sativa, in Dept. Agr. Univ., Philip.. Additional paratypes: 1 ~, Laguna)l on Asparagusplumosus~' 1 ~, Tuguegarao, Cagayan, on paddy, in Dept. Ento. Univ. Philip.) Amblyseius (Amb/yseius) imbricatus, Ebara & Bhandhufalck, J. Fac. Ed., Tottor; Univ., Nat. Sci., 27(2) : (Thailand: on paddy: Re-described) Amblyseius (Neoseiulus) inbricatlls, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield reticulate, with 17 pairs of setae and 5 pairs of pores. Lateral setae longer than or as long as distance to the bases of setae next behind, setae on dorsal hexagonal area shorter than distance between their bases. Measurements of setae: j1-33, j..-30, i 5-35, j6-48, J a -57, J 5-14, rs-49, R1-51, ja-48, za-45, z4-50, s4-58, Z1-55, Sa-61, 84-61, S5-58, Z5-77, zs-35, Z4-67. Ventrianal shield pentagonal, imbricate with 3 pairs of preanal setae and a pair of crescentic pores. Spermatheca with bowl-shaped cervix. Macrosetae on Leg IV: genu-36, tibia-41, basitarsus-76. Fixed digit of chelicera multidentate, movable digit with 2 teeth. Ehara & Bhandhufalck (1977) described male. Philippines, Thailand. 72. Amblyseius (Neoseiulus) indicus (Narayanan & Kaur) Typlzlodromus (Amb/yseius) illdicus, Narayanan & Kaur, Proc. Indian A cad. Sci., 51 B : 2-5. (described from 12 ~ ~, 400, collected on Gllorilnoscizenla infested potato, feeding on T),ropilaglls jjlllrescclltiae ; sex of holotype and paratypes not mentioned, in NPC).

53 GUPTA: ORIENTAL PHYTOSEllDAE Amblyseius indicus, Gupta, Sci. & Cult., 36 : 98 (West Bengal: wheat) Amblyseills indicus, Sandhu, Kaushal & Gupta, Sci. & Cult., 39 : (Punjab: maize, feeding on all stages of phytophagous mites) Amblyseius in dicus, Gupta, Internal. J. Acaro/., 1(2) : 37. (West Bengal: Zea mays, found feeding on eggs of O. indicus) Amblyseius indicus, Gupta, Indian J. Acar., 1 : 32 (Punjab: wheat) Amblyseius bindrai Gupta, Indian J. Acar., 1 : Amblyseius (Neoseiulus) indicus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield with 17 pairs of setae. Measurements of setae: j1-12, j4' j6' J 2, J 5-9 each, ja, Z2' Z each, s4:-16, Z1-10, Sa' S5-18 each, Z5-56, z5-9, Z4-23. Sternal shield almost as long as broad. Ventrianal shield pentagonal, with 3 pairs of preanal setae. Spermatheca with long, narrow cervix. Leg IV with macroseta only on basitarsus-45 long. India (New Delhi, West Bengal, Punjab). 73. Amhlyseius (Neoseiulus) longispinosus (Evans) Typhlodromus /ongispinosus Evans, Ann. Mag. nat. Hist., (12) 5: (Holotype ~, Indonesia: on Manihot until1isima, in BM (NH) Typhlodromus longispinosus, Evans, Ann. Mag. nat. Hist., (12) 6 : Amblyseius longispinoslls, Corpuz & Rimando, Philipp. Agric., 50 : (Philippines: Ipomoea sp., rose, Carica papaya, Psophocarpus tetragonolobus, Boehmeria nivea, Elusine indica. Asystacia gangetica, Glycine max, Dolichos lab lab, Verbena sp., Wedelia biflora, Canavalia nzaritima, Passiflora foetida. Phaseolus lunatus. Saccharum officinarum, rose, papaya. C. maritima) Amblyseius longispinosus, Gupta, Sci. & Cult., 36: 98 (West Bengal: Datura metel) Amblyse;us (Neoseiulus) longispinosus, Ebara, Mush;, 43 (6): 56. (Taiwan: Ampelopsis heterophylla, Momordica conchinchinensis, Paederia chinensis) Amblyseius longispinosus, Gupta, Internal. J. Acarol., 1 (2): 38 (West Bengal: Datura sp., Tabernaemontana coronaria, feeding on Tetranychus sp.) A~blyseius longispinosus, Lo, Bull. Sun.Yat Sen. Cult. Found.,S: (Taipei: Ipomoea batatas with T. cinnabarinus, Arachis hypogaea with Tetranychus sp., mulberry with Tetranychus sp. and Eotetranychus sp., Salix sp. with Tetranychus sp., pear with Tetranychus sp., sugarcane with O/igollychus sp., citrus with Panonychus citr;. Chrysanthemum sp.) Amb/yseius longispinosus, Gupta, Oriental Ins., 11 ~ 631 (A.ndaman lsi. on paddy).

54 40 REC. ZOOL. SURV. INDIA, OCC. PAPER No Amblyseius (Amblyseius) longispinosus, Ebara & Bhandhufalck, J. Fac. Ed. Tottori Univ. Nat. Sci., 27(2) : 54. (Thailand: beans, long bean, mulberry, grass, Cassava durian, Ficus hir!a, Achyrallthes aspera, Bridelia sp., Urena lobata, Sida rhombi/olio, Dioscorea sp., citrus) Neoseiulus longispinosus, Gupta, Oriental Ins., 12 : (Tamil Nadu : Bau~inia purpuria, castor, Zinia sp., beans, lady's finger, Pondicherry : Bauhinia purpuria) Amblyseius longispinosus, Gupta, Indian J. Acar., 2(2) : 66 (Re-described) (Orissa: guava; Bihar: rose) Amhlyseius longispinosus, Gupta & Nahar, In Contrib to Acar. in India. p. 9 (Bihar: paddy, infested with S. andropogoni) Amblyseius (Neoseiulus) longisp;nosus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Di~gnosis: Dorsal shield 325 long, 190 wide, smooth, well sclerotized, with 17 pairs of setae, mostly long excepting ji' J ls and SlS' which are shorter. Measurements of setae: ji-20, j4-54, j5-67, j6-72, J a -72, 15-7, ja-49, za-69, z4-76, s4-82, ZI-76, Sa-76, S4-59, S5-18, Z5-78, zls-31, Z4-69. Sternal, metasternal and genital shields normal. Ventrianal shield triangular, reticulate, longer than broad, with 3 pairs of setae and a pair of preanal pores. Spermatheca with funnel-shaped thick cervix. Macroseta only on basitarsus of leg IV-64. India (West Bengal, Bihar, Tamil Nadu, Pondicherry~ Andaman IsIs., Orissa), Philippines, Taiwan, Taipai, Thailand, South America, Tasmania, Indonesia, Malayasia. Remarks: Mallik (1974) studied its predatory importance. 74. Amblyseius (Neoseiulus) paspalivorus (De Leon) Typhlodromus paspa!pivorus De Leon, Fla En!., 40: (Holotype ~, Paratypes ~ ~ (1), USA: Florida, Coral Gables, on Paspalum sp.; holotype-in De Leon's Collection, paratypes in Univ. Fla. Collection. Gainsville ) A. paspalivol'us, Corpuz & Rimando, Philipp. Agric., 50: (Philippines: Oryza sativa) A. lula, Gupta, Sci. & Cult., 36: 98. (W. Bengal: Salix elegans)-misidentifi.. cation Amblyseius (Neoseiulus) paspalivorus, Ghai & Gupta, Bull. zool. Surv. India, 6(1-3) ; Amblyse;us (Neoseiulus) paspalivorus, Gupta, Fauna of India (Acari: MOIOIti~mata) Family Phrtoseiidae, p Q,

55 GUP~A : ORIENTAL PHYTOSElIDAE 41 Diagnosis: Dorsal shield 400 long, 193 wide, reticulate, specially posteriorly, except Z5' which is long, thickened and slightly pilose, others are small. Measurements of setae: jl. -11, j4-11, j5-l1, j6-l2, J a -13, J 5-12, js-12, za-10, Z4-9, s4-15, Z1-l3, S2-16, S4-30, S5-30, Z5-89, z~-ll, Z4-25. Ventrianal shield imbricate, longer than wide with 3 pairs of preanal setae and a pair of pores. Fixed digit of chelicera with 3 teeth, anterior to pilus dentilis, 2 teeth posterior to it; movable digit with one tooth. Cervix of spermatheca bell-shaped, atrium rounded. Macroseta on leg IV : basitarsus USA, Philippines, India (Delhi). 75. Amblyseius (Neoseiulus) rangatensis Gupta Amblyseius rangatensis Gupta, Oriental Ins., 11 : (Holotype ~, Paratype 1 ~, India: Rangat lsi., Yerata, on brinjal, both in ZSI.) Amb/yseius (Neoseiulus) rangatensis, Gupta, Fauua of India (Acari: Mesostigmata) Family Phytosoiidae, p Diagnosis: Dorsal shield 316 long, 200 wide, smooth, with 17 pairs of setae, all being short (6-10 long) except j1' ja' S4' Z5' Z4 which measure 16, 20, 32, 60 (weakly serrate) and 40 long, respectively. Sternal, meta sternal and genital shields normal. Ventrianal shield 100 long,92 wide, triangular with 3 pairs of preanal setae and a pair of pores. Macrosetae on leg IV : genu-40, tibia-nil, basitarsus-56. India (Andaman Nicobar IsIs.). 76. Amblyseius (Neoseiolus) siamensis Ehara & Bhandhufalck Amblyseius (Amblyseius) siamensis Ehara & Bhandhufalck, J. Fac. Ed. Tollor; Univ. Nat. Sci., 27 (2) : 63. (Holotype ~, Paratypes 2 ~ ~, Thailand; Prew, on Citrus; 1 ~, Thon BurL on citrus; 3 ~ ~, Thon Buri, on litchi: 1 ~ Thon Buri, on Acaripha sp., all in BioI. Inst. Fac. Ed. Tott, Univ.). Diagnosis: Dorsal shield heavily sclerotized, somewhat rounded with 17 pairs of setae and 7 pairs of pores. Zts sparsely barbed. Measurements of setae: ii-ii, j4-i each, J 5-9, ra-11, R l -10, is-10, z,-15, z.-lo, s4r-12, Zl-9, Sa-10, S4-10, Sts-B, Zts-4l. Ventrianal shield longer than wide. Fixed digit of chelicera with 2 small subapical teeth ~

56 42 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 and 6-7 middle teeth; 2 teeth on movable digit. Ventrianal shield longer than wide with 3 pairs of preanal setae. Leg IV with macrosetae on genu-12, tibia-13, basitarsus-30. Thailand. 77. Amblyseius (Neoseiulus) stolid~s Chaudhri Amblyseius sto/idus Chaudhri, Acar%gia, 10 : (Holotype ~, Pakistan : 6 miles south of Sheikhupura, on paddy, in UAL ; Paratypes 4 ~ ~, same data as for holotype, other ~ ~, on sweet patato, brinjal, citrus, mulberry, lucerne, deposited in USNM & BM (NH). Diagnosis: Dorsal shield 347 long, 182 wide, with 17 pairs of setae. Measurements of setae: j1-18, j4-j5-17 each, j6-19, J 2-25, J 5-14, js' za-24 z4-25, s4-26, Zl' Sa-S4-25 each, S5-24, Z5-53, (weakly serrate), z5-20, Z4-40 (weakly serrate), rs-25, R Ventrianal shield slightly concave near the anterior part, transversely striated with 3 pairs of preanal setae and a pair of preanal pores. Metapodal plates 2 paired. Cervix of spermatheca 4 times as long as wide. Fixed digit of chelicera with 5 subapical teeth and a pilus dentilis, one tooth onmovable digit. Leg IV with macroseta on basitarsus-60 long. Pakistan. 78. Amblyseius (Neoseiulus) taiwanicus Ehara Amblyseius (Amh/yseius) taiwanicus Ehara, Mushi, 43 (6): (Holotype ~, Bioi. Inst. Fac. Ed. Tott. Univ.). Paratypes 2 ~ ~, 3 0' 0', Taiwan : Fengshan, on pineapple, in Amh/yseius (Amh/yseius) taiwanicus Ehara & Bhandhufalck, J. Fac. Ed. Tottor; Univ. Nat. Sci., 27 (2) : 54 (Thailand: rice, grass, papaya). Diagnosis: Dorsal shield reticulate, with 17 pairs of setae and 7 pairs of pores; Z5 sparsely barbed. Measurements of setae: il-10, i4-j6-9 each, J a -10, J 5-8, r a -10, js-11, za-10, z4-10, s4-11, Z1-11, 8 a -12, S4-13, S5-16, Z5-44, z5-9, Z4-17. Ventrianal shield broadly rectangular, with 3 pairs of preanal setae and a pair of minute pores; 2 pairs of metapodal plates. Spermatheca with bell-shaped cervix. Basitarsus IV with macroseta-20 long. Taiwan, Thailand. In addition to the above species, some more species are known but those could not be included in key for the reasons as mentioned for Amblyseius (Alnb/yseius) s.p. on lla~e-14.

57 GUPTA: ORiENTAL PHYTOSElIDAE Amblyseius (Neoseiulus) venustus Chaudhri Amh/yseius venustus Chaudhri, Acarologia, 10 : (Holotype ~, Pakistan, 4 miles north of Sheikhupura, on Cynodon dactylon, in UAL ; Paratype 1 ~, same locality, on paddy; 1 ~, on fig (Paratypes in USNM and BM (NH). Diagnosis: Dorsal shield 340 long, 270 wide, with 17 pairs of setae and 3 pairs of pores. Measurements of setae: j 1-17, j4 -j 5-15 each, i 8-1S, , J 5-12, js-20, z2""19, z4-22, s4-24, , S2-26, S4-21, S5-20, Z5-54 (weakly serrate), z5-17, (weakly serrate, longer than the distance between its base and that of 2 5 ), rs-1s, R l -17. Ventrianal shield 117 long, 97 wide, triangular, sharp inward curve near anterolateral side with 3 pairs of preanal setae and a pair of pores. Cervix of spermatheca long. Leg IV with macroseta on basitarsus-73. Pakistan. so. Amblyseius (Neoseiulus) amantis (Chaudhri, Akbar & Rasool) Ansaria amantis Chaudhri, Akbar & Rasool, Rept. PL 480 Project, Univ. Agril. Faisalabad, Pakistan, p. 63. (Holotype ~, Pakistan: Charrapani, on Tamarix aphyl/a, in UAF). Diagnosis: Dorsal shield reticulate, setae blunt ended. Measure ments of setae: ]1-11, j4-j~-18 each, j6-21, J~.. 23, J 5-13, is, za 18 each, 'z4:-21, s4-23, Zl-21, S2-26, S4-26, Z5-31, z5-18, 24-23, ra-13, R l -16. Ventrianal shield 96 long, 52 wide. Macrosetae on basitarus IV 26 long. Pakistan. S1. AmbJyseius (Neoseiulus) bheraensis (Chaudhri, Akbar & Rasool) Neoseiulus hheraensis Chaudhri, Akbar & Rasool, Rept. PI 480 Project, Univ. Agril. Faisalabad, Pakistan, p. 76. (Holotype ~,Pakistan : Bhera, on Mangl1era indica, in UAF). Diagnosis: Dorsal shield 343 long, 146 wide. Measurements of setae jl-8, j4-8, h;-8, i6-8, J 2-10, J 5-10, i s -lo, z2-8, z4-s, S4-10, , S2-13, , Z5-62, z5-8, , ra-s, R l -10. Ventrianal shield 104 long, 91 wide. Spermatheca cup or bowl-shaped; cervix poculiform, atrium nodular. Macroseta on basitarsus IV -31. Pakistan.

58 44 REC. ZOOL. SURV. INDIA, OCC. PAPER No Amblyseius (Neoseiulus) caruncula (Chaudhri, Akbar & Rasool) Neoseiulus caruncula Chalidhri, Akbar & Rasool, Rept. PL 480 Project, Univ. Agril. Faisalabad, Pakistan, p. 83. (Holotype ~,Pakistan: Swat, on undet. plant, in UAF). Diagnosis: Dorsal shield 326 long, 185 wide. Measurements of setae: jl-16, j4-13, j5-16, j6' J a-18 each, i a -18, Za -= Z4 = 16, 84 =Zl = 23, S2-21, S4-18, Z5-52, z5-16, s4-18, Z4-36, ra-18, R l -16. Ventrianal shield 117 long, 104 wide. Macrosetae on basitarsus IV-65. Pakistan. 83. Amblyseius (Neoseiulus) curvus Wei-nan & Zhao-Quan Amhlyseius (Alnh/yseius) curvus Wei-nan & Zhao-Quan, Acta Zootaxonomica Sin ica, 10 (4) : (Holotype ~,Paratype 1 ~,South China: Jinafengling, Hainan lsi., on undet. plant). Diagnosis: Dorsal shield long, wide. Measurements of setae: jl-27-30, j , i , j6-42, J a -53, , 1a-42, za-40-42, Z , S , ZI-52, S , S , S&-52-65, Z , z , Z , ra-45-47, R l , JV South China. 84. Amblyseius (Neoseiolus) eremicos (Chaudhri, Akbar & Rasool) Neoseiulus eremicus Chaudhri, Akbar & Rasool, Rept. PL.480 Project, Univ. Agril. Faisalabad, Pakistan, p. 83. (Holotype ~, Pakistan: Faisalabad, on Gossypium hirsutum, in UAF). Diagnosis: Dorsal shield 282 long, 194 wide. Measurements of setae: jl-19, j4-16, j5-15, is, Ja-18 each, 1 5-8, i s -23, za-24, z4-26, s4-32, ZI-26, S2-26, S4-26, Z5-42, z5-16, Z -18, r a-24, Rl -13. Ventrianal shield 97 long, 48 wide. Spermatheca with cervix long, 48 wide. Macroseta on basitarsus IV.. 2S. Pakistan.

59 GUPTA: ORIENTAL PHYTOSEllDAE Amblyseius (Neoseiulus) Myrtia (Chaudhri, Akbar & Rasool) Neoseiulus myrtia Chaudhri, Akbar & Rasool, Rept PL 480 Project, Univ Agril. Faisalabad, Pakistan, p. 81. (Holotype ~,Pakistan: Mian Channu, on Arundo donax, in UAF). Diagnosis: Dorsal shield 388 long, 204 wide. Measurements of setae: jl-26, i 4, j6-16 each, J a-18, J 5-13, ja = S4-29 each, Za = Z4-21 each, Zl = Sa = S4-26 each, ZI5-86, zl5-23, Z4-34, rs-23, R Ventrianal shield 133 long, 104 wide. Pakistan. Macroseta on basitarsus IV Amblyseius (Neoseiulus) reductus (Chaudhri, Akbar & Rasool) Kashmiricus reductus Chaudhri, Akbar & Rasool, Rept. PL 480 Project, Univ. Agril. Faisalabad, Pakistan, in UAF). (Holotype ~, Pakistan: Chenari, on undet. plant, in UAF). Diagnosis: Dorsal shield 364 long, 198 wide, with 17 pairs of state: i 1-16, j4-13, jg-13, i 6-13, Ja-16, J5-18; ja-21, za-21, z4-16, s4-23, Zl-18, Sa-13, Z5-60, z5-13, Z4-18, ra-21, R l -16. Ventrianal shield quadrate, convex anteriorly, cncave laterally, with 3 pairs of preanal setae. Macroseta absent on leg IV. Cervix of spermatheca poculiform. Dis tribution: Pakistan. 87. Amblyseius (Neoseiulus) tabularis (Chaudhri, Akbar & Rasool) Neoseiulus tabu/oris Chaudhri, Akbar & Rasool, Rept. PL 480 Project, Univ. Agril. Faisalabad, Pakistan, P. 78. (Holotype ~,Pakistan: Pirowala forest, on an undet. plant, in UAF). Diagnosis: Dorsal shield 337 long, 143, wide. Measurements of setae: jl-10, j4-ja-8 each, J 2-10, J5-10,ja-s4-8each, Z1-10, S2-21, S4-18, Z5-70, zc;-8, Z4-18, r a -13, R 1-8. Macroseta on basitarsus IV -31. Male; Pakistan.

60 46 REC. ZOOL. SURV. INDIA, Occ. PAPER No. 95 Subgenus Parapbytoseius Swirski & Shechter Key to the species of Paraphytoseios 1. Zu z... serrate... sclerolicus Zu z. smooth... multidentatus 88. Amblyseios (Paraphytoseios) moltidentatos (Swirski & Shechter) Paraphytoseius multidentatus Swirski & Shechter, Israel J. agrie. Res., 11(2) : (Holotype ~,Hong Kong: on Bambusa sp., Paratyp~s 2 ~ ~,Hong Kong, on Jasm;llum sp., 1 ~ on Pomelo, 1 ~, on Ageratum houstonianum, Urena lobata, Cyclosorus subtescens, all in Univ, Inst. Agr., Israel) Typhlodromus (Amh/yseius) orientalis Narayanan, Kaur & Ghai, Proc. natn. Inst. Sci. India~ 26 (B) (6) : 394. (This was described based on 6 ~ ~, 20' C, without "designating the holotype and paratypes in the original publication, in NPC) Amblyseius (Paraphytoseius) narayanani Ehara & Ghai, Ehara, Mushi, 40(6) : 7 ' Amblyseius bhadrakaliensis Gupta, Bull. En!., Ent. Soc. India, 10(3): (West Bengal ~ Datura metel) Amblyseius horrifer, Gupta, Sci. & Cult., 36 : 98. (West Bengal: Polygonum sp.) Amb(vseius horri/er, Gupta, Internat. J. Acarol., 1(2) : 37. (West Bengal: Polygonllm sp.) Amblyseius hhadrakaliensis, Gupta, Internat. J. Acarol., 1(2) : Amblyseius narayanani, Gupta, Internat. 1. A carol., 1(2) : Amblyseius bhadrakaliensis, Gupta, Oriental Ins., 11 : 626. (Andaman lsi. : paddy, undet. plant, Cajanas cajan, Rubus sp., brinjal) Amblyseius (Paraphytoseius) multidentatus, Ehara & Bhandhufalck, J. Fac. Ed. Tottori Unlv. Nat. Sci., 27(2) : (Thailand: Urena lobata, bamboo, Cratoxylum, Eupatorium odoratum, Triumfetta rhomboides, Dioscorea sp.) Ambiyseius nicobarensis Gupta, Oriental Ins., 11 : Nicobar lsi. : on undet. plant) A111biyseius bhadrakaliensis, Gupta, Indian J. Acar., 2(2) : 6t. (Bihar: brinjal; Orissa: brinjal; Meghalaya: Undet. plant: Assam: Undet. plant, fern, Shorea robusta; Tripura : Undet. plant) Paraphyloseius (Paraphytoseius) bhadrakaliellsis, Gupta, Bull. zool. Surv. India, 2(1) : (Tamil Nadu: banana, pear) beans, sunflower, cashewnut, brinjal, Eucalyptus, tea, rose; Kerala : rose; Karnataka: Undet. plant; Andhra Pradesh: cashewnut. ).

61 GUPTA: ORIENTAL PHYTOSEIIDAE Paraphytoseius multidentatus, Gupta, Bull. zool. Surv. India, 2(1) : Paraphytoseius multidentatus, Chaudhri, Akbar & Rasool, PL 480 Project, Univ. AgriI. Faisalabad, Pakistan, p Amblyseius (Paraphytoseius) nicoharensis, Gupta & Ray, Bull. zool. Surv. India, 4( 1) : Amhlyseius (Paraphytoseius) narayanani, Gupta & Ray, Bull. zool. Surv. India, 4(1) : (Tripura: Eupatoriuin odoratum) Amblyseius (Paraphytoseius) mu[tiden to tus, Gupta & Ray, Bull. zool. Surv. India, 4(1) : (Tripura : Eupatorium odoratum) Paraphytoseius hhadrakaliensis, Gupta & Nahar, In Contrib. to Acar in India, p. 10. (Bihar: Hibiscus mutabi/is) Amh/ysejus (Paraphytoseills) bhadrakaliensis, Gupta, Bull. zool. Surv. India, 4(1) : 45. J986. Amh/yseius (Paraphyloseius) mullidenlallls, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield incised near S4' with 14 pairs of setae and some pairs of pores, those near z5 very large. Setae j1' js' S4' Z~, Z4 being long, thick and serrate measuring 30, 88, 120, 130 and 70, respectively; other setae small; S~-7 long; rs and R1 on interscutal membrane. Ventrianal shield 95 long, 55 wide, lateral margins slightly concave, with 3 pairs of preanal setae and a pair of pores. Fixed digit of chelicera multidentate, movable digit 3 dentate. Macrosetae on leg IV : genu-29, tibia-38, basitarsus-45, distitarsus-45, tip with hyaline envelope. Ehara & Bhandhufalck (1977) figured spermatophoral process. Hong Kong, Thailand, Philippines, Pakistan, India (Tripura, West Bengal, Tamil Nadu, Kerala, Karnataka, Andhra Pradesh, Assam, Meghalaya, Andaman and Nicobar Is1., Bihar, Orissa). 89. Amblyseius (Paraphytoseius) sclerotic us Gupta & Ray Amblyseius (Paraphytoseius) sclerotjeus, Gupta Ray, Bull. zool. Surv. India, 4(1) : (Holotype ~, India: Uttar Pradesh, Barkot, on an undetermined plant, in ZSI.) Amblyseius (Paraphytoseius) scleroticus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 260 long, 135 wide, with 14 pairs of setae; setae i 1, is, S4' Z4' Z5 and rs being long, thick and serrate and measure;

62 48 REC. ZOOL. SURV. INDIA, Dcc. PAPEa No , 56, 69, 45, 69 and 20, respectively; za-11, (also serrate). Sternal and genital ~hields normal. Ventrianal shield 90 long, 50 wide, with 3 pairs of preanal setae. Macrosetae on leg IV: genu-1s, tibia-29, basitarsus-34 and distitarsus-34; all with knobbed tip. India (Uttar Pradesh). In addition to the above, one more species i~ known which is not included in key because its status is doubtful. The species is as below : 90. Amblyseius (Paraphytoseius) cracentis (Corpuz & Rimando) Ptenoseius cracentis Corpuz & Rimando, Philipp. Agr., 50 : (Holotype ~, Philippines: Gamu, Isabela, on Achyranthes aspera ; in Dept. Ento. Univ. Philip.) Paraphytoseius cracelltis, Schicha & Corpuz-Raros, Internat J. Acarol., 11(2): Diagnosis: Dorsal shield 300 long, 130 wide. Me~surements of setae: j1-35, is-96, s4-144, Z4-86, Z5-112 Metasternal plate trape~ zoidal. Leg IV with thickened macroseta on genu-20, tibia-42, basitarsus-52.. distitarsus-48, all with bulbous tip. Cervix of spermatheca with shallow arc. Philippines, New Cledonia. Subgenus Phytoscutella Muma Key to the species of Phytoscutella 1. Dorsal shield rugose caudad of setae j6 ; ventrianal shield with 3 pairs of preanal setae salebrosus. Dorsal shield smooth, caudad of setae j&; ventrianal shield with S pairs of preanal setae wongs;rii. 91. AmbJyseius (Phytoscutella) salebrosus (Chant) Typhlodromus (Amblyseills) salebrosus Chant, Can. Ent., 92 : (Holotype ~, paratypes 2 ~ ~, 20' 0, India: Assam, J orhat, on citrus) hl Canadian National Collection, Belleville; another paratype ~, abov~ locality and habitat, in Citrus Expt. Stn., Riverside, California) Amblyseius sa/ebrosus, Gupta, Bull. Sun- Yat Sen Cult, Found., 5 : (Taiwan : citrus).

63 GUPTA: ORIENTAL PHYTOSElIDAE Amblyseills (Phytoscutella) salebroslis, Ehara & Bhandhufalck, J. Fac. Ed. Tollori Univ. Nat. Sci., 27(2) : (Re-described). (Thailand: Sugarcane, citrus, Hibiscus sp.) Amblyseius (Phytosclltella) salebrosus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield rugose, sclerotized, with 15 pairs of setae, jls' 1s absent. Measurements of setae: jl-12, j4-4, j6-i1, J 5-4, ja-16, za-9; z4-13, s4-134, ZI-9, Sa-52, S4-16, S5-16, ZlS-202, z5-4, Z Ventrianal shield massive, reticulate, with 3 pairs of preanal setae. Spermatheca with sacular cervix. Leg IV with macrosetae on genu-34, tibia-45, basitarsus-24. Chant (1960) described male. India (Assam), Taiwan, Thailand. 92. Amhlyseius (Pbytoscutella) wongsirii Ehara & Bhandhufalck Amblyseius (Phytoscutella) wongsirii Ehara & Bhandhufalck, J. Fac. Ed. Tottori Univ. Nat. Sci., 27(2) : 75. (Holotype ~, Paratype 1 ~, Thailand: Prew, on citrus, in BioI. Inst. Fac, Ed., Tott. Univ.). Diagnosis: Dorsal shield sclerotized, lateral margins rugose, with 15 pairs of setae; j5' 1a absent. Measurements of setae: jl-28, j4t-9, ;6-24, J 5-13, ja-64, za-65, Z4-53, S4-169, ZI-26, Sa-174, S4-48, S5-48, , z5-10, Z Ventrianal shield wider than long, about twice as wide as genital shield. Fixed digit of chelicera multidentate, movable digit tridentate. Macrosetae on leg IV: genu-loa, tibia-70, basitarsus-57. Thailand. Subgenus Proprioseiopsis Muma 93. Amhlyseius (Proprioseiopsis) peltatus Van der Merwe Amblyseius (Amblyseius) peltatus Van der Merwe, Ent. Mem. Dept. Agr. Tech. Servo Repub. S. Afr., 18 : Amblyseius rosel/us, Gupta: Sci. & Cull., 36 : 98. (West Bengal: fern) Amblyseius (Proprioseiupsis) peltatus, Ebara & Bhandhufalck, J. Fac. Ed. Totlori Univ. Nat. Sci., 27(2) : (Re-described). (Thailand: Combretum quadrangulare, grass, AcI,yranthes aspera, Bridelia sp., soy-bean, Dioscorea, Citrus, Corn). 7

64 50 REC. ZOOL. SURV. INDIA, Oce. PAPER No IphiseillS punicae Gupta, 1980, EIlIOI1IO/ogisfs' Mag., 115: 213 (West Bengal: Punlca granatuln) Amb/yseius (Proprioseiopsis) peitallls, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 315 long, 232 wide, lateral margins sclerotized. Measurements of setae: jl-27, j4' j6-4 each, js-54, za-3l, z4, 22, S4-93, Z1-18, S9-22, S4-10, S~-8, Z~-80, zr;-4, Z4-93. Sternal shield reticulate, Ventrianal shield robust, triangular, reticulate, 90 long, 108 wide, with 3 pairs of preanal setae and a pair of preanal pores. Spermatheca with funnel-shaped cervix. Macrosetae on leg IV: genu-53, tibia-37, basitarsus-76. India (West Bengal), Thailand, South Africa. In addition to the above species, the following are also known but could not be included in the key for the same reasons as nlentioned for Amblyseius (Amb/yseius) spp. on page Amblyseius (Proprioseiopsis) arunachalensis Gupta Anlblyseius (Proprioseiopsis) arullachalensis Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p (Holotype ~, India: Arunachal Pradesh, Likabali, on an undet. plant. in ZSI). Diagnosis: Dorsal shield 336 long, 245 wide, with 17 pairs of setae. Measurements of setae: jl-24, j4 -j6-minute, js-15, za-8, Z4-8, S4-201, Sa-S5-minute, Z5-400, z5-minute, Z4-194, rs, R l -13 each. Ventrianal shield 112 long, 120 wide, with 3 pairs of preanal setae. Macrosetae on leg IV : genu-138, tibia-67, basitarsus-30. India (Arunachal Pradesh). 95. Amblyseius (Proprioseiopsis) beatus Chaudhri Amblyseius beatus Chaudhri, Acarologia, 10: SSI-553. (Holotype ~, Pakistan: West Pakistan Agril. Univ. campus, on Triticum aestivum, in UAL; Paratypes 1 ~, on : rass ; I ~ on debris, in USNM and B.M. (N.H.). Diagnosis: Dorsal shield 367 long, 279 wide, oval with 16 pairs of setae and 3 pairs of small pores. Measurements of setae : jl-26, j*-j~-8 each, j6-16, J~-missing, J~-13, j8-42, z2-28, z4-62, s4-78, Zl-29,

65 GUPTA: ORIENTAL PHYTOSEIIDAE 51 5 a -36, , S5-26, Z5-78 (weakly serrate), z&-8, Z4-75 (serrate), ra, R 1-16 each. Ventrianal shield reticulate, triangular, with 3 pairs of preanal setae..macrosetae on leg IV : genu-36, tibia-53, basitarsus-81. Male; Pakistan. 96. Amblyseius (Proprioseiopsis) bulga (Chaudhri, Akbar & Rasool) Proprioseiopsis bulga Cha udhri, Akbar & Rasool, Rept. PL 4~0 Project, Univ. Agril. F aisala bad, p. 50. (Holotype ~, UAF). Pakistan, Abbotabad, UAF campus, on Ocimuln basilicum, in Diagnosis; Dorsal shield 367 long, 260 wide. Measurements of setae: jl-13, j4-j5-nlinute, js-8, J 5 -minute, ja-49, za-26, z4-34, s4-68, Zl == Sa== S4-8 each, Z5-96, Z4-75, r a -26, R I -13. Ventrianal shield pentagonal. Macrosetae on leg IV : genu-55, tibia-30, basitarsus-65. Pakistan. 97. Amblyseius (Proprioseiopsis) dentatus (Chaudhri, Akbar & Rasool) Proprioseiopsis delllalus Chaudhri, Akbar & Rasool, Rept. PL 480 Project, Univ. Agril. Faisalabad, p. 52. (Holotype ~,Pakistan, UAF can1pus, on Ca/otl'opis, in UAF). Diagnosis: Dorsal shield 347 long, 244 wide. Measurements of setae: i I - 29, j4-js-minute, J 5 -minute, js-47, za-36, z4-62, s4-83, ZI-41, 5 a -36. S4-29, Z&-75, Z4-68, ra-23, R I -16. Macrosetae on leg IV : genu-31, tibia-31, basitarsus-83. Pakistan. 98. Amblyseius (Proprioseiopsis) synachattiensis Gupta Anzblyseius (Proprioseiopsis) synachattiellsis, Gupta, Entomon, 10(3) : 211. (Holotype ~, India: Uttar Pradesh, Garhwal Himalaya, Synachatti t, on grass, in ZSI) Amblyseius (Proprioseif1psis).iYllachaltiensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p

66 o 52 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 95 Diagnosis: Dorsal shield 504 long, 292 wide, rugose. Measurements of setae: j1-34, j4-5, j6-20, js-54, za-36, z4-52., S4-90, Z1-25, Sa-18, S4-13, SI)-16, ZI)-121, zl)-8, Z4-90, ra-31, R Spermatheca ~ith bell-shaped cervix. Macrosetae on leg IV: genu-20, tibia-40, basitarsus-34, all pointed. India (Uttar Pradesh). Subgenus Proprioseius Chant 99. Amhlyseius (Proprioseius) Kumaonensis (Gupta) Proprioseius kumaollensis Gupta, Indian J. Acar., 6: (Holotype ~,India : Uttar Pradesh, Kumaon Garhwal Himalayas, on undet. plant, in ZSI) Alnblyseius (Proprioof\eius) ]{ui11aonellsis, Gupta, Fauna of India (Acari: Mesostign1ata) amily Phytoseiidae, p Diagnosis: Dorsal shield 312 long, 252 wide, highly rugose, well sclerotized with 14 pairs of setae. Measurements of setae: jl-36, i4:-5, jl)-5, i 6-7, J 5-4, js-53, zs-15, z4-27, s4-87, Sa-17, , Z5-72, Z6-5, Z4-50, ra-20. Ventrianal shield 96 long, 60 wide, lateral margins concave with 3 pairs of preanal setae. Macrosetae present on genu, tibia and basitarsus of leg IV, all being spatulate. India (Uttar Pradesh). Subgenus Typhlodromalus Muma Key to the species of Typhlodromalus Ventrianal shield vase-shaped Ventrianal shield squarish or pentagonal js and Z2 equal or j. only slightly longer than Z2 js more than 2 times as long as Z2 Z, reaches almost upto bas~ of Za Z, much shorter than distance between i.ts base and that of ZI5 ja and Z2 almost equal or ja only slightly longer ja 2 times as long as Z2 Z2 and Z4 equal Z, 2 alid l/2 times as long as Z2 Z4, reaches beyond base of S 15 Z., much smaller than the distance between its base and that of S ]{a!ilnpongellsis 3 jarooa sorghumae S eucalypticus 6 Inanipurensis lablabi newsami

67 GUPTA: ORIENTAL PHYTOSEUDAE Amhlyseius (Typhlodromalus) eucalypticus (Gupta) Typhlodromalus eucalypticus Gupta, Oriental Ins., 12 : (Holotype ~, India: Karnataka. Bandipur Sanctuary, on Eucalyptlls sp., in ZSI) Amblyseius (Typhlodromalus) eucalypticus Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield with anterior lateral margins imbricate with 17 pairs of setae. Measurements of setae: jl-24, j3-20, s4-28, Zlr 64, Z4-24; other setae 5-6 long. Ventrianal shield longer than broad, with 3 pairs of preanal setae. Spermatheca with elongated flaskshaped cervix, bulbous atrium and long duct. Leg IV with macrosetae on genu-48, basitarsus-60. India (Karnataka) Amblyseius ( Typhlodromalus ) jarooa Gupta Al1lblyseius jarooa Gupta, Oriental Ins., 11 : 629. (Holotype ~, Andaman lsi... Wandrau, on undetermined plan, in ZSI ; Paratypes 2 ~ ~,lis, Ferrargunj, Manpur, on an undet. plant, in ZSI) Amb/yseius swaga, Gupta, Indian J. Acar., 2(2) : 73 (Assam: Tectona grandls) Amb/yseius (Typhlodromalus) jarooa Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis; Dorsal shield 320 long, 180 wide, with 17 pairs of setae and a pair of pores above the level of Z4. Dorsal setae nlostly small except jl' ja, S4' Z5 and Z4: which are longer and measure: 24, 20, 40, 88 (serrate) and 48, respectively. Sternal and genital shields normal; metasternal plate absent. Spermatheca funnel-shaped. Leg IV \vith macrosetae on geuu-53, tibia-38 and basitarsus-53. shields. Distribution.: Gupta (1977) illustrated the ventrianal and sternitigenital India (Andaman IsIs.) Amblyseius (Typhlodromalus) kalimpongensis Gupta Amblyseius kalimpongensis Gupta, Bull. ent., Ent. Soc. India, 10: (Holotype ~ 1 Paratypes 1 ~, 1 ~, India: West Bengal, KaUmpong, on Citrus reticulata. in ZSI) Amb/yseius hitna, Gupta, Sci. & Cult., 36 : 98. (West Bengal: Rupellia sp.)-misidentification Anlblyseius kalimpongens;s, Gupta, Internal. J. Acal'oi., 1(2): 37 (West Bengal: Hibiscus mutabilis).

68 54 REC. ZOOL. SURV. INDIA, OCC. PAPER No Amblyseius hima, Gupta, Internat. J. Acoro!., 1(2) : 37. (West Bengal: Hibiscus mutabilis) Amblyseius kalimpoligensis, Gupta, Indian J. Acar., 1 : 32. (Gujarat : Undet. plant, citrus, weed} guava) Typhlodromalus hima, Gupta, Oriental Ins., 12: 331 (Tamil Nadu : Magnolia grandijlora, maize, Bougainvillea sp., Ladys' finger) Typhlodromalus kalimpongensis, Gupta, Oriental Ins., 11: 331 (Tamil Nadu : banana, brinjal, Karnataka : Undet. plant; Andhra Pradesh: dahlia) Amblyse;us kalimpongensis, Gupta, Indian J. Acar., 2(2) : 63 (Bihar: Jasminum, Orissa: Shorea sp., cashew, citrus; Assam: Undet. plant) Amblyseius (Typhlodrolnalus) Kalbnpongensis, Gupta, rauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield smooth with 17 pairs of setae and 5 pairs of pores. Measurements of setae: j1-j each, J 5-6, ja-27, za-24, Z4-27, S4-31, Z1-17, Sa-20, S4-18; S5-26, Z5-40, z5-16, Z4-18. Sternal and genital shields normal, metasternal plates absent. Ventrianal shield vase-shaped, strongly concave laterally, with 3 pairs of preanal setae and a pair of preanal pores. Leg IV with macroseta only on basi tarsus. Gupta (1969) figured the spermatophoral process and ventrianal shield. India (West Bengal, Gujarat, Tamil Nadu, Karnataka, Andhra Pradesh, Bihar, Orissa, Assam) Amblyseius (Typhlodromalus) lablabi Ghai & Menon Amblyseius lablabi Ghai & Menon, Orielltal. Il1s., 1 : (Holotype ~ J several paratypes ~ ~, India: Karnataka, Bangalore, on Dolichos lablab, in NPC) Amblyseius lablabi, Gupta, Internat. J. Acaro/., 1 (2) : Amblyseius (Typhlodronlalus) lablabi, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield reticulate, with 17 pairs of setae; setae j S l>jl =za ; Z4 and S4 progressively longer, j4' j5 and z~ shorter than rest of the dorsal setae. Posterior margin of sternal shield lobed. Ventrianal shield reticulate, penta8onal, longer than broad, anterior margin rounded, lateral margins concave, with 3 pairs of preanal setae and a pair of pores. Leg IV with pointed macrosetae on genu, tibia and basitarsus, that on tibia. being shorter. India (Karnataka).

69 GUPTA: ORIENTAL PHYTOSElIDAE Amblyseius (Typhlyodromalus) manipurensis Gupta Amblyseills nlol1ipurensis Gupta, Indian J. Acar., 2(2) : (Holotype ~,India: Manipur, Ukhral, Hibiscus esculentus) Amb,yseius (Typhlodromalus) lnanipurensis, Gupta, Fauna of India (Acari : Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 308 long, 180 wide, smooth with 17 pairs of setae. Measurements of setae: j1-22, j4-j6-7 each, 1a-12, J 5-8, js-12, za-8, z4-24, S4-40, Z1,-12, 8a-16, each, Z5-68 (gently serrare), zr;-8, Z4-48 (smooth). Ventrianal shield 92 long, 64 wide, with 3 pairs of preanal setae and a pair of pores. 8permatheca with tubular cervix. Macrosetae on leg IV : genu-28, tibia-28, basitarsus-64. Ind ia (Manipur) Amblyseius (Typlodromalus newsami Evans Typhlodromus newsami Evans, Ann. Mag. nat. Hist., (12)6 : (Holotype ~,Paratype 1 (J, Malaya, on rubber, in BM(NH» Amblyseills (Amblyseius) newsami, Ebara & Bbandhufalck, J. Fac. Ed. Tolt. Univ. Nat. Sci., 27(2) : (Thailand: citrus, litchi, okra, banana). Diagnosis: Dorsal shield reticulate only along anteror lateral margins with 17 pairs of setae and 10 pairs of pores; Z 5 smooth. Measurements of setae: j1-24, j4-j each, 1a-14, 1 5-8, rs-13, R l -13, is-11, za-10, z4-11, , Z1-11, S2-14, S4-13, S5-13, Z5-45, z5-9, Z4-14. Metasternal setae on plates. Ventrianal shield vase-shaped, widest at anal region with 3 pairs of preanal setae and a pair of conspicuous cresentic pores. Cervix of spermatheca tubular. Fixed digit of chelicera multidentate, movable digit with a few weak teeth. Macrosetae on leg IV : genu-50, tibia-38, basitarsus-49. Malaya, Hong Kong, Thailand Amblyseius (Typblodromalus) sorghumae Gupta Amb/yseius sorghumae Gupta, Oriental Ins., 11 : (Holotype ~, India: Car Nicobar lsi., on maize; Paratypes 2 ~ ~ same data as for holotype, all in ZS]) Amb~vseius (Typhlodromalus) sorghllmae, Gupta, Fauna of India (Acari: Mesos.. tigmata) Family Phytoseiidae" Diagnosis: pa irs of setae. Dorsal shield 330 long, 225 wide, smooth, with 17 Measurements of setae: jl-18, j4-j 6 1 J 2, J each,

70 56 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 ja-11, z2' z4-9 each, S4-16, 2 1, S2-S fs-7 each, 25-72, z&-5, Z4-17 ~ Sternal, metasternal and genital shields normal. Ventrianal shield 92 long, 64 wide, lateral margins concave at preanal region. Spermatheca with filamentous cervix. Leg IV with macro setae on genu-48, tibia-"40, basitarsus-36. India: Andaman-Nicobar IsIs. In addition to the above, the following species-of this subgenus are also known from the Oriental region but could not be included in key for late publications of those species AmhJyseius (Typhlodromalus) chikmagalurensis Gupta A 111blyseius (Typhiodromaills) chikmagalurensis Gupta, Env. & Ecol., 3(3): (Ho!otype ~,India : Karnataka, Chikmagalur, on undet. plant. in ZSI) Amblyseius (Typhlodronlalus) chikmagalurensis, Gupta, Fauna of Indja (Acari: Mesostigmata) Family Phytosei;dae, p Diagnosis: Dorsal shield 274 long, 150 wide. Measurements of setae: jl-22, j4-22. je-18, 1 a-16, 15.8, js-22, z2-25, Z4-29, S4-37, ZI-16, S2-13, S4-22, S5-18, 2&-45, z5-21, Z4-18. Ventrianal shield 78 long, 50 wide. Macrosetae on leg IV : basitarsus-49. Male; India (Karnataka) Amblyseius (Typhlodromalus) cbitradurgae Gupta A,nblyseius (Typhlodromalus) chitradurgae Gupta, Env. & Ecol., 3(3) : (Holotype ~, Parat) pes 11 ~ ~ 1 is, India: Karnataka, Chitradurga. in ZSI) Amblyseius (Typhlodromalus) chitradurgae. Gupta, Fauna of India (Acari: Meso~tigmata) Family Phytoseiidae. p Diagnosis: Dorsal shield 292 long, 146 wide, rugose. Measurements of setae: jl-20, j4-l6, j5-16, je-16, l a -18, 15-5, js-27, za-27, z4-29, s4-35, , S2-25, S4-24, S&-22, ZG-44, z5-20, 24-22, rs 25, R Ventrianal shield 78 long, 56 wide. Macroseta on basitarsus IV-31. Gupta (1985) figured the spermatophoral process. India (Karnataka).

71 GUPTA: ORIENTAL PHYTOSElIDAE AmbIyseius (Typhlodromalus) ficusi Gupta Alnblyseills (Typhlodron:alllJ) jiclisi Gupta, Otiell/cli Ins., (In press). (Holotype ~ Paratypes 3 ~ ~, India: Arunachal Pradesh, Kambang, on Ficus sp., in ZSI) Anlblyseius (Typhlodr. malus) ficusi, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis; Dorsal shield 292 long, 157 wide. Measurements of setae: j1-13, j4-13, j5-11, is-13, Ja-IO, 1 5-5, ja-16, za-11, z4-20, s4-25, Zl-11, Sa-13, S4-9, S5-9, Z~-49, z5-9, Z4-22, r a -13, R1-IO, Ventrianal shield 78 long, 50 wide, with 3 pairs of preanal setae. Macrosetae on leg IV: genu-25, tibia-25, basitarsus-47. India (Arunachal Pradesh) Amhlyseius (TyphlodromaIus) Iaaensis Gupta Amblyseius (Typhlodromalus) laaensis Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p (Holotype ~,Paratype 1 ~, India: Arunachal Pradesh, Laa, on tea, in ZSI). Diagnosis: Doral shield 270 long, 130 wide, weakly reticulate anteriorly. Measurements of setae: jl-16, j4-j5-4-5 each, is-10, J 5-8, ja-29, za-7, z4-22, S4-40, ZI-13, S2-27, S5-13, Z5-40, z5-6, Z4-25, rs-22, R1-lj. Ventrianal shield 90 long, 56,vide, with 3 pairs of preanal setae. Macroseta on basitarsus IV -36. India (Arunachal Pradesh). Subgenus Typblodromips De Leon Key to the species of Typhodromips 1. J 2 absent J 2 present 2. Besides Z" and Z." other setae being subequal Besides Z" and Z 4 there are other long setae 3. Z2 and Z4 equal Z2 and Z4 unequal 4. Zs 3 times the length of Z2 Zs less than 2 times the length of Z2 s. Z4 much longer (over 4 times) the length of S5 Z4 as long as 85 8 len is po ten tillae 5 6 Inangiferot:

72 58 REC. ZOOL. SURV. INDIA, OCC. PAPER No S'l over 4 times the length of Z 1 S2 only slightly longer than Z 1 7. Z4 and S4 almost equal. z. and s. unequal 8. Z 4 reaches almost upto the base of Z;; Z. shorter than the distance between Z4 and Z,'l 9. Macroseta on basitarsus IV over 1 ~ times as that of genu IV Macroseta on basitarsus IV much less than It times as that of genu IV 10. Sternal shield with indentation posteriorly Sternal shield without indentation posteriorly 11. Z. quite long at least 3 times the length of s" Z. not so long 12. Ventrianal shield as long as wide Ventrianal shield longer than wide t 3. Ventrianal shield notched on lateral sides Ventrianal shield not notched 14. Macroseta on genu IV longer than that on tibia IV Macroseta on genu IV shorter than that on tibia IV 15. Z. distinctly serrate Z. smooth 16. Z" reaches almost upto base of S fi Z. much shorter than the distance between its base and that of Z Macroseta on leg IV knobbed Macroseta on leg IV simple t 8. Spermatheca obscured Spermatheca prominent 19. Dorsal shield reticulate throughout Dorsal shield smooth or reticulate marginally 20. Z. reaches almost upto the base of ZfJ Z. much shorter than the distance between its base and that of Z5 21. Macrosetae on leg IV knobbed Macrosetae on leg IV smooth 22. Macrosetae on genu IV knobbed, others mostly smooth Macrosetae on genu. tibia and basitarsus of leg IV knobbed 23. Macrosetae on gen~ I V and tibia IV almost equal Macrosetae on genu IV much longer (over 2 times) than that of tibia IV 24. Cervix of spermatheca funnel-shaped Cervix of spermatheca tubulclf ". tefranychivorus simi/is 9 8 crotalariae neocrotalariae 10 eujeniae hidakai guajavae IS sapienticoia ghanii ojji.cinaria polyantheae eghaiayensis 17 iabis are cae johoreae 19 syzyg;; suknaensis asiaticus Iineari.4: 24 perfectus okinawanus vignae

73 OurTA: ORIENTAL PHYTOSEI1DAE 59 Ill. Amblyseius (Typblodromips) areeae Gupta Amblyseius arecae Gupta, Oriental Ins., 11 : (Holotype ~, Paratypes 5 ~ ~, 1 C, Andanlan IsIs., Aniket, on arecanut, in ZSI) Alnb(vseius (TyphlodroI11ips) arecae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield well sclerotized with 17 pairs of setae, mostly small except Z5 and Z4 which are longer measuring 90 and 26, respectively. Sternal, metasternal and genital shields normal. Ventrianal shield squarish, concave at preanal region with 3 pairs of preanal setae. Spermatheca with elongate cervix and knobbed atriun. Macrosetae on leg IV : genu-44, tibia-32 and basitarsus-48. Gupta (1977) figured its spermatophoral process Amblyseius (Typhlodromips) asiatic us (Evans) Typhlodromlls asia/ieus Evans, Ann. Mag. nat. His!., (12)6: (Holotype ~, Indonesia : Bogor, on leaves infested with tetranychids, in BM(NH) Typhlodromus (Al1zblyseius) asia/jeus, Narayanan, Kaur & Ghai, Proc. Nat. Ins!. I Sci. India, 26(B) (6) : 389. (Karnataka : arecanut)-identification doubtful Amblyseius asia/ieus, Gupta, Sci. & Cult., 36 : 98. -Misidentifica ti on Amblyse;us asiatieus, Gupta, Internal. J. Acorol., 1(2) : 32. (West Bengal: Helianthus annuus, Anaeardiunz occidentalis)-misidentification Amblyseius (Anzb/yseius) asiaficll.f, Ehara & Bhandhufalck, J. Fae. Ed. To tto ri Univ. Nat. Sci., 27(2) : (Redcscribed). (Thailand & Malay: Cambre/um quudrangu/are, EupatoriulIl odoratul11, Co/ells blumei, soy-been, bamboo, guava, Dioscorl'ea, citrus, Triul1zjella rlzonlboides, tea, Celosj~ argentea, corn). Diagnosis: Dorsal shield imbricate near lateral margins, with 17 pairs of setae. Measurements of setae: jl.20, j4-8, j5-8, js-10, , 1 5-8, r s-13, R l -l1, js-15, za-14, Z4-16, S4-21, ZI-11, Sa-15, S 4-13, Z5-73 z5-9, Z4-51. Ventrianal shield shield-shaped with 3 pairs of setae. Spermatheca with cervix elongate, tubular, gradually sidening towards mouth and often coiled. Fixed digit of chelicera multidentae ; movable digit 3-dentate. Macrosetae on leg IV, genu-27, tibia-22, basitarsus-50. Chaetotaxy same as in fernale. Spermatophoral 'process inverted L-shaped. Indonesia, Malaya, India?, Thailand.

74 60 REC. ZOOL. SURV. INDIA, OCC. PAPER No Amblyseius (Typblodromips) crotalariae Gupta Al1lblyseius crotalariae Gupta, Entoln%gists' mono Mag., 112: (Holotype ~, India: West Bengal, ])arjeeling Dist., Sukna, Crotalaria sp., in :lsi) Amblyseius crotalariae, Gupta, Indian J. Acar., 2(2): (Meghalaya: fern Assam : fern ; Tripura; Nyctanthes arbortristis) Amb(vseius (Typhiodromips) crotaiariae, Gupta, Fauna of India (Acari: Mesos-' tigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield smooth with 17 pairs of setae and 5 pairs of pores. Excepting setae ii' i a, s4' Z4 and ZI)' other setae being very small. Setae measure: ji-18, ia-22, S4-36, Z4-45, Z5-63 (serrate). Sternal shield slightly reticulate; almost as long as broad. Ventrianal shield elongate, lateral margins co~tricted with 3 pairs of setae, Macrosetae on leg IV : genu-46, tibia-36, basitarsus-55. Gupta (1978) figured its spermatophoral process. India (West Bengal, Assam, Tripura) Amhlyseius (Typblodromips) eujeniae Gupta Alnblyseius enjeniae Gupta, Orielltal Ins., 11 : (Holotype ~, India: Andaman IsIs., Ferrargunj, on black berry; Paratype 1 ~, Ferrargunj, on banana, all in ZSI) Amb~vseius (Typhlodromips) eujeniae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, Diagnosis: Dorsal shield smooth, with 17 pairs of setae. Measurements of setae: il-20, j4' j6-5 each, ja-5, ja-35, za-8, Z4-8, S4-52, Z1' Sa S4-8 each, 21)-80, z{;-8, Z4-56. Ventrianal shield squarish with 3 pairs of preanal setae; 2 pairs of metapodal plates present. Macrosetae on leg IV : genu-52, tibia-28, basitarsus-64. India ( Andaman & Nicobar IsIs ) Amblyseius (Typblodromips) gbanii (Muma) Cydllodrol11us ghanii Muma, Fla. E"I., 50(4) : 274. (Holotype ~,Pakistan, Hyderabad, on GossypiU111 sp., in USNM). Diagnosis: Dorsal shield 330 long, 180 wide, slightly sclerotized, with 17 pairs of setae, all being short, except Z4 and Z{; which are longer and the latter serrate. Fixed digit of chelicera with 2 teeth. Distribution; Pakistan.

75 GUPTA: ORIENTAL PHYTOSEliDAE " AmhJyseius (Typhlodromips) guajavae Gupta Anlblyseius guajavae Gupta, Indian J. Acar., 2(2) : 63. (Holotype ~,Paratypes 5 ~ ~, India: Meghalaya, Dalu, on guava, in ZSl) Amblyseius (Typhlodromips) guajavae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield with 17 pairs of setae. Measurements of f. setae: jl-32, js-36, 84-64, Z5-BO, Z4-60; other setae measure between 6 8 long. Sternal, metasternal and genital shields normal. Ventrianal shield pentagonal, much longer than broad, with 3 pairs of preanal setae. Spermatheca with broad tubular cervix and short atrium. Leg IV with macrosetae : genu-40, tibia-30, basitarsus-76. Distribution; India ( Meghalaya ). 117 Amblyseius (Typhlodromips) hidakai Ehara & Bhandhufalack Amhlyseius (Amb/yseius) hidakai Ehara & Bhandhufalck, J. Fac. Ed. Tollor; Univ. Nat. Sci., 27(2) : (Holotype ~, Paratypes 4 ~ ~, 10, Thailand: Fang, on an undet. tree, in BioI. Inst. Fac. Ed. Tott. Univ.). Diagnosis: Dorsal shield reticulate along anterior margin, with 17 pairs of setae and 5 pairs of pores ; j3) S4' Z5' Z4 stout, other setae small. Measurements of setae: j1-24, j4-8, i5-7, j6-10, J 2-12, J 5-8, rs-14, R l -9, ja-38, za-11, z4-11, s4-65, Z1-10, Sa-12, S4-8, S5-8, Z5-84, zes-8, Z4-63. Ventrianal shield longer than wide, with 3 pairs of preanal setae and a pair of pores. Fixed digit of chelicera multidentate. Macrosetae on leg IV : genu-56, tibia-39, basitarsus-82. Chaetotaxy similar as in female AmbJyseius ( Typhlodromips ) johoreae ( Muma ) Typhlodronzips johoreae Muma, Fla. Ellt., 50 : 272. (Holotype ~, Malaya: Johore, on oil palnl feeding on red spider, in USNM). Diagnosis: Dorsal shield 320 long, 200 wide, lightly sclerotized, imbricate on anterior fourth fifth, with 17 pairs of setae; all the setae small except 24 and Z5 which are thick and serrate. Fixed digit of chelicera with 7-9 teeth, movable digit with 2-3 teeth. Cervix of spermatheca a short tubular structure. Chaetotaxy of genu II 23 -& 1, genu III Malaya.

76 62 H.EC. ZOOL. SURV. INDIA, Oce. PAPER No Amblyseius Cfyphlodromips) lenis Corpuz & Rin1ando A111blyseills lei/is Corpuz & Rimando, Philipp. Agl'ic., 50: (Holotype ~, Philippines: College calupus, Los Banos, Laguna; on Citrus Ilohilis ; Paratypes 2 ~ ~, san1e locality as for ho!otype, on Digitaria sanguillulis ; 1 ~, same locality, Elusille Indica; 1 ~, Pel1ll;setul11 sp. ; all in Dept. Ento, Univ. Philip.). Diagnosis: Dorsal shield smooth, 363 long, 232 wide, with 17 pairs of setae ; j l' j3' S4' 2 4, 25 moderately long measuring 22, 29, 58, 80 and 75, respectively; others being minute; r 3, Rl on interscutal membrane. Matasternal plates rectangular. Ventrianal shield pentagonal, reticulate, with 3 pairs of setae and a pair of pores. Peritreme extends anteriorly upto j3. Fixed digit of chelicera ll1ultidentate, movable digit unidentate. Spermatheca with cervix a narrow arc, atrium slender. Macrosetae on leg IV : genu-46, tibia-24, basitarsus-55. Philippines Amblyseius (Typhlodromips) labis Corpuz & Rimando Al1lblyseius labis Corpuz & Rimando, Philipp. Agric., 50: (Holotype ~, Para types 5 ~ ~, 4 d' d', Philippines, Gamu, Isabela, on Tabernaemolltana pandacaque. Other Paratypes: 3 ~ ~, Villaverde, Nueva Vizcaya, Prel1l1la odorala; 1 ~, College Canlpus, Laguna, Los Banos, on Ceiba pelltandra, all in Dept. Ent. Univ. Philip. Diagnosis: Dorsal shield 328 long, 220 wide, with linear anterior and lateral reticulations, with 17 pairs of setae and 7 pairs of crescentic pores. All setae rather short except Z4 (20) and Z5 (78), both thick and the latter serrate. Metasternal setae present on metasternal plates. Peritreme extends beyond jl. Fixed digit of chelicera with 10 teeth, movable digit with 3 teeth. Leg IV with 3 macrosetae, all knobbed. Spermatheca with very short and wide cervix, atrium swollen. Similar to female in chaetotaxy ; spermatophoral process inverted L-shaped. Disli ibution: Philippines Amblyseius (Typhlodromips) Iinearis Corpuz & Rimando AI11bly.,eius linearis Corpuz & Riolando, Philipp. Agric., 50: (Holotype ~, Philippines: Gaolu, Isabela, on Achyrallllzes aspera ; Paratypes 4 S,-?, 3 00, san1e data as for holotype; 6 exs (sex not specified) Los Banos; 1 ~, on Achyral11iJes (J~pt.'ra ; 4 ~ ~, on nlulberry, all jn Dept. Ento. Univ. Philip.).

77 GUPTA: ORIENTAL PHYTOSEllDAE 63 Diagnosis: Dorsal shield 324 long, 214 wide, margins with linear reticulations, with 17 pairs of setae; all setae short except Z4 and 2 5, which are long, thick and serrate. Metasternal plates triangular. Peritreme extends upto i 1 Ventrianal shield pentagonal with 3 pairs of preanal setae. Leg IV with macro setae on genu-24, tibia-20, and basitarsus-46. Cervix of spermatheca slender, cylindrical, gradually wtdening like a funnel; atrium cylindrical. Fixed digit of chelicera multidentate with pilus dentilis, movable digit tridentate. Similar to female in chaetotaxy. Spermatophoral process inverted L-shaped. Philippines Amblyseius (Typhlodromips) mangiferae Ghai & Menon Amblyseius magiferae Ghai & Menon, Oriental. Ins., 1 : (Holotype C, Paratype ~, and 2 more specimens (sex not specified), India: Delhi, on malformed mango infested with eriophyids, in N PC) Amblyseius magijerae, Gupta, Internat J. Acarol., 1 (2): Amblyseius (Typh/odromips) mangijerae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield smooth, with 1 7 pairs of setae, Z 5 being longest and thickest; ji' ;a, Za, Z4' S4' is, J a quite long, rest small. Setae ja-z4 being shorter than the distance between their bases. Ventrianal shield with 3 pairs of preanal setae and a pair of circular pores. Leg IV with macrosetae on genu, tibia and basitarsus. India ( Delhi) Amblyseius ( Typhlodromips ) meghalayensis Gupta Ambiyseius meghaiayensis Gupta, Indian J. Acar., 2(2) : (Holotype ~, Paratypes 4 ~ ~, Megbalaya: Dalu, guava, in ZSI; additional paratypes 1 ~,Assam, Dhubri, undet plant, in ZSI) Amblyseius (Typhiodromips) meghalayensis, Gupta, 1 auna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 300 long, 162 wide, reticulate with 1 7 pairs of setae. Measurements of setae: i l -20, i4 -if)-8 each, js, J a-10 each, 1 5-6, ja-13, za-10, z4-16, s4-24, Zl-12, Sa-17, S4-S5-16 each, Z5-64, Z5-8, Z4-41; all setae being smooth and pointed. Ventrianalshield 88 long, 70 wide, with 3 pairs of preanal setae and a pair of pores. Spermatheca with tubular cervix. Macrosetae on leg IV : genu-24, tibia-20, basitarsus-48.

78 64 REC. ZOOL. SURVE. INDIA, OCC. PAPER No. 95 Gupta (1978) figured its spermatophoral process. India: Meghalaya Amblyseius (Typhlodromips) neocrotalariae (Gupta) Typhlodromips neocrotalariae Gupta, Oriental Ins., 12 : 335. (Holotype ~ Paratypes 2 ~ ~, 10, India: TamilNadu, Kodai Kanal, Oothu, on Datura sp., in ZSI) Amblyseius (Typhlodromips) neocrotaiariae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 324 long, 212 wide, reticulate, with 17 pairs of setae measuring: j1-20, j4 -j5-8 each, j6' J a-12 each, J 5-8, js-28, za, z4-9 each, s4-30, ZI' Sa-12 each, S4-S5-8 each, Zf)-68 (weakly serrate), z5-8, Z4-40. Ventrianal shield 108 long, 80 wide, with 3 pairs of preanal setae. Macrosetae on leg IV : genu-52, tibia-40, basitarsus-60. India ( Tamil Nadu ) Amblyseius ( Typhlodromips ) okinawanus Ehara Amblyseius (Amblyseius) okinawanus Ebara, Mushi, 40(6) : (Holotype ~, Paratypes 6 ~ ~, 200, Japan: Okinawa lsi., Tomigu8uku, on Verbena officinalis. Additional Paratypes 1 ~, 10, Tomigusuku, on Plantago asiatica; 1 ~, 10', same locality, Brachlaria mir;ca; 1?, Nakijin, on Gerbera sp., all in Zool. Inst. Fac. Sci., Hokk. Univ.) Amblyseius okillawanus, Lo, Bull. Sun- Yat. Sen Cult. Found., 5 : 56 (Taiwan: on citrus witb P. ulmi) Amblyseius okinawalllls, Ebara & Bhandhufalck, J. Fac. Ed. Tottori Un;v. Nat. Sci., 27(2): 58. (ThaiJand : Ageratuln conyzoides, Polygonum chinens;s, corn, citrus). Diagnosis: Dorsal shield reticulate near lateral margins with 17 pairs of setae and 6 pairs of pores. Setae Z5 and Z4 stout and serrate. Measurements of setae: jl-18, j4-10, j5-9, je-11, J s-12, J f)-8, rs-12, R 1-11 js-19, za-14, Z4-15, S4-19, ZI-12, S2-17, S4-17, Sf)-15, Z5-57, z5-10, Z4-30. Ventrianal plate shield-shaped, longer than wide, with 3 pairs of preanal setae ; metasternal plate with seta present. Macrosetae on leg IV, genu-29, tibia-24, basitarsus-54. Spermatheca with funnel-shaped cervix. Spermatophoral process hammer-shaped. Japan, Taiwan, Thailand.

79 GUPTA: ORIENTAL PHYTOSElIDAE Amhlyseius (Typhlodromips) officinaria Gupta Amblyseills ojficinaria Gupta, Indian J. Acar., 1(2) : 39. (Holotype ~, Paratypes 4 ~ ~, 1 d, India: West Bengal, Chinsurah, on Saccharunl ojficinarull1, in ZSI) Amb/yseills (Typhlodrolnips) ofjicinaria, Gupta, Fauna of India (Acari: Mesostigmata) Fan1i1y Phytoseiidae, p Diagnosis: Dorsal shield 302 long, 200 wide, margins slightly imbricate with 17 pairs of setae and 4 pairs of pores. Measurements of setae: jl-18, j4,-4, j5-6 each, j6~10, J a -12, J 5-7, j3' ZS) z4-9 each, , Z'1'-9, Sa-14, S4-11, S5-10, Z5-82, Z4-30. Sternal, metasternal and genital shields normal. Ventrianal shield 104 long, 72 wide, with 3 pairs of preanal setae. Spermatheca with cup-shaped cervix. Macrosetae on leg IV : genu-23, tibia-27, basitarsus-46. Gupta (1975) figured the spermatophoral process. India (West Bengal) Amblyseius (TyphIodromips) perfectus Chaudhri Ambiyseius perfectus Chaudhri, Acar%gia, 10: (Holotype ~, Pakistan : 4 miles North of Sheikhupura, on loquat, in UAL ; Paratype 1 ~, 6 miles South of Sheikhupura, on paddy, repository not mentioned). Diagnosis: Dorsal shield 333 long, 186 wide, with 17 pairs of setae. i 1-13, i 4, i5-10 each, j6-14, l a -15, 1 5-8, j3-23, za, z4-18 each, s4-26, Z1-17, Sa-20, S4-S5-14 each, Z5-58 (weakly serrate), z5-12, Z4-39, (weakly serrate), r 3-18, R l -16. Ventrianal shield rectangular, 117 long, 88 wide, with 3 pairs of setae and a pair of elliptical pores. Two pairs of metapodal plates present. Spermatheca with long funnel-shaped cervix. Chelicera with 5 subapical teeth on fixed digit and 1 subapical tooth on movable digit. L,eg IV with macrosetae on genu-41, tibia-17, basitarsus-61. Pakistan Amblyseius ( Typhlodromips ) polyantbeae Gupta Amblyseius polyantheae Gupta, Internal. J. AcaJ'ol., 1(2) : (Holotype ~, India: West Bengal, on Po/yanthea tuberosa, in ZSI) Amb(vseills (Typhlodrondps) poi),ol1theae, Gupta, Fauna of India (Acari: Mesostigmata) r-amily Phytoseiidae p Diagnosis: 9 Dorsal shield 325 long, 250 wide, with 17 pairs of setae

80 66 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 Measurements of setae: jl-18, j4-j6' Jat J each, ja-14, za-12, Z4,-S, 84-27, Z1-18, Sa-14, S4-14, 5 5-9, Zrs-57, z5-6, Z4-27. Ventrianal shield longer (104) than broad (81), with 3 pairs of preanal setae. Macrosetae on leg IV : genu.. 27, tibia-31, basitarsus-36. India (West Bengal) Amblyseius (Typhlodromips) potentillae ( Garman) Amblyseius potentillae Garman, Ann. en!. Soc. Am., 51 : (Holotype ~,Holland, on Potentilla sp., in USNM) Amblyseius rykei Pritchard and Baker, Gupta, Sci. & Cult., 36: 98 (West Bengal: Ficus) Amblyseius charui Gupta, Bull. En!., En!. Soc. India, 10 : (West Bengal: Magno/es chanlpa; Ca/ophy//um inophyllum) Amblyseius charui, Gupta, Internat J. Acarol, 1(2): 33 (West Bengal: Punica granatum). 198\. Alnblyseius potentil/ae, Gupta, Indian J. Acar., S : 47 (Re-described). (Jammu & Kashmir: on cucurbitaceous plant) Amblyseius (Typhlodromips) potentillae, Gupta} Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 300 long, 172 wide, with 17 pairs of setae and 3 pairs of pores. Setae j1' js' S4' Z.. and Z5 being long, measuring 20, 44, 72, 60 and 84, respectively. Ventrianal shield 100 long, 84 wide, anterior margin rounded, lateral margins gently concave with 3 pairs of preanal setae and a pair of pores. Fixed digit of chelicera multidentate, movable digit 3-dentate. Spermatheca with bell-shaped cervix. lviacrosetae on leg IV : genu-40, tibia-40, basitarsus-60.? Europe, USA, India ( Jammu & Kashmir ) Amblyseius (Typhlodromips) sapienticola Gupta Amblyseius sapient;cola Gupta, Orienta/Ins., 11 : (Holotype ~,Paratypes 4 ~ ~, India: Andaman IsIs., Ferrargunj, on banana, in ZSI) Amblyseius (Typhlodromips) sapient;eo/a, Gupta, Fauna of India (Acari: Mesos-. tigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 308 long, 188 wide, reticulate. Measure-: ments of setae; j1-16, j4:-j6' J 1 -J 5-8 each, js-12, Za, Z4-8 each, s4t-16t.~ Zl' S4z-S5-8 each, Sa-12, 25-64, z1)-8, Ventrianal shield 100 long, 108 wide, lateral margins notched, with 3 pairs,of preanal setae. Macrosetae on leg IV: genu-48, tibia-33, basitarsus-48.

81 GUPTA: ORIENTAL PHYTOSElIDAE 67 India (Andaman IsIs) Amblyseius (Typhlodromips) similis (Koch) Zercon similis Koch, Fasc. 27. (Holotype ~, Germany, lost) Typhlodromus (Amh/yseius) simi/is, Chant, Can. EIlI., 91 (Suppl. 12) : Diagnosis: Dorsal shield with 17 pairs of setae; js, 54' Z5' Z4 long; Z4' Z2' ZI' S4 and S5 minute but equal to one another; S2>Zl' Ventrianal shield long, narrow, lateral margins parallel, with 3 pairs of preanal setae and a pair of pores, Leg IV with macrosetae on genu, tibia and basitarsus. Germany, Italy, France, Belgium, Austria, Indonesia, Holland, Mexico, California, Texas. Remarks: Koch's holotype is lost Amblyseius (Typhlodromips) suknaensis Gupta ~mblyseius suknaensis Gupta, Orienta/Ins., 4: (Holotype ~, Paratypes 3 ~ ~, 26' C, India: West Bengal, Sukna, on Colocasia sp., in ZSI) Amblyseius suknaensis, Gupta, Internat. J. Acarol., 1(2) : Amblyseius daturae Gupta, Internat. J. Acaro/., 1(2): (W. Bengal Datura) Amhlyseius suknaellsis, Gupta, Oriental Ins., 11: 636 (Andaman lsi. : Tee/olla gralldis, Rangat lsi. : Undet. plant, Car Nicobar : Undet. plant) Neoseiulus suknaensis, Gupta, Oriental Ins., 12: 337 (Kerala : rose) Alnblyseius suknaensis, Gupta, Indian J. Acar., 2(2): 73. (Or issa: SIzorea robusta ; Meghalaya : Undet. plant. ; Assam: fern; Tripura : Undet. plant) Amblyseius (Typhlodromips) suknaellsis, Gupta, Fauna of India (Acari: Mesostigmata) Fanli1y Phytoseiidae, p Diagnosis: Dorsal shield 287 long, 175 wide, with 17 pairs of setae and 3 pairs of pores. Measurements of setae: jl-18, j4 -js, J g-j each, ja-13, za-9, z4-11, s4-13, ZI' S2-S5-9 each, Z5-67, Z5-6, Z4-4'. Sternal, metasternal and genital shields normal. Ventrianal shield pentagonal, 90 long, 85 wide, with 3 pairs of preanal setae and a pair of pores. Macrosetae on leg IV : genu-27, tibia-18, basitarsus-48. Gupta (1970) figured its spermatophoral process and ventrianal shield. India (West Bengal, Andaman & Nicobar IsIs., Kerala, Orissa, Meghalaya, Assam, Tripura).

82 68 REC. ZOOL. SURV. INDIA, OCC. PAPER No Amblyseius (Typhlodromips) syzygii Gupta Anlblyseius syzygii Gupta, Internal. J. Acarol., 1(2) : 4(-45. (Holotype ~,India: West Bengal, on Syzygiunl cul1lini, in ZSI) Amblyseius syzygii, Ebara & Bhandhufalck, J. Fac. Ed. Tollori Ullil'. Nat. Sci., 27(2) : (Thailand: citrus, guava, Cratoxyium, litchi, bamboo, sugarcane, soybean, papaya, Eupatorium odoratum, Triumfetta rhomboides, longan, corn, Hibiscus sp.) Amblyseius syzygi;, Gupta, Indian J. Acar., 2(2) : 74. (Orissa: leaf litter ; Tripura: Undet. plant) Amblyseius syzygii, Gupta & Nahar, lit Contrtb. to Acar. in India, p. 8. (Bihar: nlaize, beans, sugarcane) Amblyseius (Typhlod1omips) syzygii, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytos~iidae, p Diagnosis: Dorsal shield reticulate, 345 long, 210 wide. Measurements of setae: j1-15, j4-j6' la-lis 7-10 each, ja-16, Za-z4-9 each, S4-20, Z1' Sa-S5-10 each, Z5-67 (serrate), Z4-31 (serrate), zls-9. Sternal, metasternal and genital shields normal. Ventrianal shield reticulate, llq long, 85 wide, with 3 pairs of preanal setae and a pair of pores. Macrosetae on leg IV : genu-45, tibia-3!, basitarsus-49. Ehara & Bhandhuflack (1977) described male. India (West Bengal, Orissa, Tripura), Thailand, Indonesia Amblyseius (Typhlodromips) tetranychivorus (Gupta) Typhlodrol1zips tetrallychivorlls Gupta, Oriental IllS., 12: (Holotype ~,India: Tamil Nadu : Coimbatore, Agril. Univ. Campus, on brinjal, infested with T. cillllabarillus; Paratypes 1 ~, 1 d, Kerala, Trivandrum, Nayer Dan, on undet. plant, all in ZSI) Typhlodromips letrallychivorus, Channa Ba~avanna, Final Rept. ICAR Research Schen1e on Studies on the spidermite family Tetranychidae ( ). Univ. Agril. Sci. Bangalore, p Alnblyseills telranychivorlls, Gupta & Nahar, Ill. Contrib. to Acar. in India, p-9. (Bihar: Hibiscus 111Iltabilis) Anlblyseills (Typhlodrolnips) tetrallychivol'us, Gupta, Fauna of India (Acari; Mesostigmata) Fan1i1y Phytoseiidae, p Diagnosis: Dorsal shield 320 long, 192 wide, anterior lateral margins imbricate, with 17 pairs of setae measuring j1-28, j4-j6' J a -J each, is-44, za-17, z4-36, , Z1",,8, Sa-36, S4-8, S5-B, ZIS-80, z5-8, Z4-66. Sternal shield with postlateral projections. Ventrianal shield longer (100) than broad, anterior margin rounded, lateral margins straight with 3 pairs of preanal setae. Macrosetae on leg IV : genu-48, tibia-40 and basitarsus-68. Spermatheca with cup-shaped cervix.

83 GUPTA: ORIENTAL PHYTOSEIlDAE 69 Gupta (1978) figured its spermatophoral process. India (Tamil Nadu, Kerala, Karnataka, Bihar) Amblyseius (Typhlodromips) vignae Lai-rong & Li-sheng Amblyseius vignae Lai-rong & Li-sheng, Fudan J. (Nat. Sci.), 20(2) : (Holotype ~, Fujien Long, on Vigna sinensis, repository unknown). Diagnosis: Dorsal shield 350 long, 250 wide, with 17 pairs of setae. Measurements of setae: j1-19, j4-8, j5-4, j6-6, J a -ll, J 5-6, z5-8, Z4-47, j s -13, Z a -13, Z4-13, s 4-19, Z 1-8, Sa -1 7, S 4-1 7, S 5-17, Z 5-81, r a -11, R 1-11, JV Macrosetae on leg IV : genu-26, tibia-24, basitarsus-59. South China. In addition to the above, the under mentioned species are also known but could not be included in key for reasons as mentioned for A.(A.) sp. on p Amblyseius (Typhlodromips) baiyunensis Wei-nan Amhlyseius (Amhlyseius) baiyunensis Wei-nan, Acarology, 6(1) : 224. (Holotype ~, Paratype 1 ~, South China: Baiyun Mountains, Guangzhou, on 1). Diagnosis: Idiosoma long, wide. Measurements of setae: j1-32, za-18, z4-18, s4-35, Z1-15, S2-22, S4-30, S5-31, Z5-67, ze-10, Z4-18, ra-17, R 1-15, ]V 5-37 South China Amblyseius (Typhlodromips) draconis (Chaudhri, Akbar & Rasool) Typhlodrol1zips dracol1is Chaudhri, Akbar & Rasool, Rcpt. PL 480 Project, Univ. Agril. Faisalabad, p. 74. (Holotype ~, Pakistan: Mansehra, on [Ililia cuspidafa, in UAF). Diagnosis: Dorsal shield 364 long, 182 wide. Measuren1ents of setae: j1-29, j4:-21, j5-26, j6-36, J 2-49, J 5-13, ja-39, za-39, z4-44, s4-52, Zl-57, Sa-57, S4-42, Z5-60, z5-26, Z4-62, r a -39, R Spermatheca poculiform. Macrosetae on leg IV : genu-34, tibia-36, basitarsus-68. Pakistan.

84 70 REe. ZOOL. SURV. INDIA, OCc. PAPER No Amblyseius (Typblodromips) neogbanii Gupta Al1lhlyseius (Typhlodronlips) neoghanii Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p (Holotype ~, India: Ar unachal Pradesh, Ziro, Yazuli, on undet. plant, in ZSI). Diagnosis: Dorsal shield 358 long, 250 wide. Measurements of setae: jl-18, j4-9, h~-10, j6-12, 19-14, 15-10, ja-11, za-7, z4-7, s4-16, ZI-11, 8a-l1, , 8 5-9, Z~-95, z5-9, Z4-22, f S ' R 1 9 each. Ventrianal shield 134 long, 118 wide, with 3 pairs of preanal setae. Macrosetae on leg IV: genu-3!, tibia-40, basitarsus-92. India (Arunachal Pradesh) Amblyseius (Typblodromips) oguroi Ehara Amb/yseius oguro; Ebara, J. Fac. Sci. Hokkaido Uldv. (6) 15 (3) : (Holotype ~,Paratype 16, Japan: Sendai, on Mallolotus japonicus, in Zool. Inst. Fac. Sci. Hokk. Univ.) Amhlyseius (Amh/yseius) oguro;, Wei-nan, Acarology 6(1) : 222. Diagnosis: Dorsal shield reticulate along lateral margins. Measurements of setae: i l -23, 'j4-9, j~-9, i 6-9, l a -12,15-7, js-22, za-13, z4:-17, s..-24, ZI-12, ,84, -14, S~-13, Z~-87, z5-10, Z4-60, rs-14, RI-10, JV Macrosetae on leg IV: genu-37, tibia-32, basitarsu~-55. Dorsal setae shorter than those of males. Ehara (1964) described male in detail. Japan, South China Amblyseius (Typhlodromips) repletus Wei-nan & Zhao-Quan Alnblyseius (Amhlyscius) rep/elus ~ei-nan & Zhao-Quan, Acto Zoolaxonamica SinicQ, 10(3) : (Ho!otype ~, Paratypes 8 ~ ~ South China: SheD Nong Zia,' Hubei Province, on Rhus sp.) Vilis adslricta & Prunus salicina). Diagnosis: Dorsal shield long, 215~225 wide. Measure-' ments of setae: i l -17, j4-13, j5-12, je-12-15, Ja-17-18, J , js-17, za-13, z4-16, s4:-17-21, ZI-15-16, Sa-18-20, S , S , Z , z5-12, Z , fs-ls-20, RI-13-16, )V 5-22.? South China.

85 GUPTA: ORIENTAL PHYTOSEllDAE Amblyseius (Typhlodromips) sijiensis Gupta Amblyseius (Typhlodromips) sijiensis Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p (Holotype ~, India ~ Arunachal Pradesh, Siji, on undet. plant, in ZSI; Paratypes 1 ~, 1 C, Daporizo, on undet. plant, in ZSI). Diagnosis: Dorsal shield long, wide, recticulate. Measurements of setae: jl-22, j4-j6-7-9 each, la-ii, IG-6, ja-18-21, za-11-12, z , , Z1-9-12, S2-16, S , , Z , z5-8, Z4-56, rs, R 1-16 each. Macrosetae on leg IV: genu-24-30, tibia-20-26, basitarsus Gupta (1986) described male. India (Arunachal Pradesh). Genus Anathia Chaudhri, Akbar & Rasool Allathia Chaudhri, Akbar and Rasool, Rept. PL 480 Project, UAF, p. 48. Type: Anathia rancidus Chaudhri, Akbar & Rasool, Anatbia rancidus Chaudhri, Akbar & Rasool Allathia rancidus Chaudhri, Akbar & Rasool, Rept. PL 480 Project. UAF, p.48. (Holotype ~,Pakistan : Lahore, on undet. plant, in UAF). Diagnosis: Dorsal shield reticulate, 489 long, 265 wide. Measurements of setae: j1-39, ja-18, j4-16, j5-22, i 6 - I 8, , J 5-13, j3-44, z2-21, z4-23, s4-31, Z].-26, S2-28, S4-23, 8 G -21, 26-83, z5-18, Z4-5 Ventrianal shield pentagonal, 161 long, 133 wiqe, with 3 pairs of preanal setae. Macroseta on basitarsus IV-31 long. Pakistan. Genus Indoseiulus Ehara lndoseius Ghai & Menon, Oriental /I1S., 3 : Indoseiulus Ehara, Appl. En!. Zool., 17 : Type: Indoseius ricini Ghai & Menon, Indoseiulus ricini (Ghai & Menon) Indoseius ricini Ghai & Menon, Oriental Ins., 3 : 348. (Holotype ~, Paratypes 8 ~ ~, India: Coimbatore, on Ricinus COllll11Ullis, in NPC).

86 72 REC. ZOOL. SURV. INDIA, OCC. PAPER No /Ildoseills ricilli, Gupta & Dhooria, Abs~r. India" Sci. COllgr., 1974, p. 69. (Punjab and Himachal Pradesh) lndoseius ricin;, Gupta, Indian J. Acar., 1 : 18. (Gujarat : Castor; Uttar Pradesh: Castor) Illdoseius ricin;, Gupta & Nahar, In Contrib. to Acar. in India, p. 10. (Bihar: guava), Indoseililus ricini, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield oval, 311 long, 208 wide, does not cover whole of dorsum, reticulate with 16 pairs of setae measuring jl-21, j4-8, j5-10, j6-10, la-12, J 5 /8, js-io, za-10, z4-16, s4-16, Z1-16, S2-16, S 5-14, Z5-24" z5-12, Z4-12. Sternal, metasternal and genital shields normal. Ventrianal shield absent, only preanal setae present. Macrosetae on leg IV : genu-32, tibia-40" basitarsus-36. India (Tamil Nadu, Gujarat, Bihar, Uttar Pradesh) Indoseiulus eharai Gupta Indoseiullls eharai Gupta, Orienta/Ins. (in press). (Holotype ~ Paratype 1 ~ India: Arunachal Pradesh, Khonsa, on papaya in ZSI) Indoseiulus eharai, Gupta, Fauna of India (Acari : Mesoltigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 425 long, 250 wide, poorly sclerotized and highly reticulate, with 16 pairs of setae and 2-3 pairs of pores. Measurements of setae: jl-26, j4-9, j5.7, j6-11, la-ii, 1 5-4, ja-7, z.l-lo, z4-9, s4-13, , S2-13, S5-14, Z5-22, Z5-4, Z4-15, rs-14, Rl -12. Ventrianal shield appears to be absent, 3 pairs of preanal setae present. Macrosetae on leg IV : genu-71, tibia-67, basitarsus-56, that on genu , tibia , genu ; genu 1 with 3 macrosetae, 44, 33, 38 long, respectively. India (Arunachal Pradesh). Genus Iphiseius Berlese Iphise;us Berlese, Redia, 12 : Phytoseiulel/a Muma, Bull. Fla. St. Mus., 5(1) : Iphiseiini Schuster & Pritchard, Hilgardia, 34: Type: Sejus degenerans Berlese, 1916.

87 GUPTA: ORIENTAL PHYTOSElIDAE 73 Key to the species of Iphiseius 1. All setae but Z4 short Z 4' Z4" s, long or very long andamanicus bakeri 145. Iphiseius (Iphiseius) andainanicus Gupta Iphiseius andamanicus Gupta, Entomologists' mon. Mag., 115 : (Holotype ~, Paratypes 9 ~ ~, India: Andaman Isis., on Dolichos lablab, in ZSI) lphiseius (/phiseius) andamanicus, Gupta, Fauna of India (Acari: Mesostigmat a) Family Phytoseiidae, p Diagnosis: Dorsal shield oval, 320 long, 243 wide, smooth with 17 pairs of setae; all being short (5-8 long) except Z5 which is 27 long. Sternal shield as broad as long ; meta sternal plates big. Ventrianal shield pentagonal, 99 long, 68 wide, constricted at anal region; small platelets present between genital and ventrianal shields, with 3 pairs of preanal setae. Macrosetae on leg IV : genu-11, tibia-14, basitarsus-50. India (Andaman IsIs.) Iphiseius (Iphiseius) hapoli Gupta Jphiseius (Iphiseius) hapoli Gupta, Oriental Ins. (Holotype ~, India: Arunachal Pradesh, Hapoli, on apple, in ZSI) Iphiseius (Iphiseius) hapoli, Gupta, Fauna of India (Acari: Mesostigmata) Family"Phytoseiidae, p Diagnosis: Dorsal shield highly sclerotized with 17 pairs of setae, all being very small except Z5' js, j1 which are slightly longer and measure 12, 18, 14, respectively; others measure 6-8 long. Ventrianal shield reticulate, 100 long, 112 wide with 3 pairs of preanal setae. No macroseta present on leg IV. Gupta (1986) described it. India (Arunachal Pradesh) Iphiseius (Trochoseius) bakeri Gupta Iphiseius bakeri Gupta, Entomologists' mon. Mag., 115 : (Holotype 0, India: Andaman lsi., on arecanut, in ZSI) Iphiseius (Trochoseius) bakeri, Gupta, Fauna of India (Acari: Mesostigmata) F amity Phytoseiidae, p Diagnosis (Known from 0 only): Dorsal shield 288 long, 255 wide, 10

88 74 REC. ZOOL. SURV. INDlA, OCC. PAPBR No.. 95 highly sclerotized, reticulate, with 15 pairs of setae 05' J a lacking), all setae small (5-10 long), excepting S4' Z5 and Z4 which measure 117" 189 and 135, respectively. Ventrianal shield reticulate with 3 pairs of preanal setae and a pair of pores. Macrosetae on leg IV : genu~54, tibia-45, basitarsus-36. Female: India (Andaman IsIs.). Genus Paraamblyseius Murna Paraamblyseius Muma, Fla. Ent., 45 : 8. Type: Paraamblyseius lunatus Muma, Key to the species of Pllraamblyseius 1. Metasternal plate round, J 2 longer than Z 1 Metasternal plate kidney-shaped, J ~ equal to Zl mumai fragariae 148. Paraamblyseius fragariae Gupta Paraamblyseius SPa Gupta, Sci. & Cult., 36 : 98. (West Bengal: strawberry) Paraamhlyseius fragariae Gupta, Entomologists' mona Mag., 115 : (Holotype ~,India : West Bengal, Fragaria nilgerrensis, in ZSI) Paraamblyseius formosanus Gupta, Entomologists' mona Mag., 115: 35. (Assam: Undet plant; Manipur: Undet. plant) Paraamblyseiusfragariae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield highly sclerotized, reticulate, 320 long, 230 wide, with 17 pairs of setae measuring: jl-12, j4-14, j5-16, j6-33, J a -40, J 5-12, is-is, za-22, z4-22, s4-33, Z1-28, Sa-36, S4-29, , Zrs-38, Z4-45. Sternal shield reticulate, wider than long. Ventrianal shield massive, sculptured, with 3 pairs of preanal setae and a pair of pores. Macroseta absent on leg IV. India (West Bengal) Paraamblyseius formosanus (Ehara) Jphiseius formosanus Ehara, Mushi, 43 : (Holotype ~, Paratypes 20' 0' J 11 ~ ~, Taiwan, Kenting, on Calamus magarilae, in BioI. Inst. Fac. Ed. Tott. Univ.). J976! Iphiseills formosqnus, Tsen~, I. Agr. Ass. China, 94:

89 GUPTA: ORIENTAL PHYTOSEilDAE 75 Diagnosis: Idiosoma heavily sclerotized, dorsal shield with rninute pores, margins with reticulations. Measurements of setae: ii-ii, j4,-13, j5-14, ja-22, J 2-27, J 5-14, j3-14, z2-20, z4-23, s4-26, ZI-31, S2-27, 54,-17, S5-15, Z5-29, z5-11, Z4-60. Macroseta on basitarsus IV-1410ng. Ehara (1970) described it in detail. Taiwan, South China Paraamblyseius mumai Gupta Paraamblyseius mumai Gupta, Entomologists' mono Mag., 116: (Holotype ~,India: Meghalaya, Mawphlong, on undet. plant, in ZSI) Paraamblyseius mumai, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield heavily sclerotized.. punctate, 325 long, 232 wide, with 17 pairs of setae. Measurements of setae: jl-16, j4-16, jls-16, ja-40, J ~-48, J 5-14, j3' z2-16, Z4-20, S4-24, ZI-38, Ss-38, , S5-17, Z5-32, z5-36, Z4-40. Sternal shield much wider than long. Ventrianal shield 110 long, 168 wide, with 3 pairs of preanal setae and a pair of pores. Metapodal plates large, triangular. Macroseta absent on leg IV. India (Meghalaya). Genus Platyseiella Muma Platyseiella Murna, Bull. Fla. St. Mus., 5(7) : 280. Type: Platyseiella platypilis Chant, Platyseiella mumai Ray & Gupta Platyseiella mumai Ray & Gupta, Bull. zool. Surv. India., 4(3) : (Holotype ~, India: Tripura, Agartala, Amtali, on undet. plant, in ZSI) Platyseiella mumai, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield rugose, 285 long, 128 wide, with 14 pairs of setae. Setae j4' j5) j6' J 5 and Z5 small and simple, the remaining. setae long, thick and serrate. Measuremen ts of setae : j 1-25, j4-7, j ~.. 5, j8-5, J 5-5, j3-34, z2-11, z4-9, ,86-61, Z5-90, z&-s, Z4-90. Ventrianal shield 100 long, 67 wide, with 3 pairs of preanal setae. Fixed digit of chelicera multidentate. Macrosetae on leg IV : genu-38, tibia-61, basitarsus-34, distitarsus-33, all being spatulate.

90 76 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 India (Tripura) Platyseiella formosanus Tseng Platyseiella (N.) formosanus Tseng, J. Agr. Ass. China,94: Distribution : Taiwan. Remarks: Due to nonavailability of original literature, the particulars about diagnosis, type data, etc. could not be given. This appears to be a doubtful species. Genus Okiseius Ehara Okiseius Ehara, Mushi, 40: 77. Type: Okiseius subtropicus Ehara, Key to the species of Okiseius 1. Incision present on dorsal shield posterior to Rl Incision absent on dorsal shield posterior to Rl 2. State j4, and j6 very small (6-12 microns), simple Setae j4, and j6 longer than above, serrate 3. Ventrianal shield much more than 1 t times as IODg as wide, lateral mareins distinctly constricted Ventrianal shield less than 1~ times as long as wide, lateral margins almost straight, or only slightly constricted subtropicus 2 3 himalayana )1azuliensis sikkimensis 153. Okiseius subtropicus Ehara Okiseius subtropicus Ehara, Mushi, 40 : (Holotype ~,Okinawa Island, Itoman, on Hibiscus tiliaceus, in Zool. Inst. Fac. Sci. Hokk. Univ.) Platyseiel/a (Neoledius) subtropicus, Tseng, J. Agr. Ass. China, N. S.,94: Okiseius subtropicus, Ehara, Rev. Pl. Prot. Res., 10 : 38. Diagnosis: Dorsal shield heavily sclerotized, with 15 pairs of setae, J a absent ; all (except J 5) being stout, serrate, and set on tubercles. Measurements of setae: jl-1?, j4-15, j5-16, j6-26, J 5-5, rs-24, R I -24, ja-21, za-24, z4-25, s4-29, Zl-33, Sa-38, S4-14, ZI)-44, zg-19, Z4-44. Ventrianal shield much longer than wide, lateral margins slightly concave with 3 pairs of preanal setae. Cervix of spermatheca short. Movable digit of chelicera with a tooth. Macrosetae on leg IV : basitarsus-15, distitarsus-13.? Distribution; Japan, Taiwan.

91 GUPTA: ORIENTAL PHY1'OSElIDAE Okiseius chinensis Wei-nan Okiseius chinensis Wei-nan, Entomotaxonomia, 5: 15. (Holotype ~, South China: Yunnen Province, on rubber tree, repository unknown). Diagnosis: Measurements of setae: jl-17, ja-28, j4-18, j5-17, j6-22, J~-37, r a -30, R l -25, s4-37, Sa-32, S~-16, za-32, z4-32, zr,-27, Zl-37, Z4-40, Z5-31, JV South China Okiseius bimalayana Gupta Okiseius himalayana Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p (Holotype ~, India: Uttar Pradesh, Garhwal, Fata village, on an undet. shrub, in ZSI). Diagnosis: Dorsal shield highly rugose, sclerotized, 315 long, 157 wide. Setae J a, S4 absent. Measurements of setae: j1-10, j4-16, j&-16, j6-33, J 5-7, ja-23, z'l-27, z4-34, s4-38, Z1-40, , S~-16, Zs-47, z5-20, Z4-50, ra-27, R l -27. Ventrianal shield about Ii times as long as broad, with 3 pairs of setae. Macrosetae on leg IV: genu-indistinguishable, tibia-indistinguishable, basi tarsus-l 6, all with spa tula te tip. India (Uttar Pradesh) Okiseius sikkimensis Gupta Okiseius sikkimensis Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p (Holotype ~,India : Sikkim, Nathula Road, on undet. plant, in ZSI). Diagnosis: Dorsal shield rugose, sculptured, with L 7 pairs of setae. Measurements of setae: jl-25, j4-5, jls-5, j6-9, J 5-5, ;3-45, za36, Z4t-45, , Zl-47, Sa-56, Sf)-18, Z5-83, z5-6, Z4-65, rs-36, R l -22. Ventrianal shield 105 long, 75 wide, with 3 pairs of preanal setae. Macrosetae on leg IV: genu-25, tibia-21, basitarsus-45, all with spatulate tip. India (Sikkim).

92 78 REC. ZOOL. SURV. INDiA, Dcc. PAPER No Okiseius yazuliensis Gupta Okiseius yazuliensis Gupta, Oriental Ins. (Holotype ~. India : Arunachal Pradesh, Ziro, Yazuli, on an undet. plant, in ZSI) Okiseius yazuliensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 358 long, 212 wide, heavily sclerotized, with 16 pairs of setae. Measurements of setae: jl-22, j4:-j each, j6-23, J 5-7, js-34, za-29, z4:-34, s4-40, ZI-35 Sa-45, SD-16, ZD-45, zd-14, Z4-40, rs-33, R I -22. with 3 pairs of preanal setae. basitarsus-23, disti tarsus-23. Ventrianal shield 112 long, 60 wide, Macrosetae on leg IV : genu-7, tibia-7, India (Arunachal Pradesh). Subfamily Phytoseiinae Berlese Phytoseiinae Berlese, Redia, 12: Phytoseiinae, Murna, Bull. Fla. St. Mus., 5 : Phytoseiinae, Muma & Denmark, Fla. Dept. Agr. & Con. Serv.,6: 115. Diagnosis: Dorsal shield undivided, 5-6 pairs of prolateral setae, 2 pairs of sublateral setae either on lateral integument or on dorsal shield. Ventrianal shield with 1-4 pairs of preanal setae; 0-3 macrosetae on leg IV. Type genus: Phytoseius Ribaga, Key to the genera of Phytoseiinae 1. Both setae r a and R 1 on dorsal shield r a may be on dorsal shield, Rl if present, always on lateral integument 2. r a on dorsal shield, genu II with 7 setae, genu III with 6 setae r a on lateral integument, genu II with 7 or 8 setae; genu III with 6 or 7 setae. Indodromus 2 Phytoseius Typhlodromus Genus Indodromus Ghai & Menon Illdodrol11US Ghai & Menon, Oriental Ins., 3 : Type; lndodromus meerutensis Ghai & Menon, 1969.

93 GuPTA: ORIENTAL PHYTOSElIDAE Indodromus meerotensis Ghai & Menon Indodromus meerutensis Ghai & Menon, Oriental Ins., 3: 349. (Holotype ~, Paratypes 9 ~ ~, India: Uttar Pradesh, Meerut, from malformed mango samples, in NPC) Indodromus meerutensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield reticulate, with 18 pairs of setae; setae being progressively longer posteriorly. Setae Sa' S4' 2 5, 24 being serrate, S 5' J 5 being tiny ; r 3 and Rl both on dorsal shield. Sternal shield with 2 pairs of sternal setae ; 3rd pair on inters cut al membrane and 4th pair on metasternal plates. Ventrianal shield longer (92) than broad, margins irregular. Leg IV without macroseta. India (Uttar Pradesh). Genus Phytoseius Ribaga Phyloseius Ribaga, Riv. Pat. Veg., 10: 175. Type: Phytoseius plumifer Canestrini & Fanzago, 1876 by subs-equent designation (Vitzthum, 1941). Key to the subgenera of Phytoseius 1. Posterior sublateral seta (R 1 ) present Posterior sub lateral seta (R i) a bsen t Pellnaseius Phyloseills Subgenus Pennaseius Pritchard & Baker Key to the species of Pennaseius 1. Dorsal shield without notocephalic pores Dorsal shield with notocephalic pores 2. Z2 and z.. equal, both simple Z2 shorter than Z4., the latter serrate 3. Dorsal shield notched at the level or r:1 Dorsal shield not notched at the level of r s 4. js 2 times as long as Zs - ja about I! times as long as Zs 5. Macrosetae on genu, tibia and basitarsus of leg IV of same length Macrosetae on basitarsus IV much longer than those on genu and tibia IV 2 3 diutius improcerlls 4 5 hongkollgellsis kapuri In inlltlis cui'valll$

94 80 REC. ZOOL. SURV. INDIA, OCC. PAPER No Phytoseius (Pennaseius) curvatus Chaudhri Phytoseius (Pennaseius) curvatus Chaudhri, Pak. J. Zool., 5 : (Holotype ~, Cotoneaster bacci/laris, in UAL). Paratypes 20 d", Pakistan: Sunny Bank, Murre Hills, on Diagnosis: Dorsal shield rugose posteriorly with 15 pairs of setae and a large notocephalic pores near zg. Measurements of setae: i 1-23, i4-8, j5-8, i6-10, J 15-10, is-49, za-16, zs-42, z4-18, s4-88, s6-88, ZG-70, zg-10, Z4-66, rs-47, R Besides Za' i4:-j6 and ZG' other setae serrate. Spermatheca with cervix bowl-shaped. Ventrianal shield longer than wide with 3 pairs of preanal setae. Macrosetae on leg IV : genu-16, tibia-60, basitarsus-24; all with slightly knobbed tip. Metapodal plates 1 paired. Pakistan Phytoseius (Pennaseius) diutius Corpuz Phytoseius diutius Corpuz, Philipp. Agric., SO : (Holotype ~, Philippines: Los Banos, Laguna, on Achyranthes aspera, in Dept. Ento. Univ. Philipp.). Diagnosis: Dorsal shield slender, oblong, 280 long, 124 wide, with 15 pairs of setae. Setae i 1, js, zs, S4' S6' Z4' Z5 long, thick, serrate; other setae small. Measurements of setae: il-20, is-lo, zs-44, s4-84, s6-76, Z4:-64, Z5-64. Metastemal setae on plates. Ventrianal shield longer than wide, laterally constricted with 3 pairs of preanal setae, without pores. Cervix of spermatheca slender, cylindrical at basal half, the distal half widening gradually like a bell; atrium small, swollen. Macrosetae on leg IV on genu, tibia, basitarsus and distitarsus; tip splitted, membraneous and bulbous. Philippines Phytoseius (Pennaseius) hongkongensis Swirski & Shechter Phytoseius (Phytoseius) hongkongensis Swirski & Shechter, Israel J. agrie. Res., 11 : (Holotype ~, Paratype 10, Hongkong: Victoria Mt. Forest, on Heterosmilax gaudichaudiana and 1 paratype ~, locality same, on Urena lobata, all in Univ. Inst. Agr., Israel) Phytoseius (Pennaseius) hongkongensis, Tsens, J. Afr. Assoc. China, N. S., 94 ; 87 (Taiwan).

95 GUPTA: ORIENTAL PHYTOSEIlDAE Phytoseius (Pennaseius) hongkongensis, Ehara, Rev. Pl. Prot. Res 10 ~ Phyloseius (Phytoseius) hongkongensis, Ebara & Bhandhufalck. J. Fac. Ed, Tollori Un;v. Nat. Sci., 27(2) : 46. (Thailand: on lime tree. green apple. Cratoxy/um, Ficus sp.). Diagnosis: Dorsal shield long, with 16 pairs of setae. Measurements of setae: il -25, ;4-i6, zcs-4-b each, 1 a -9-12, 1 G-8-9, ji-64, zl-14, zs-38, z4.-10, s4-102, s6-76, Z5-55, Z4-76, rs-43, R l Ventrianal shield elongate, lateral margins concave, long, wide, with 3 pairs of preanal setae. Leg IV with 4 macrosetae on genu-30, tibia-32, basitarsus-24, distitarsus-26, all with broadened and divided tip. Leg chaetotactic formula: genu IT 2 a a f, genu III 1 A * 1. Chaetotaxy same as in female. Hong Kong, Taiwan, Thailand Phytoseius (Pennaseius) improcerus Corpuz Phyto~eius improcerus Corpuz, Philipp. Agric., 50 : (Holotype ~, Philippines: Maquiling, Laguna, on Schizostachyum lima, in Dept. Ent. Univ. Philipp.). Diagnosis: Dorsal shield 268 long, 118 wide, with 15 pairs of setae; ;1-is' Zs, s4.' s6' Z4' Zcs long, thick and serrate and other setae small. Peritreme extends upto S4. Cervix of spermatheca entirely slender, cylindrical, being thickened only for a short length at the mouth of vesicle; vesicle oblong. Fixed digit of chelicera 2-3 teeth, movable digit unidentate. Philippines Phytoseius (pennaseius) kapuri Gupta Phytoseius (Phytoseius) kapuri Gupta, Israel J. agric. Res., 17 : (Holotype ~, Paratypes 2 ~ ~, 10, India : Howrah Botanical Garden, on Syzygium javanica, in ZSI) Phytoseius (Phytoseius) kapuri, Gupta. Oriental Ins.) 11 : 636. (Andaman IsIs. : Undet. plant) Phytoseius (Phytoseius) kapuri, Gupta, Indian J. Acar., 1 : 12. (Gujarat : Sugarcane, guava; Rajasthan: brinjal ; Punjab: brinjal) Phytoseius (Phytose;us) kapuri, Gupta, Indian J. Acar., 2 : 10. (Orissa: Tendu ; Meghalaya : Undet. plant, brinjal ; Assam: Shorea robusta) Phytoseius (Phytoseius) kapuri, Gupta. Bull. zool. Surv. India, 3 (1 & 2) : 51. (Pondicherry : unknown plant; Kerala: unknown plant; Tamil Nadu: brinjal). 11

96 82 REC. ZOOL. SURV. INDIA, OCC. :PAP-E~ No Phytoseius (Phytoseius) kapuri, Gupta & Nahar. In Contrib. to Acar. in India, p. 10 (Bihar : guava) Phytoseius (Phytoseius) kapuri, Gupta, Fauna of India (Acari: Mesostigmata) Family Pbytoseiidae, p Di~gnosis: Dorsal shield 270 long, 135 wide, notched, at the level of rs, with 16 pairs ()f setae and 4 p$lirs of pores, of those,. ~h~ 9ne associated with Z5 being the largest. Measurements of setae: j1-26, j4,-j8-4 S each, J a -9, J~-4, js-65, za-12, zs-40, z4:-6, s4-14, s6-18, ZG-76, Z4c-76, rs-45. Ventrianal shield 73 long, 47 wide, lateral ~rgj~s concave, with 3 pairs of preanal setae. Spermatheca with funnel7s~aped cervix. Macrosetae on leg IV: genu-27, tibi~-30, basitarsus-24, distitarsus-24. Gupta (19~9) illustrated the spermatophoral proc~ss and ventrianal shield. India (West Bengal, Andaman lsi., Gujarat, Rajasthan, Punjab, Orissa, Meghalaya, Assam, Pondicherry, Kerala, Tamil Nadu, Bihar) Phytoseius (Pennaseius) minutus Narayanan, Kaur &. Ghai PhyllJseius minutus Narayanan, Kaur & Ghai, Pror. natn. Inst. Sci. India, 26B(6) : (Described from 7 ~ ~,So 0, New Delhi, on Hibiscus esculentus. No mentioning has been made about sexes of types, all in NPC) Phytoseius (Phytoseius) minutus, Gupta, Sci. & Cult., 36 : 98. (West Bengal: lady's finger) Phytoseius (Dubininellus) minutus, Gupta & Dhooria, Abstr. Indian Sci. Congr., 1974: 69. (Punjab : Himachal Pradesh) Phytoseius (Pennaseius) minutus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield smooth, with 16 pairs of setae and 8 pairs of small pores, the one associated with Z5 being the largest; setae i4:' id' i 6, 1St J IS, ZIS' Zs, Z4 small, smooth; other setae long and serrate. Measurements of setae: i 1-30, i 4-4, j5-3, j6-5, J a -8, J 5-5, )3-71, za-17, za-52, z4-6, S4:' s6-86 each, ZD-83, Z4-79, r Ventrianal shield longer than wide, with 3 pairs of preanal setae. Spermathe~~ &lender, cervix with bulbous atrium. Macrosetae on leg IV : genu-30, tibia-28, basitarsus-2 7. Dorsal chaetotaxy siil)il~r Distrib"4tion: lkn~al)t to female but setae being -shorter. India (New Delhi, Punjab, Himachal Pradesh, West

97 GUPTA: ORIENTAL PHYTOSEI1DA 83 In addition to the above species, the following are also known hut could not be included in key for the reasons as given for Amblyseius (Amblyseius) spp. on p Phytoseius (Pennaseins) incisus Wei-nan & Zhao-Quan Phytoseius (Phytoseius) incislls Wei-nan & Zhao-Quan, Acta Enlomologica Sinica, 27(4) : (Holotype ~, Paratypes 1 &, 4 ~ ~, South China: Neng Gang, Guangxi, on F,irmiana simpiexus). Diagnosis: Dorsal shield long, wide. Measurements of setae: j , ia-37-40, j4-5, j5-5, i6-7, 15-6, za-7, zs-25, Z4-12, z5-5, Z , Z , S , s , r a-33-37, R l -12, JV South China Phytoseins (Pennaseius) amba Pritchard & Baker Phytoseius (Pennaseius) amba Pritchard & Baker, Hilgardia, 33(7) : (Holotype ~, Belgian Congo, on Tithonia speciosa, in USNM. Para types 3 ~ ~, Belgian Congo, Kysenyi, on Pycnostachus sp. ; 1 ~, same locality, on Pholox; 1 ~, Ruanda Urundi, on Cherimoya; 1 ~, on Bixa ; 10, 1 ~, Ruwinzori, OD Hoslundia opposita ; 7 ~ ~, same locality and habitat, in USNM) Phytoseius (Ptenoseius) ambo, Tseng, J. Agr. Ass. China, 94: Diagnosis: Dorsal shield with 16 pairs of setae. Idiosoma 25~ long, 126 wide. Measurements of setae: i l -25, j4: -j6' J,,-7 each, , j&-42, zs-7, zs-16, z4-13, 54-79, se-57, Z5-79, zls-7, Z4:-55, ra-39, R l -16. Ventrianal shield with 3 pairs of preanal setae, lateral sides concave. Leg IV with macrosetae on genu-is, tibia-20, basitarsus-27. Pritchard &. Baker (1962) described it. Belgian Congo, Taiwan Phytoseius (Pennaseius) namdapbaedsis GUpta Phytoseius (Pennaseius) namdaphaensis Gupta, Oriental Ins. (In Press). (Holotype ~, Para types 2 ~ ~, India: Arunachal Pradesh, Namdapha Sanctuary Area, on an undet. plant, in ZSI) Phytoseius (Pennaseius) namdaphaensis, Gupta, Fauna of India (Acari: Mesos tigmata) Family Phytoseiidae, p Diagnosis: pairs of setae. Dorsal shield smooth, 302 long, 145 wide, with 14 Measurements of setae: jl-24, j4-j6' JIS-very small,

98 84 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 9S is-49, zg-9, Zs -22, Z4-15, S4-95, s8-72, Za-71, z~-small, Z Ventrianal shield 100 long, 56 wide, with 3 pairs of preanal setae. Macrosetae on leg IV : genu-24, tibia-56, basitarsus-17, all with broad tip. India (Arunachal Pradesh) Phy toseius (Peonaseius) taianensis Lai-rong & Li-sbeng Phytoseius taianensis Lai-rong & Li-sheng, Entomotaxonomia, 3(3) : 237. (Holotype ~, Paratypes 2 ~ ~, South China: Tian, Shandong Province, on an undet. plant). Diagnosis: Dorsal shield 306 long, 154 wide. Measurements of setae: i l -27, i a -J8, i4-6, ia-5, i8-6, J 5-6, za-12, zs-31, Z4: -18, z&-5, Z4: -46, Z&-74, s4-67, s8-71, ra-41, R l -17, JV Macrosetae on tibia IV-61, basitarsus IV -24. South China. Subgenus Phytoseios Ribaga Key to the species oj Pbytoseius 1. Macrosetae absent on leg IV Macrosetae present on leg IV 2. s. and Z. large, flat and divided s. and Z. not flat and divided 3. r I longer than j a r. and j. equal 4. Genu IV without macroseta... Genu IV with macroseta s. Macroseta on basitarsus IV subequal to that of tibia IV. Macroseta on basitarsus IV smaller than that of tibia IV 6. Macroseta on tibia IV as long as segment.. - Macroseta on tibia IV shorter than that of segment.. 7. ZII and z. shorter, smooth - Zi and z. longer and serrate 8. Cervix of spermatheca cylindrical Cervix of spermatheca gradually enlarging towards mouth 9. Cervix of spermatheca 2 times as long as wide Cervjx as long as or only slightly longer than wid" s" longer than 9ft s. as long as or shorter than S6 intermedius crinitus neofelox S 10 macrosetoslu roseua mixtus 9 mansehraensis neocorniger jujuba 19 11

99 GUPTA: ORIENTAL PHYTOSEIiDAE 11. j. only slightly longer than ZI j. much longer than Zs 12. Posterior margin of sternal shield lobed... Posterior margin of sternal shield straight 13. Za longer than Z4 Za shorter or equal to Z4 14. Macroseta on genu IV very small, about 1/4 the length of that of basitarsus IV Macroseta not as above IS. Cervix ofspermatheca broad towards atrium Cervix of spermatheca narrowed towards atrium 16. Macroseta on genu IV and basitarsus IV equal Macroseta on genu IV and basitarsus IV unequal 17. Cervix of spermatheca uniformly cylindrical Cerv ix of spermatheca not uniformly cylindrical 18. Cervix bell-shape d Cervix elongated, widening at the mouth 19. su Z6 and Z4 almost equal or subequal... Se, Za and Z4 unequal 20. Macroseta on genu IV and basitarsus I V equal Macrosetae on genu IV and basit arsus IV unequal Cervi x of spermatheca Y -shaped Cervix of spermatheca as shallow arc 22. Macrosetae on leg IV spatulate Macrosetae on leg IV not spatulate 23. Z4 Ii times as long as ZI Z4 as long as ZI 24. S4 much longer than S6 5. only slightly longer than Se 25. Macroseta on tibia IV 4 times as long as that on genu IV Macroseta on tibia IV 2 times as long as that on lenu IV 26. Macroseta on leg IV bulbous Macr05eta on leg IV with hyaline membrane brevicrinis meyerae rimandoi 15 jaunpurensis punjabensis indicus 17 swirskii 18 coheni huangi macropilis bandipurensis glareosus wainsteini corniger 25 hawa;iensis 26 rachelae rasllis spathulatus 169. Phytoseius (Phytoseius) bandipurensis Gupta Phytoseius (Dubininellus) bandipurensis Gupta, Bull. zool. Surv. India, 3(1-2) : S3-S4. (Holotype ~,India : Karnataka, Bandipur Sanctuary, On undet. plant, in 2SI) Phytoseius (Phytoseius) bandipurensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis; Dorsal shield heavily sculptured with 15 pairs of setae; i1 very close to each other. Measurements of setae: jl-24, ja-28 II and z..-8 each, za-28, s4-88, so-56, Z~-64, Z4-72, rs-36. Ventrianal

100 86 REe. zooi. SURV. INDiA, OCC. PAPER No. 95 shield short, lateral margins deeply concave with 3 pairs of preanal setae. Spermatheca with a Y -shaped cervix and a long duct. Fixed digit of chelicera with 2 teeth, none on movable digit. Macrosetae on.leg IV : genu-18, tibia-40, basitarsus-14, distitarsus-20; all with spatulate tip. India: Karnataka Phytoseius (Pbytoseius) brev~crinis Swirski & Shechter Phytoseius (Dubininellus) brevicrinis Swirski & Shechter, Isreal J. agric Res., 11(2): (Holotype ~, Hong Kong: Tai Po, on Bambusa, sp., Paratype 1 ~,Hong Kong, Sha Tin, on Bambusa, all in Univ. Inst. Agr., Israel) Phytoseius (Plzytoseius) brevicrinis, Ebara & Bhandbufalck, J. Fac. Ed. Totiori Univ. Nat. Sci., 27(2): (Redescribed ~,and described male for first time-thailand, on bamboo). Diagnosis: Dorsal shield rugose, ii' rs, S5' S6 and Z5 stout, strongly serrate, Z4 stout, narrower than Z 5' weakly serrate, is, Zs and' z~ smaller, more or less serrate others smooth, small. Measurements of setae: jl-22, j4-10, j5-9, j6-11, 15-7, ra 30, js-14, za-11, zs-18, z4-38, s6-41, Z5-44, z5-11, Z4-42. Ventrianal shield much longer than wide, lateral margins concave, with 3 pairs of preanal setae. Fixed digit of chelicera with 3 subapical and 1 middle tooth, movable digit with 1 tooth. Chaetotactic formula of leg: genu II 2 a a * 1, genu IiI 1 1. Leg IV with macrosetae on genu-8, tibia-30, basitarsus-19, distitarsus-19, all with hyaline envelope. Chaetotaxy same as in female. Hong Kong, Thailand Phytoseius (Phytoseius) eoheni Swirski & Shechter Phytoseius (Dubinille/lus) macropilis cohen; Swirski & Shechter, Israel J. agrie. Res., 11(3-4): (Holotype ~, Paratypes 11 ~ ~, Hong Kong: on Pomelo, in Univ. Inst. Agr. Israel) Phytoseius (Dubininelllls) coheni, Gupta, Sci. & Cult., 36: 98. (West Bengal on Diploknema butyracea)-misidentification Phytoseius coheni~ Gupta et al., Sci. & Cult., 37: 298. (India: Punjab" on guava, feeding upon Brev;pa!pus sp. )-misidentification Phytoseius (Dubininellus) coheni, Lo, Bull. Sun-Yat-Sen Cult. Found., No. S: (Taiwan : Tai Pai on citrus, guava).

101 GVPTA: ORIENTAL PHYTOSEIIDAE 87 Diagnosis: Dorsal shield 295 long, 180 wide, well sclerotized, ru~ose, with 15 pairs of setae, setae ji' j4' j5' Z5 and J 5 short and smooth. Measurements of setae: jl-29, j4-7, j5-6, je-9, J 5-8, z5-8, js-2b, za-14, zs-29, z4-15, S4-90, s6-82, Z5-',0, Z4' r8-41 each. Metastemal s~tae on metasternal plates. Ventrianal shield longer than wide, with 3 pairs of preanal setae. Spermatheca with bell-shaped cervix and slightly enlarged atrium. Chaetotaxy similar to that of female. process with shaft Ii times as long as foot. Hong Kong, Taiwan Pbytoseius (Pbytoseius) corniger Wainstein Spermatophoral Phytoseius (Dubininellus) corniger Wainstein~ Zool. Zh. Ukr.,38: (Holotype and Para type not mentioned in original publication. However, types were from the following localities: Tadzikistan, South Kazakhstan, Shelkovitsa, Inzhir, Isblonia, Vinograd) Phytoseius (Dubininellus) corniger, Denmark, Fla. Dept. Agr. Bul/., 6 : (Pakistan: Behrain, on fig leaves) Phytoseius (Phytoseius) corniger, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 260 long, 140 wide, rugose with 14 pairs of setae. Measurements of setae: ji-28, ja-48, za-32, Z2' z~ -12 each, S4-128, s6-27, Z5-64, Z4-84. Ventrianal shield elongate, narrow, lateral margins deeply concave with 3 pairs of preanal setae. Fixed digit of chelicera with 3 teeth, movable digit with one tooth. Spermatheca with elongate cervix and a fairly long and broad duct. Macrosetae on leg IV : tibia-34, basitarsus-23. Chaetotaxy shorter than female. Shaft of spermatophoral process 1~ times as long as foot. Pakistan, USSR Phytoseius (Phytoseius) crinitus Swirski & Shechter Phytoseius (Duhininel/us) crinitus Swirski & Shechter, Israel J. agrie. Res., 11(2): (Holotype ~,Paratypes 13 ~ ~, Hong Kong: Victoria. Mt. forest, on lfomalium conchinchinense, in Univ. Inst. Agr., Israel) Phytqseius (Phytoseius) crinitus, Ray and Gupta, Rec. zoo!. Surv. Illdia, 80 : (Assam: on undet. plant)-redescribed Phytoseius (Phytoseius) crinitus, Gupta, Fauna of India (Acari: Mesostismata) Family Pbytoseiidae, p

102 88 REC. ZOOL. SURY. INDIA, OCC. PAPER No. 95 Diagnosis: Dorsal shield 280 long, 168 wide, with 15 pairs of setae. Measurements of setae: j1-29, j4,-ie, zd' J D-4-5 each, js-40, za-18, zs-31, Z4: -11, s4-10, s6-67, Z5-65, Z4-83, r,s -38. Ventrianal shield longer (90) than wide (56), with 3 pairs of preanal setae. Fixed digit of chelicera 2-dentate, movable digit I-dentate. Macrosetae on leg IV : genu-31, tibia-50, basitarsus-2 7, distitarsus-22, all being spatulate. India: Assam, Hong Kong Phytoseius (Pbytoseius) glareosus Corpuz Phytoseius g/areoslls Corpuz, Philipp. Agrie., SO: 738. (Holotype ~, Philppines: College campus, Los Banos, Laguna, on Oryza sativa, in Dept. Ento., Univ. Philip.) Diagnosis: Dorsal shield coarsely pebblicate, 288 long, 160 wide, with 15 pairs of setae, except Za' j,. -j5' J D' other setae thick, long and serrate. Ventrianal shield longer than wide, slightly constricted laterally at midway, with 3 pairs of preanal setae. Spermatheca with cup-shaped cervix. Fixed digit of chelicera multidentate, movable digit unidentate. Macrosetae on leg IV present on genu# tibia, basitarsus and distitarsus, all strongly bulbous at tips. Philippines Pbytoseius (Pbytoseius) bawaiieosis Prasad Phytoseius hawaiiensis Prasad, Ann. ent. Soc. Am., 61 : (Holotype ~,U. S. A. : Oahu, Hawaii, Poinsettia leaf, in B. P. Bishop Museum., Honolulu. Paratypes: in BM (NH); BP. Bishop Musoum: Canadian Nat. Coll; Univ. Hawaii; Inst. Acarology? Wooster, Ohio; US Nat. Mus., ZSI (Number and sex of specimens sent in different Museums not specified by author in oriaina1 publication) Phytoseius (Phytoseius) hawaiiensis, Ebara & Bbandhufalek, J. Fac. Ed. Totlor; Univ. Nat. Sci., 27(2) : (Thailand on A/angillm sa/vifoiium, Pinus kesiya, Bridelia sp., Bridelia retusa. Coleus blumei), (Redescri bed ~, 0 described for first time). Diagnosis: Dorsal shield rugose, with 2 pairs of conspicuous proes. Setae S4, se, ZD' Z4t and rs, stout, strongly serrate; j1' is, ZSh Zs and z.. serrate, zs, Z4: small. Measurements of setae: jl-26, j4:-9, j&-9, je-10, J a -9, r3-36, ja-22, zj-13, zs-28, z4-13, S4.-80, s6-67, ZD-61, za-10, Z6-53 Ventrianal shield mllch longer than wide, narrower than senita!

103 GUPTA: ORIENTAL PHYTOSEllDAE 89 shield, lateral margins concave, with 3 pairs of preanal setae and a pair of pores. Chaetotactic formula: genu II 2 a g 1. Macrosetae on leg IV: genu-21, tibia-4, basitarsus-23, distitarsus-21, all with hyaline tip. Ehara & Bhandhufalck (1977) described it. Hawaii, Thailand Phytoseius (Phytoseius) hu8ngi Ehara Phytoseius (Phytoseius) huangi Ehara, Mushi, 43(6) ~ 62. (Holotype ~ I Taiwan: Fengshan, on Pithecolobium dulce, in Bioi. Inst. Fac. Ed. Tott. Univ.). Diagnosis: Dorsal shield 275 long, 150 wide, with 15 pairs of setae. Measurements of setae: i}-24, j4 -j6' J long, jg-22, zg-13, zg-26, Z4-9, 54-75, , Z5-60, Z4-51. Ventrianal shield elongate, lateral margins constricted with 3 pairs of preanal setae. Fixed digit of chelicera with 3 teeth, movable digit with one tooth. Macrosetae on leg IV: genu-15, tibia-38, basitar5us-22. Taiwan Phytoseius (Phytoseius) intermedius Evans & Macfarlane Phytoseius (Dubininel/us) intermedius Evans & Macfarlane, Ann. Mag. nat. Hist., (13)4 : (Holotype ~, Paratypes 2 ~ ~, South Rhodesia: Mazoe, on Aloe; all in BM(NH) Phytoseius (Dubininellus) intermedius, Denmark, Fla. Dept. Agr. Bull., 6 : (Pakistan: papaya) Phytoseius (Dubininel/us) intermedius, Gupta, Sci. & Cult 36: 98. (West Bengal: Undet. plant) Phytoseius (Dubininellus) intermedius, Gupta, Oriental Ins., 11 : (Andaman lsi. : Uundet. plant) Phytose;us (Phytoseius) intermedius, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 275 long, 150 wide, with 15 pairs of setae. Measurements of setae: j 1-22, j4 -j6' J each, js~22, zg-18, zs-31, z4-13, 54-54, s6-72, Z )-54, z5-5, Z4-76. Sternal, metasternal and genital shields normal. Ventrianal shield 76 long, 45 wide, with 3 pairs of preanal setae. Macrosetae absent on leg IV Spermatheca with bellshaped cervix. 12 Denmark (1966) described it. India (West Bengal), Pakistan, South Rhodesia.

104 90 REC. ZOOL. SURV. INOlA, Oce. PAPBR No Phytoseius (Phytoseius) jujuba Gupta Phytoseius (Dubininel/us) jujuba Gupta, Indian J. Acar., 1 : 13. (Holotype ~,India : Gujarat, Zizyphus jujuba. Paratypes 3 ~ ~, data as for holotype, all in ZSI). 1 0, same Phytoseius (Phytoseius) jujuba, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 270 long, 130 wide, with 15 pairs of setae, all the setae being minute measuring 5-<, except js, Zg, s4' Z5' Z4' ra which measure 36, 33, 66, 69, 72 and 40, respectively. Sternal and genital shields normal, metasternal plates absent. Ventrianal shield longer (68) than wide. Spermatheca with flask-shaped cervix and elongate duct. Macrosetae on leg IV : genu-nil, tibia-36, basitarsus-20. Gupta (1977) collected male but did not describe it. India (Gujarat) Phytoseius (Phytoseios) macropilis (Banks) Sejus macropilis Banks, Proc. ent. Soc. Wash., 11 : 135. (Holotype ~, Canada: Guelph, Ontario, attacking Eriophyes sp., on willow, in USNM) Phytoseius macropilis, Cunliffe & Baker, Pinellas BioI. Lab. Pub, No.1: Phytoseius macropilis, Narayanan, Kaur & Ghai, Proc. natn. Inst. Sci. India, 26 B(6) : (New Delhi : on fig, mulberry, Compositae, Lantana camara) Phytoseius (Dubininellus) woodburyi, Gupta, Sci. & Cult., 36 : Phytoseius macropilis, Gupta, En!. Rec., 86 : Phytoseius (Dubininel/us) macropilis, Gupta, 1977, Indian J. Acar., 1 : 9. (Orissa: Shorea robusta) Phytoseius (Dubininel/us) woodburyi, Gupta, Indian J. Acar., 6: Phytoseius (Phytose/us) macropilis, Gupta, Fauna of India (Acari: Mesostig-) mata) Family Phytoseiidae, p Diagnosis: Dorsal shield rugose, 280 long, 150 wide, with 15 pairs of setae. Measurements of setae: jl-23, j4' js' z5' j5 minute (5-6 each), i s -36, za-5, zg-25, Z4-5, s4-100, 8s-76, ZD-70, Z4-70, r g-44. Sternal, metasternal and genital shields normal. Ventrianal shield elongate, vase-shaped, lateral margins concave, 80 long )2 wide, with 3 pairs of preanal setae. Macrosetae on leg IV : genu-20, tibia-48, basitarsus-32. Denmark (1966) described it. India (Andhra Pradesh, Orissa, New Delhi) ; Canada; and many other parts of the world (Gupta, 1986).

105 GUPTA: ORIENTAL PHYTOSElIDAE Phytoseius (Phytoseius) macrosetosus Gupta Phytoseius (Dubininellus) Illacrosetosus Gupta, indian J. Acar., 1: (Holotype ~, Paratype 1 ~, India: Madhya Pradesh, Shivpuri, on Tendu, all in ZSI) Phytoseius (Phytoseius) macrojetosus, Gupta, Fauna of India (Acari: Mesostigmata) Family Pbytoseiidae, p Diagnosis; Dorsal shield 265 long, 130 wide, with 15 pairs of setae. Measurements of setae; j1-20, j4' js, J 5' z6-5 each, ja-36, za-16, zs-32, z4-24, s4-62, ss-64, Z5-50, Z4-52, rs-32. Sternal and genital shields normal. Metasternal plate absent. Ventrianal shield 56 long, 40 wide, with 3 pairs of preanal setae, lateral sides concave. Macrosetae on leg IV: genu-nil, tibia-18, basitarsus-20, both with spatulate tip. DistributiQn: India (Madhya Pradesh) Phytoseius (Phytoseius) mansebraensis Chaudhri Phytoseius (Phytoseius) mansehraensis Chaudhri, Puke J. Zoo/., 5(1) : (Holotype ~, Pakistan: 2 miles south of Mansehra, on Inula cuspidata, in UAL). Diagnosis: Dorsal shield rugose, wi th a pair of pores near Z ~ and posterior to S6 laterally, with 15 pairs of setae. Measurements of setae: jl-34, j4-j6' Z5' J 5-minute, js-55, za-16, zs-34, Z4-23, S4-127, 5673, Z5-83, Z4-86, rs-47. All setae except Za' j4-j6' J 5, Z5 serrate. Ventrianal shield longer than wide, with 3 pairs of preanal setae and a pair of pores. Spermatheca with cervix funnel-shaped. Macrosetae on leg IV : genu-absent, tibia-52, basitarsus-26, all blunt. DistributiQn: Pakistan Pbytoseius (Phytoseius) meyerae Gupta Phytose;us (Dubininellus) meyerae Gupta, Indian J. Acar., 2 : 7. (Holotype ~,India : Meghalaya, Tura, on weed ; Paratypes 1 0, same data as for holotype. 1 ~,Assam, Gossaigaon, banana, all in ZSI) Phytoseius (Phytoseius) meyerae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p DiagnQsis: Dorsal shield rugose, 260 long, 170 wide, with 15 pairs of setae, all being thin, gently serrate. Measurements of setae: j1-20, j4-;6' 1 5, zf)-8 each, ja-20, za-12, za-20, Z4-20, S4-20, s6-60, Z~-37,

106 92 REC. ZOOL. SURV. INDIA, OCC. PAPER No.9; r s -28, Z4-44. Sternal and genital shields normal. Ventrianal shield 72 long, 50 wide, with 3 pairs of preanal setae. Spermatheca with thick flask-shaped cervix. Macrosetae on leg IV: genu-7, tibia-36, basitarsus-20, all with spatulate tip. Gupta (1977) illustrated its spermatophoral process. India (Assam, Meghalaya) Phytoseius (Phytoseius) mixtus Chaudhri Phytoseius (Phytoseius) mixtus Chaudhri, Pak. J. Zool., 5(1): (Holotype ~, Pakistan: North of Haripur, on mulberry, in UAL) Phytoseius (Phytoseius) mixtus, Ray & Gupta, Rec. zool.. Surv. India, 80(3-4) : (Redescribed) (Uttar Pradesh, on grass) Phytoseius (Phytoseius) mixlus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagonosis: Dorsal shield rugose, 250 long, 135 wide, with 15 pairs of setae and a few pairs of pores. Measurements of setae: j1-29, j4-j6, J 6, 4-5 each, js-56, z2-16, zs-36, z4-20, s4-130, s6-78, z5-5, Z5-72, Z4-95, rs-52, ; 24 thicker than ZIS. Ventrianal shield longer than wide with constriction little below the level of 3rd pair of preanal setae. Fixed digit of chelicera with 2 teeth anterior to pilus dentilis, movable digit with 1 tooth. Spermatheca with tubular cervix and knobbed atrium, major duct long. Macrosetae on leg IV: tibia-56, basitarsus-31, distitarsus-31. India (Uttar Pradesh), Pakistan Pbytoseius (Phytoseius) indicus Bhattacharyya Phytoseius (Phytoseius) indicus Bhattacharyya, J. Born. Nat. Hist. Soc., 65(3) : , (Holotype ~,India : W, Bengal, Sonarpur, Sitala, on Hibicus sp., in ZSI 1) Phytoseius (Dubininelllls) neglecta Gupta, Israel. J. agric. Res., 19: (West Bengal: Saraca india) Phytoseius (Dubininel/us) neg/ecta, Gupta & Dhdoria. Proc. Indian Sci.Congr., 61 : Phytoseius /leg/ecta, Prased, A catalogue of mites of India, p PIzytoseius indicus, Prasad, A catalogue of mites of India, p Phytoseills (Plzytoseius) illdicus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p

107 GUPTA: ORIENTAL PHYTOSEllDAE 93 Diagnosis: Dorsal shield 299 long, 145 wide, rugose, with 15 pairs of setae. Setae j4-j61 J 5' Za' Z4' Z5 short and simple, other setae being long, thick and serra tee Measurements of setae: j 1-27, j a -27, Z a -29, s4-72, s6-72, Z5-50, Z4-65, ra-36. Ventrianal shield twice as long as broad, with 3 pairs of preanal setae. Spermatheca with cup-shaped cervix. Macrosetae on leg IV: genu-18, tibia-40, basitaruss-18, distitarsus-22. India (West Bengal) Phytoseios (Phytoseius) neocorniger Gupta Phyloseius (Dubillinellus) neocorniger Gupta, Indian J. Acar., 1 : (Holotype ~,India: Rajasthan, Pratapgarh, on Lantana. Pratypes 2 ~ ~, sa~e data as for holotype, in ZSI) Phytoseius (Phyloseius) neocorniger, Gupta, Fauna of India Acari: Mesostig mata) Family Phytoseiidae, p Diagnosis: Dorsal shield highly sclerotized, 284 long, 155 wide, with 15 pairs of setae. Excepting setae j4 -i 6, J 5' Za' Z4' Z5' which are small, other setae thick and serrate. Measurements of setae: j1-28, ja-44, za-12, zs-33, z4-12, s4-120, s8-80, Z5-70, Z4-75, rs-40. Sternal and genital shields normal; metasternal plates absent. Ventrianal shield 60 long, 40 wide, constricted at preanal region with 3 pairs of preanal setae. Spermatheca with tubular flask-shaped cervix. Macrosetae on leg IV : genu-nil, tibia-40, basitarsus-26. India (Rajasthan) Phytoseiu.s (Phytoseius) neofero~ Ehara & Bhandhufalck Phytoseius (Phytoseius) neoferox Ebara & Bhandhufalck, J. Fac. ed. Tottori Univ. Nat. ~c;., 27(2) : (Holotype ~,Thailand, Fang, on persimmon, in BioI. Inst. Fac. Ed. ToU. Univ. Paratype 1 ~, same locality, on coffee; repository-not specified) Phytoseius (Phytoseius) neoferox, Gupta, Fauna of India (Acari; Mesostigmata) Family Pbytoseiidae, p Diagnosis: Dorsal shield rugose, pores present behind ZI5 with 15 pairs of setae; j1' js, ZS' S4' S6' Z5' Z" and rs stout and serrate and ra divided, other setae small. Measurements of setae: j1-28, j4-4, j5-4, i8-5, ia-38, za-12, zs-26, Z4-9, S4-93, s6-69, ZI5-63, z5-4, Z4-74. Ventrianal shield much longer than wide, lateral margins constricted at preanal

108 94 REC. ZOOL. SURV. INDIA, Occ. PAPER No. 95 level with 3 pairs of preanal setae and a pair of pores. Spermatheca with cervix tubular. Macrosetae on leg IV: genu-22, tibia-44, basitarsus-23, distitarsus-21. Thailand Phytoseius (Phytoseius) punjabensis Gupta Phytoseius (Dubininellus) pubjabensis Gupta, Indian J. Acar., 1: 16. (Holotype ~,Indla : Punjab, on "fig, in ZSI) Phytoseius (Phytoseius) jaunpurens;s Gupta, Rec. zoo/. Surv. India, 79: (Uttar Pradesh: Jaunpur, on Zizyphus jujuba) Phytoseius (Phytoseius) punjabensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield rugose, 275 long, 140 wide, with 15 pairs of setae. Setae jl' js' zs' S4' 56' Z5' ra, Z4 being long, thick and serrate, other setae minute and simple. Measurements of setae: i l - 24, js-48, zs-32, Z4-10, S4-97, s6-97, Z5-64, Z4-72, ra-44. Ventrianal shield 76 long, 98 wide, with 3 pairs of preanal setae. Spermatheca with elongate funnel-shaped cervix. Macrosetae on leg IV: genu-16, tibia-36, basitarsus-24. India (Punjab) Phytoseius (Phytoseius) rachelae Swirski & Shechter Phytoseius (Dubininellus) rache/ae Swirski & Shochter, Israel J. agrie. Res., 11 : (Holotype ~,Hong Kong: on Rhus chinensis. Paratypes 2 ~ ~, Hong Kong, on Glochidion criocarpum, in Univ. Inst. Agr., Israel) Phytoseius (Dubininellus) reehelae, Gupta, Bull. zool. Surv. India, 3(1-2) : 52. (Kerala: Hibiscus) Phytoseius (Phytoseius) rache/ae, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, Diagnosis: Dorsal shield 268 long, 128 wide, with 14 pairs of setae. Setae ji = js ; S4 being longest; s6>z5>z4 ; measurements of setae: i l - 28, i 4 -i6' each, js-28, zs-8, zs-20, z4-14, s4-100, s6-80, Z&-76, z&-5, Z4-64, rs-40. Ventrianal shield 64 long, 44 wide, with 3 pairs of preanal setae. Spermatheca with 'Y-shaped cervix. Macro setae on leg IV : genu-12, tibia-48, basitarsus-20, all spatulate. India (Kerala), Hong Kong.

109 GUPTA: ORIENTAL PHYTOSEIIDAE Phytoseius (Phytoseius) rasilis Corpuz Phytoseius rasilis Corpuz, Philipp.;Agric., 50 : (Holotype ~,Philippines: Bigger Balatero, Puerto Galera, on Wedelia bijlora, in Dept. Ento. Univ. Philip. Paratypes 1 ~, Nasugbu, on Desmodium gangeticum; 1 ~, on Flemingiu strohilifera, in Dept, Ento. Univ. Philip.). Diagnosis: Dorsal shield 288 long, 152 wide, with 15 pairs of setae: j1-16, js-33, zs-24, s4-116, s6-66, Z4-80, Z5-72, r 3-40 ; other setae small. Metasternal setae on plates. Ventrianal shield longer than wide, constricted at anterior third, widening abruptly at anal region, with 3 pairs of preanal setae and a pair of pores. Metapodal plates absent. Fixed digit of chelicera 2-dentate, movable digit I dentate. Leg IV macrosetae on genu-10, tibia-46, basitarsus-26, distitarsus-25, all but that on distitarsus with bulbous tip. Cervix of spermatheca slender, cylindrical, widening slightly into the vesicle. Male; Philippines Phytoseius (Phytoseius) rimandoi Corpuz Phytoseius rimandoi Corpuz, Philipp. Agric., 50 : (Holotype ~, Philippines: College Campus, Los Banos, Laguna, Bambusa sp., in Dept. Ento. Univ. Philip. Paratypes 12 ~ ~, same data as for holotype ; 3 ~ ~, Mt. Maquiling, Laguna, 300 Mt. elevation, on Schizostachyum lima, in Dept. Ento. Univ. Philip.). Diagnosis: Dorsal shield with pebble-like marking, 332 long, 186 wide, with 15 pairs of setae. i 1-26, js-16, za-24, z4-22, s4-48, se-58, Z4-56, Z5-52 long, ra-36 thick and serrate; others small. Metasternal setae on plates. Ventrianal shield longer than wide, constricted laterally at the middle, with 3 pairs of preanal setae. Peritreme extends anteriorly upto j 1. Fixed digit of chelicera 3-dentate, movable digit I-dentate. Leg IV with macrosetae on genu-6, tibia-42, basitarsus-24 and distitarsus-24; all thick and with bulbous tip. Cervix of spermatheca cup-like, atrium a small globular chamber. Philippines Phytoseius (Phytoseius) rugosus Denmark Phytoseius (Dubininel/us) rugosus Denmark, Fla. Dept. Agr. Bull., 6 : 100. (Holotype-O', Pakistan: Rawalpindi, name of plant not mentioned, in USNM. Paratype-l nymph, Sialkot, on Zzyphus sp., Florida St. CoUectjoQ of Arthropods, GainsviUe).

110 96 REC. ZOOL. SURVE. INDIA, Oce. PAPER N Phytoseius (Dubininellus) rugosus, Gupta, Indian J. Acar., 2: 7-8 (Redescribed). (Orissa: TeYltliltalia arjuna) Phytoseius (Phytoseius) rugosus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: (known from conly): Dorsal shield 200 long, 155 wide, rugose, with 15 pairs of setae, of those, j4 -j6' J 5' ZI5' Za, Z4 small, smooth, other setae being long, thick and serrate. Measurements of setae: jl-19, js-18, zs-24, s4-45, s6-44, ZI5-29, Z",,-26, r s-26. Ventrianal shield with 3 pairs of preanal setae. Macrosetae on leg IV : genu-6 (peg-like), tibia-7, basitarsus-28. Distribution; India (Orissa), Pakistan Pbytoseios (Pbytoseios) roseus Gupta Phytose;us (Dubinine/lus) roseus Gupta, IJrael J. agric. Res., 19 : (Holotype ~,India: West Bengal, on rose, in ZSI) Phytoseius (Phytoseius) rugulosus Chaudhri, Pak. J. Zool., 5(1) : (Pakistan : Daphar, on Canabis sativa) Phytoseius (Dubininellus) roseus, Gupta, Indian J. A car., 1 : (Gujarat : rose; Punjab: milkweed, Zizyphus sp., fig) Phytoseius (Dubininel/us) roseus, Gupta, Bull. zool. Surv. India, 3(1-2) : SI-S2. (Tamil Nadu : Papaya) Phytoseius (Phyloseius) roseus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dors al shield 285 long, 145 wide, rugose, with 15 pairs of setae, of those, i 4, j6' J 5' ZI5' Za, Z4.' small, smooth; other setae being long, thick and serrate. Measurements of setae: jl-27, js-40, zs-31, s4, -108, s6-70, Z5-81, 24-81, rs-45. Ventrianal shield 81 long, 50 wide, with 3 pairs of preanal setae. Spermatheca with bowl-shaped cervix. Macrosetae on leg IV: tibia-36, basitarsus-22, distitarsus-27; those of the first two being spatulate. Pakistan. India (West Bengal, Gujarat, Punjab, Tamil Nadu), 193. Phytoseios (Phytoseios) spathulatos Chaudhri Phytoseius (Phytoseius) spathulatus Chaudhri, Pak. J. Zool 5(1) : (Holotype ~, Pakistan: 3 miles west of Hyderabad, on undet. plant, in UAL). Diagnosis: Dorsal shield rugose with a pair of pores neal' Z 15' with 15 pairs of setae. Measurements of setae: jl-29, j4-j6' J IS' zls-minute, is-44, za-13, zs-34, z4-13, s4:-114, s6-75, ZI5-73, Z4-19, r8-42. Besides z~-z4' j4 -j6' z~, other setae thick and serrate, V ~ntrianal ~hield longer

111 GUPTA: ORIENTAL PHYTOSEUDAB 97 than wide with 3 pairs of preanal setae. Cervix of spermatheca long, tubular. Metapodal plate 1 paired. Leg IV with macrosetae on genu-10, tibia-39 and basitarsus-23, all spatulate with hyaline membrane. Pakistan Phytoseius (Phytoseius) swirskii Gupta Phytoseius (Dubininellus) swirskii Gupta, Entomologists' mono Mag., 115: (Holotype ~,India : West Bengal, Burdwan Agr. Farm, on Ficus sp., in ZSI) Phytoseius (Phyroseius) swirskii, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield rugose, 280 long, 170 wide with 15 pairs of setae. Measurelnents of setae: jt-20, ja-25, za-26, Z4-12, S4, -70, S6-70, Z5-58, Z4-47, ra-32, all being thick and serrate; other setae tiny. Ventrianal shield longer (82) than broad, with 3 pairs of preanal setae. Metapodal plates one paired. Macrosetae on leg IV : genu-ii, tibia-36, basitarsus-21, all being spatulate. India (West Bengal) Phytoseius (Phytoseius) wainsteini Gupta Phytoseius (Dubinine/lus) wainsteini Gupta, Indian J. Acar., S : (Holotype ~, Paratype 1 ~,India: Jammu & Kashmir, Pahalgam, on undet. plant, in ZSI) Phytoseius (Phytoseius) wainsteini, Gupta, Fauna of India (Acari: Mesotigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 270 long, 145 wide, rugose, with 15 pairs of setae. Measurements of setae: ji-20, j4-j6' lrs small, js-56, za-16, zs-32, z4-28, s4-12, s6-68, Z5-72, Z4-84, rs-36; Z4 thicker than Zts. Ventrianal shield elongate, 80 long, 40 wide with 3 pairs of preanal setae. Fixed digit of chelicera with 3 teeth, movable digit with one tooth. Macrosetae on leg IV : genu-is, tibia-67, basitarsus-32. India (Jammu & Kashmir). In addition to the above, the following species are also known but could not be included in the key for the reasons as mentioned for Amblyseius (Amhlyseius) spp. on p

112 98 REc. ZOOL. SURV. INDIA, OCC. PAPER No Phytoseius (Phytoseius) domesticus Rather Phytose;us domesticus Rather, Acar%gia, 26(1) : (Holotype-Particulars not available to the author). Diagnosis: Due to non-availability of the concerned literature the diagnostic characters could not be mentioned. 1 India (Kashmir) Phytoseius (Phytoseius) longus Wei-nan & Zhao-Quan Phytose;us (Dubin;nellus) longus Wei-nan & Zhao-Quan, Acta Zootaxonomica Sinica, 10(4) ~ Diagnosis: Dorsal shield 255 long, 155 wide. Measurements of setae: jl-22, j4' jrs' J rs' zrs-5 each, j6-l0, js-37, za-12, zs-30, z4-12, ZB-90. Not known. South China Phytoseius (Phytoseius) nipponicus Ehara Phytoseius (Duhinine/lus) nipponicus Ehara, Jap. J. apple Ent. & Zool., 6(1) : S5-S8. (Holotype ~,Japan : Den-n-Chofu, Tokyo, on chrysanthemum, in Zool.lnst. Fac. Sci. Hokkaido Univ.) Diagnosis: Dorsal shield faintly reticulate. Idiosoma 360 long, 230 wide. Measurements of setae: il-31, js-31, zs-31, S4-93, s6-85, Zrs-79, Z4.-73, rs-50, JV rs-54 long. Macrosetae present on tibia and basitarsus IV. Ehara (1962) described it. Japan, South China Phytoseius (Phytoseius) petentis Chaudhri, Akbar & Rasool Phytoseius (Phytoseius) petent;s Chaudhri, Akbar & Rasool, Rept. PL 480 Project, UAF., p.233. (Holotype ~,Pakistan : Faisalabad, on cotton, in UAF). Diagnosis: Dorsal shield rugose. Measurements of setae: jl-29, js-40, zs-13, zs-39, z4-13, 8'6-74, Zrs-68, Z4-82, rs-45. Ventrianal shield 90 long, 52 wide, with 3 pairs of preanal setae. Macrosetae on leg IV: tibia-39, basitarsus-25, distitarsus-45. Spermathecal atrium nodular.

113 GUPTA: ORIENTAL PHYTOSEIIDA 99 Pakistan Phytoseius (Phytoseius) qianshanensis Lai-rong & Li-sheng Phytoseius qianshanensis Lai-rong & Li-sheng, ~ntomotaxonomia. 3(3) : 237. (Holotype ~,Paratypes 3 ~ ~, South China: Qianshan, Liaoning Province, on undet. plant). Diagnosis: Dorsal shield 308 long, 163 wide. Measurements of setae: i 1-32, i s -30) i 4-8, i 5-8, j6-9, J 5.. 9, za-13, zs-32, z4-18, z5-8, Z4:-65, Z5-81, s4,-76, ss-72, r s-46, JV Macrosetae on leg IV: genu-19, tibia-52, basitarsus-32, distitarsus-32. South China Phytoseius (Phytoseius) rugatus Tseng PhytoseiuJ (Dubininellus) rugatus Tseng, J. Agr. Ass. China, 94: (Holotype : Particulars unavailable to the author). Diagnosis: literature. Not available to the author for want of original Taiwan Phytoseius (Phytoseius) ruidus Wei-nan & Zhao-Quan Phytoseius (Dubininellus) ruidus Wei-nan & Zhao-Quan, Acta Entoffl%gica Sinica,27(4) : 460. (Holotype ~,Paratype 1 ~, South China : Long Chuan, Yunnan Province; 1 ~ Da Ming Shan, Guangxi). Diagnosis: Dorsal shield 275 long, wide. Measurements of setae: jl-22-29, js-20-.5, i 4-6-7, i 5-6-7, js-7, J 5-6-8, za-20-22, zs , z , z5-6-7, 24:-72-82, r a , JV~ South China Phytoseius (Phytoseius) silvaticus Wei-nan & Zhao-Quan Phytoseius (Dubinillellus) silvaticus Wei-nan & Zhao-Qu3:D, Acta Entonl%gica Sinica, 27(4) : (Holotype ~, Paratypes 4 ~ ~, 200' South China: liu Hu Ping, Hubei, on Tectona grandis ; 2 ~ ~, Song Bai. Vitis adstricta).

114 100 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 Diagnosis: Dorsal shield long, wide. Measurements of setae: jl-27-30, ja-20-22, j4-7-l0, i ls -6-8, j6-8-l2., 1 1S -8-l0, ja-33-35, z4-l6-l7, z5-8-10, Z , Z , , , ra-37-40, JV Distribution; South China. Genus Typhlodromus Scheuten Typhlodromus Scheuten, Arch. Naturges, 23: Seius (Seiulus) Berlese, Acari Myriapoda et Scorpionides, Fasc. 41(3) : Seiulus Oudemans, Tij. Ent., 4S: Iphidulus Ribaga, Riv. Pat. Veg., 10 : Echinoseius Ribaga, Riv. Pat. Veg., 10: Neoseiulus Hughes, In. Min. Agr. Fish. Lond. Pub., T'yphlodromus (Typhlodromus) Chant, Can. Ent., 89 : Typhloseiopsis De Leon, Ent. News., 70: Anthoseius De Leon, Ent. News., 70 : Typhlodromidae Karg, Z. angew Ent., 47 : Cydnodromella Muma, Bull. Fla. St. Mus., 5: Amblydrome/la Muma, ibid., p Paraseiulella Muma, ibid., Meta~'eiulus Muma, ibid., Neoseiulella Muma, ibid., Austaliseiulus Muma, ibid., Clavidromus Muma, ibid., Clavidromina Muma, ibid., Amblydromus Muma, ibid., \. Typhlodromina, Muma, ibid., Galendromimus Muma, ibid., Galendromus Muma, ibid., Typhlodromella Muma, ibid., Typhloctonus Muma, ibid., Paraseiulus Muma, ibid., Typhlodromini Wainstein, Acar%gia, 4 : Chanteius Wainstein, Acarologia,.4 : Melodromus Wainstein, Acar%gia, 4 : Chilise;us Gonzalez & Schuster, Bull. Univ. Chile Agr. Exp. Sta., 16: Cydnoseius Muma, Fla. Ent., 50 : Typhlodromus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Key to the subgenera of Typblodromus 1. Dorsai shield' with 11 pairs of setae Doral shield with less than 11 pairs of setae 2. Dorsal shield with 10 pairs of setae Dorsal shieid with less than 10 pairs of setae Typhloctonu& 2 4 3

115 GUPTA: ORIENTAL PHYTOSEIIDAE 3. Sternal shield entire and distinct Sternal shield not entire and or slightly indistinct pairs of median setae 2 pairs of median setae S. Ventrianal shield with 3 pairs of preanal setae Ventrianal shield with 4 pairs of preanal setae 6. Most of the lateral setae plumose Most of the lateral setae not plumose, only a few may be serrate 7. Z5 and Z.. serrate and former knobbed Not as above 8. Leg IV with macrosetae on genu, tibia and basitarsus of leg IV Leg IV with macrosetae on basitarsus IV only 101 Brethria Metaseiulus Paraseiulus 5 Anthoseius 6 Clavidromus 7 Amblydromella 8 Orientiseius Typhlodromus Subgenus Amblydromella Key to the species of Amblydromella 1. Some setae on dorsal shield knobbed None of the setae on dorsal shield knobbed 2. Macroseta present only on basi tarsus IV Macroseta present on other leg segments besides basitarsus IV 3. s.. longer than z.. s.. as long as z.. 4. Length 'f setae jl-j6' J2 vary between jl-j6' J2 much longer than S. Macroseta present on genu, tibia and basitarsus of leg IV Macroseta present on genu and basitarsus of leg IV Ventrianal shield much longer than wide Ventrianal shield only slightly longer than wide 7. Z2 reaches almost upto base of Zs Z2 much longer than the distance between its base and that of Zs 8. Pores absent around ventrianal shield Pores present around ventrianal shield 9. Macroseta on genu IV and tibia IV equal Macroseta on genu IV and tibia IV unequal 10. Macroseta on genu IV longer than that of tibia IV.. Macroseta on genu IV shorter than that of tibia IV... 1'. Macroseta on genu IV much, shorter than that of tibia IV Macroseta on genu IV only slightly shorter than that of tibia IV 12. Spermatheca corniform Spermatheca saculiform holnalii himalayensis dactyli/era 6 nlori serrulatus fleschneri umbratus gaillmmatus 10 philippinus eharai batllbusicolus

116 102 REC. ZOOL. SURV. INDIA, OCC. PAPER No Z" almost double the length of S6 Z" only slightly longer than S~ 14. Setae comparatively longer, Z" extends beyond the base ofz/s Setae comparatively shorter, Z" never reaches upto the base of Z Cervix of spermatheca funnel-shaped Cervix of spermatheca sacular 16. Macroseta present only on basitarsus IV Macrosetae present on genu, tibia and basitarsus IV 17. Ventrianal shield with concave lateral margins' Ventrianal shield with straight lateral margins 18. Macrosetae simple Macrosetae knobbed 19. Dorsal shield reticulate Dorsal shield partly reticulate 20. Z" never reaches upto S6 Z" reaches upto S~ 21. Ss and S4 equal Ss and S" unequal 22. Ss shorter than S" Ss longer than 8" 23. r a and R 1 almost equal r I longer than R S" longer than S 45 S" as long as Movable digit with 3 teeth Movable digit without teeth... ryu/cyuensis eremetides IS 16 gopali divergentis acacia zalar; octavus, cordia rhenanus 22 rhododendron; balakotiensis sica loralaiana 25 ghani; kodaikanalensis chrysanthemi 204. Typhlodromus (Amblydromella) acacia ] ie-liu, Lai-rong & Li-Sheng Typhlodromus acacia Jie-liu, Lai-rong & Li-sheng, Fudan Journal (N. S.) 19 (4) : (Holotype ~, South China: Fujian Fuzhou, on Acacia richu; repository unknown). Diagnosis: Dorsal shield reticulate with 18 pairs of setae, Z5 thick and serrate, others smooth, a conspicuous pore present associated with SG. Measurements of setae: i 1-13, j4-13, l&-11, j's-17, J~-17, J 5-8, z&-19, is-1s. ja-13, za-13, z4-13, 84-15, 80s-17, S2-19, S5-17, Z5-S3, ra-17, R l -15. Ventrianal shield with concave lateral margihs and 4 pairs of preanal setae. Knobbed macrosetaeo preseilt i;h genu, tibia, and basitarsus o( Leg IV. South China.

117 GUPTA: ORIENTAL PHYTOSEllOAE Typhlodromus (Amblydromella) balakotiensis (Chaudhri, Akbar & Rasool)-New Comb AmblydromeJla balakotiensis Chaudhri, Akbar & Rasool, Rept. PL 480 Project, UAL, p (Holotype ~, Pakistan, 1 mile East of Balakot, at 300', on Salvia sp., in UAL). Diagnosis: Dorsal shield pear-shaped, 326 long, 194 wide, concave near R 1, with 18 pairs of setae and a pair of pores. All setae shorter than distance next in line, Z4>Z4 - Z5. Measurements of setae: il-16, i4-13, j5' j6-16 each, J a -18, J 5-8, js-21, za=s6-16, Z3 -Z4' each Sa-26, S4-23, S5-13, Z5-42, z5-18, Z4-29, rs-21, R Ventrianal shield 104 long, 81 wide, pentagonal, lateral margins concave, with 4 pairs of setae. Spermatheca with cervix sacular. Macroseta on leg IV: basitarsus-29, simple. Dis tribution: Pakistan Typhlodromus (Amblydromella) bambusicolus Gupta Typhlodromus bambusicolus Gupta, Indian J. Acar., 2 : 2-4. (Holotype ~, India: Assam, Mikir hills, on bamboo, in ZSI. Paratype 1 ~, Assam, Nowgong, on citrus, in ZSn Typhlodromus (Amblydromella) bambusicolus, Gupta, Fauna of India (Acari: Mesostigrnata) Family Phytoseildae, p Diagnosis: Dorsal shield well sclerotized, reticulate, 290 long, lto wide, with 18 pairs of setae. Measurements of setae: jl-12, ;4 -j each, js-16, J a-20, J 5-8, js-15, za-12, Z s-12, Z4-16, , 58-20, 8a-20, , S5-12, Z5-49 (serrate), z5-14, Z4-24 (serrate). Ventrianal shield pentagonal, 76 long, 72 wide, with 3 pairs of preanal setae and a pair of preanal pores. 8permatheca with broad, tubular cervix. Macrosetae on leg IV: genu-8, tibia-12, basitarsus-20, all with bulging tip. DistributiDn; India (Assam) Typhlodromus (Amblydromella) cordiae (Murna) Cydnoseius cordiae Murna, Fla. En!., 50 : 276. (HolotYl'e ~, Pakistan: Karachi, on Cordia obliqua, in USNM. Paratypes 5 ~ ~, Hyderabad, same habitat; 10 ~ ~, 30' (j, Multan, on Mangi/era indica; 1 ~, 20' 0', 1 nymph paratypes, Tando Jam, Pongamia glabra ; Paratypes both in USNM and author's Collection).

118 104 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 Diagnosis: Dorsal shield 370 long, 190 wide, moderately sclerotized, imbricate, with 18 pairs of setae, Z5 long and serrate; other setae short. Fixed digit of chelicera with 4 teeth, movable digit with one tooth. * Peritreme extends anteriorly upto jl. Chaetotaxy c;lf genu II 2-8 1, genu III 1 a a 1. Spermatophoral process with short shank, very short foot, an indistinct lateral process and a truncate toe. Pakistan Typhlodromus (Amblydromella) chrysantbemi Gupta Typhlodromus chrysanthemi Gupta, Indian J. Acar., 1 : (Holotype ~,India: Gujarat, Jamnagar, on chrysanthemum, in ZSI) Typhlodromus (Amblydromella) chrysanthemi, Gupta, Fauna of India (Acari: Mesostigmata, Family Phytoseiidae, p Diagnosis: Dorsal shield reticulate, 312 long, 200 wide, with 18 pairs of setae, mostly thin and small exept Z4: and Z5' which are thick and serrate. Measurements of setae: i l -22, i4: -20, i 5-20, je-20, J a-25, J &-8, js-18, za-20, zs-20, Z4: -22, S4: -26, s6-28, SI-29, S ~-32, S5-18, Z5-45, Z5-20, Z4:-33. Ventrianal shield 108 long, 80 wide, lateral margins concave at preanal region, with 4 pairs of setae and a pair of pores. Spermatheca with an elongate cervix. Macroseta on basi tarsus IV -28. India (Gujarat) Typhlodromus (Amblydromella) dactylifera (Chaudhri, Akbar & Rasool)-New Comb Amblydromella dactyli/era Chaudhri, Akbar & Rasool, Rept. PL. 480 Project, UAL, p (Holotype ~,Pakistan, 15 miles north of Karachi, on Phoenix dactyli/era, in UAL.). Diagnosis: Dorsal shield reticulate, 326 long, 224 wide, concave near Rl with 18 pairs of setae and 4 pairs of pores. Setae js-zs longer than distance between their respective bases and those of the next setae; other setae shorter; Z4: shorter than the distance between the bases of Z4: and Z5. Measurements of setae: jl-26, j4-j5-21 each, j8-31, 1a-26, , , za-23, zs-29, z4:-31, s4-31, s6-36, 5a ==S4-39, SIS-42, Z&-49, z&-23, Z4-39, rs-34, R l -42. Metasternal plates present.

119 GUPTA: ORIBNTAL PHYTOSBIIDAE 105 Ventrianal shield 11 7 long, 81 wide, with 4 pairs of preanal setae. Macroseta on basitarsus IV -26 with bulbous tip. Pakistan Typhlodromus (AmhlydromelJa) divergentis (Chaudhri, Akbar & Rasool)-New Comb Arnblydromella divergenlis Chaudhri, Akbar & Rasool, Rept. PL 480 Project, UAL. pp (Holotype ~, Pakistan: Charrapani, 4000' Murree Hills, on Grewia sp., in UAL) Typhlodromus divergentis, Dhooria, Acar. Newsl., 10 : 4-6. Diagnosis: Dorsal shield partly reticulate, 316 long, 163 wide, concave near R 1, with 18 pairs of setae; setae i s -z 4 longer than distance between setae next in line, S4 extends beyond S~, Z4>Z..-S6. Measurements of setae: ji-18, j4-23, j:s-21, j 6-39, J a-47, J 5-10, js=zs-29, za-23, z4-31, s4-39, 56-52, Sa S4-55, , Z5-51, zr;-26, Z4-60, r s -26, R l -31. Metasternal plates present. Ventrianal shield 107 long, 65 wide, pentagonal, concave laterally, rounded anteriorly, with 4 pairs of preanal setae. Fixed digit of chelicera with 4 subapical teeth, movable digit with 1 tooth. Macroseta on leg IV: basitarsus-30. Pakistan, India (Punjab). Other collection localities: Pakistan: Alipur, Bhera,' Chakwal, Choa Saiden Shah, Haripur, Havailian, Islamabad, Jauharabad, Karachi,.Lyallpur, Mian Channu, Multan, Mazaffargarh, Nankana, Nia Lahore, Pirmahal, Rahim Yar Khan, Rajanpur, Rawalpindi, Salooni Jhal, Samasata, Sheikhupura. Other plants: Vernonia sp., Acacia arabica, Erlobotrya japonica, Luffa acutangula, Helianthus annuus, Malvastrum sp., Cupressus sempervirens, Acacia modesta, Withania somnifera, Nerium indicum, Salvadora oleoides, Jacaranda sp., Mangifera indica, Olea europea, Punica granatum, Grewia asiatica, Cestrum noctrum, Thuja o rien talis, Sesbania sesban, Chrysanthemum sp., Cordia obliqua Typhlodromus (Amhlydromella) darjeelingensis Gupta Typhlodromus insu/aris, Gupta, Sci. & Cu/t., 36: 98 (West Bengal: SYZygiU111 cumini). 14

120 106 RBC. ZOOL. SURV. INDIA, OCC. PAPER No Typhlodromus caudigians, Gupta et al., Sci. & Cult., 37 : 298. (Punjab: mango) Typhlodromus eharai, Gupta, Bull. zool. Surv. India, 1 : 54. (Karnataka: Cassia; Tamil Nadu: Oothu-coffee) Typhlodromus eharai Gupta, Entomologists' mon. Mag. 115: (Holotype ~,India: West Bengal, Darjeeling Dist" Lataguri, on an undet. plant and Paratypes 2 ~ ~, same data as for holotype, all in ZSI) Typhlodromus darjeelingensis- Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shieid 292 long, 185 wide, reticulate, with 18 pairs of setae. Setae Z!5 and Z4 serrate, former being knobbed. Measurements of setae: jl-10, j4-ja-11 each, j6-1s, J a -18, J 5-9, js-14o, za-12, za-13, Z4-15, s4-17, s6-20, Sa-22, 84-22, , Z!5-40 (knobbed), zls-12, Z4:-2I. Ventrianal shield slightly longer than wide, with 4 pairs of preanal setae and a pair of round pores. Leg IV with macrosetae on genu-b, tibia-is, basitarsus-29. India (West Bengal, Punjab). Remarks: The species T. (A.) eharai Gupta was re-named as T. (A.) darjeelingensis Gupta, Typhlodromus (Amblydromella) eremitidis (Chaudhri, Akbar & Rasool)-New Comb Alnblydromella eremitidis Chaudhri, Akbar & Rasool, Rept. PL 480 Project, UAL, pp (Holotype ~,Pakistan: 2 miles west of Pirmahal, on Tamarix aphylla, in UAL). Diagnosis: Dorsal shield reticulate, 316 long, 173 wide, concave near RI with 18 pairs of setae and 8 pairs of visible pores; Z4 longer than distance between Z4 and Z!5' Measurenlents of setae: j1-1s, j4-16, j15 = j6-18, each J a -2I, J 1S-8, js, Zs to S4 equal -23 each, zs-18. S8 = 84 = 8 1S-29 each, Sa-3I, Z15-49, z!5-16, Z4-39, rs = R I -23. Ventrianal shield with 4 pairs of preanal setae. Macrosetae on leg IV: genu-22, tibia-26, basitarsus-42, all with bulbous tip. Metastemal plates with setae present. 8permatheca sacular. Pakistan.

121 GUPTA: ORIENTAL PHYTOSElIDAE Typhlodromus (Amhlydromella) flescbneri Chant Typhlodronzus (Typhlodromus) /leschneri Chant, Can. Ent., 92 : (Holoty pe ~,Paratypes 2 ~ ~, 1 is, India : Karnataka, Mercara, on orange, in Canadian National Collection, Belleville. Paratypes 3 ~ ~, Megbalaya, Shillong, on citrus; 1 ~,Assam, Burnihat on citrus; 1 ~, Bangalore, on lemon, all in Citrus Expt. Sta., Riverside, California) Typhlodromus fleschneri, Gupta, Sci. & Cult., 36: 98. (West Bengal: on cotton) Typhlodromus fleschneri, Gupta, Indian J. Acar., 2 : 6. (West Bengal: Gossypium herbaceum) Typhlodromus (Amblydromel/a) /leschneri, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 338 long, 195 wide, reticulate, with 18 pairs of setae. Measurements of setae: i 1 -i 6, 20 each, J 2-25, J 5-10, js' Z9-Z4 24 each, S4 -s6-32 each, S2-32, S4-36, S5-16, Z5-44 ; Z5 and Z4 with bulbous tip. Sternal, metasternal and genital shields normal. Ventrianal shield 100 long, 63 wide, with 4 pairs of preanal setae and a pair of preanal pores. Macroseta on basitarsus IV-27 long. Chant (1960) made a brief mentioning of its characters. India (Assam, West Bengal, Meghalaya, Karnataka) Typhlodromus (Amhlydromella) galummatus (Chaudhuri, Akbar & Rasool)-New Comb Amblydromella ga!ummatus Chaudhri, Akbar & Rasool, Rept. PL 480 Project, UAL, p (Holotype ~,Pakistan : altitude Murree Hills, on undet. plant, in UAL). Diagnosis: Dorsal shield reticulate, 306 long, 204 wide, concave near R 1 Dorsal shield with 18 pairs of setae and 3 pairs of pores; Z4 shorter than distance between Z4 -S 1$. Measurements of setae: jl-13, i4 = i5-10, j6 = J a-16, J 5-10, ja-18, za-13, za = z.. = S4-16 each, 56-18, S2=S4-21 each, S5-19 Z5-31, z5-10, Z4-18, ra-18, R Ventrianal shield 99 long, 83 wide with 5 pairs of preanal setae. Cervix of 8permatheca tubular. Macrosetae on leg IV: genu-16, tibia-16, basitar5us-31, all with bulbous tip. Pakistan.

122 108 REC. ZOOL. SURV. INDIA, OCC. PAPER No Typhlodromus (AmblydromeIla) ghanii (Muma) Amblydromella ghanii Muma, Fla. Ent., 50 : 279. (Holotype ~, other & and ~ Paratypes, Pakistan: Multan, on Punica granatllm; 0' Paratype from Sialkot, on Zizyphus sp. Holotype in USNM, Paratypes in USNM and in author's Collection). Diagnosis: Dorsal shield 330 long, 190 wide, imbricate, entirely (indistinct behind i 6 ), with 18 pairs setae; Z4 and Z5 thick and serrate, the latter knobbed. Ventrianal shield pentagonal * with 4 pairs of setae. Spermatheca sac-like, chaetotaxy of genu II 2 ~ 1, genu III 1 && 1 Macroseta on basitarsus IV knobbed. Pakistan Typhlodromus (AmblydromeIla) gopali Gupta Typhlodromus gopali Gupta, Sci. & Cult., 35 : 277. (Holotype ~, Paratypes 3 ~ ~, India: West Bengal, Calcutta, on palm tree, all in ZSI) Typhlodromus gopa/i, Gupta & Dhooria, Abslr. Indian Sci. Congr., 1974, p.69 (PuDjab & Himachal Pradesh) Typhlodromus gopali, Gupta, Indian J. Acar., 1 : 12. (Rajasthan: palm) Typhlodromus (Alnblydrolnella) gopali, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 298 long, 192 wide, reticulate with 18 pairs of setae. Measurements of setae: jl-19, j4-22, j5-22, i6-31-3t, J 2-40, J 5-9, js-27, za-18, zs-24, z427, s4-36, s6-45, Sa-52, S4-52, S5-6, Z5-56, z&-22, Z Ventrianal shield 100 long, 63 wjde, with 4 pairs of preanal setae and a pair of preanal pores. Spermatheca with funnel-shaped cervix. Macroseta on basitarsus-iv : 27 long. Pradesh.) India (West Bengal, Rajasthan, Punjab and Himachal 217. Typhlodromus (AmblydromeIla) himalayensis Gupta Typhlodrol1llls hinlalayensis Gupta, Indian J. Acar., S: (Holotype ~, India: Himachal Pradesh, Kangra, on pear. Paratype 1 i i same data as for holotype, both in ZSI) Typhlodronlus (Amblydromella) h imala... ell sis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoselidae, p

123 GUPTA: ORIENTAL PHYTOSEI1DAE 109 Dipgnosis: Dorsal shield 298 long, 192 wide, reticulate, with 18 pairs 'of setae. Mea6urements of setae: jl-12, j4-10, j5-12, i s -16, 1 a-16 J e-8, js-14, za-18, zs-15, Z4-16, S4-20, ss-24, S2-24, S4-25, S5-17, (serrate, knobbed), z5-16, Z4-24. Ventrianal shield 100 long, 68 wide, with 4 paits of preanal setae. Macroseta on basitarsus IV-36 long (knobbed). Spermatheca with bell-shaped cervix. India (Himachal Pradesh) Typhlodromus (Amblydromella) homalii Gupta Typhlodromus homalii Gupta, 1970, Oriental. Ins., 4: (Holotype ~, Paratypes 1 ~, 10, India: West Bengal, Sibpur Botanical Garden, on Homalium tomentosum, in ZSI) Typhlodromus homalii, Gupta & Dhooria, Absrr. Indian Sci. Congr. J 1974, p Amblydromella plebius, Chaudhri, Akbur & Rasool, Rept. P L 480 project, VAL, pp (Pakistan: Xanthium strumarium) Typhlodromus homalii, Gupta, 1977, Indian J. Acar., 1 : 12. (Punjab: gall of Z,zyphus sp.) Typhlodromus homalii, Gupta, Indian J. Acar., 2 : 6. (Orissa: Ficus bengaiellsis, Meghalaya : arum, citrus) Typhlodromus homalii, Gupta, Bull. zool. Surv. India., 1: (Kerala : Vndet. plant, Tamil Nadu : Undet. plant) Typhlodromus homalii, Gupta & Nahar, In Contrib. to Acar. in India, p. 10 (Bihar: Citrus). 198~. Typhlodromus (Anlb~ydromella) homolii, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 325 long, 188 wide, reticulate with 18 pairs of setae; Z4 and Z5 serrate and the latter with bulging tip. Measurements of setae: jl-js each, 1s-22, 1 ls-9, is, ZS-Z each, S4' Ss, Sa-S each, SlS-lS, 2 ls-45, zls-18, Ventrianal shield pentagonal, lateral sides concave, 103 long, 75 wide, with 4 pairs of short preanal setae. Macroseta on leg IV: basitarsus-15 (bulbous tip). Gupta (1970) illustrated its spermatophoral process. India (West Bengal, Bihar, Kerala, Punjab, Meghalaya, Orissa, Tamil Nadu, Himachal Pradesh), Pakistan Typhlodromus (AmbJydromella) kodaikanalensis Gupta Typhlodromus kodaikanalensis Gupta, Bull. zool. Surv. India, 1 : (Holotype ~, India: Tamil Nadu, Kodaikanal, on pear. Paratypes 3 ~ ~, same data as for holotype, all in ZSI).

124 110 REC. ZOOL. SURV. INOlA, OCC. PAPER No Typhlodronlus (Amb~vdromella) kodaikana/ensis, Gupta, Fauna of India (Acari: Mesostign1ata) Family Phytoseiidae, p Diagnosis: Dorsal shield 300 long, 165, wide, reticulate, with 18 pairs of setae, Z5 thickest and serrate, others being smooth and simple. Measurements of setae: jl-21, j4-j6-16 each, J i' J 5-6 each, ja-20, za-20, zs-20, z4-20, s4,-24, s8-2r, Sa-28, S4-28, S5-16, Z5-48, z5-16, Z4-28. Ventrianal shield 92 long, 76 wide, with 4 pairs of preanal setae and a pair of preanal pores. Spermatheca with flask-shaped cervix. Macroseta only on basitarsus IV : 40 long. India (Tamil Nadu) Typhlodromus (Amblydromella) loralaiana (Muma) Amblydromella loralaiana Muma, Fla. Ellt., 50 : 578. (Holotype ~, Paratypes~ 0 and ~, 3 nymphs, 1 larva, Pakistan: Loralai, on Punica granatum; 2 ~ ~, Sialkot, on Zizyphus sp. ; 1 & Tando Jan, on Pongamia glabra. Holotype in USNM, Paratypes in USNM as well as in author's collection). Diagnosis: Dorsal shield 330 long, 200 wide, imbricate between dorsal and lateral setae, with 18 pairs of setae, 24 and Z5 long, serrate; other setae moderately long. Ventrianal shield longer than wide, lateral margins concave, with 4 pairs of setae. Chaetotaxy of genu II 2 a i 1, genu III 1 a i 1. Leg IV with macroseta only on basitarsus. Spermatophoral process with shank nearly twice as long as cheliceral fingers, slightly bifurcate at tip but with no foot, toe or lateral process. Pakistan Typblodromus (Amblydromella) mori Gupta Typhlodromus mor; Gupta, Indian J. Acar., 5 : (Holotype ~, India: Jammu & Kashmir, Udhampur, on mulberry. Paratypes 2 ~ ~, J. & K., Achhabal, on apple, all in ZSI) Typhlodromus (Amblydromella) mori, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 312 long, 172 wide, reticulate, with 18 pairs of setae. Measurements of setae: j 1-12, j4-j6-16 each, J a-20, J~-8, js-34, za-16, zs-20, z4-20, s4-24, 86-24, Sa, S4-27 each, S5-26, Z5-44, z5-16, , ra-24, R l -16. Ventrianal shield 100 long, 80 wide,

125 GUPTA: ORIBNTAL PHYTOSBllOAB 111 with 4 pairs of preanal setae. cervix. Macroseta on leg basitarsus-36 (knobbed). Spermatheca with wide flask-shaped IV: genu-l0 (Knobbed). tibia-16, India (J ammu & Kashmir) Typhlodromus (Amhlydromella) octavus (Chaudhri, Akbar & Rasool)-New Comb Amblydromella octavus Chaudhri, Akbar & Rasool, Rept. PL. 480 Project, UAL, p (Holotype C!, Pakistan: 1 mile south of Naran, 8500', Kaghan Valley, on Crotoneaster bacillaris, in UAL. Paratypes 11 S ~, same data and locality, on Cornus macrophyl/a, Dryopteris filiales, Polygonum sp., Rheum sp., Urtica dioica ; 3 ~ ~, NathiagaIi, on Youngia japonica ; 3 ~ ~, Madyan, on Cnicus sp., Dryop teris sp., exemplary deposited in USNM). Diagnosis: Dorsal shield reticulate, 306 long, 189 wide, concave near R 1, with 18 pairs of setae and 3 pairs of pores, setae Z2 = Z!.l-Za ; other setae shorter than distance to seta next in line; 24>24 -S5' Measurements of setae: jl-21, j4-10, i5 = j6-13 each, J 2-18, J 5-8, is = Zs = 21 each, za-18, Z4 = S4-23 each, s6-29, S2-34, S4-36, S5-34, Z5-57, z5-13, Z4-34, rs-27, R I -18. Metasternal setae on metasternal plates. Ventrianal shield 104 long, 88 wide, pentagonal, lateral margins concave, with 4 pairs of preanal setae and a pair of pores. Spermatheca sacular. Flxed digit of chelicera with 6 teeth, movable digit with 2 teeth. Macroseta on leg IV : basitarsus-44, simple. Pakistan Typhlodromus (Amhlydromella) philippinus Corpuz Typhlodromus philippinus Corpuz, Philipp. Agric., 50 : (Holotype ~, Philippines: Laguna Banos, College Campus, on Asparagus plumosus, in Dept. Ento. Univ. Philip. Paratypes 9 ~ ~, 1 d', same locality as for holotype, on Casuarina epuiseti!olia, in Dept. Ento. Univ., Philip.). Diagnosis; Dorsal shield notched at the level of R 1, laterally reticulate, with 18 pairs of setae, all pointed except Z5 with bulbous tip. Measurements of setae: jl-20, j4 = j5-12 each, i6 = J 2-20 each, zls-12, Z4-34, js-20, za=zs-14 each, Z4-20, S4=S6-24 each, S2=S4-24 each SIS-8, ZIS-48. Metasternal setae on triangular metasternal plates. Ventrianal shield longer than wide, constricted at mediolataral margin

126 112 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 with 4 pairs of preanal setae and a pair of pores. Fixed digit of chelicera with 3 teeth and pilus dentilis, movabl.e digit with 3 teeth. Leg IV with macrosetae on genu, tibia and basitarsus ; that on tibia is half of that on genu. Chaetotaxy similar as in female, 240 long, 120 wide, Spermatophoral process L-shaped. Philippines Typhlodromos (AmblydromeUa) rhenanus (Oudemans) Seiulus rhenanus Oudemans, Tij. Ent., 48 : 78. (Holotype-?, Germany: Bonn, on rotting leaves, in Rijksmus. van Nat. Hist. Leiden) Typhlodromus (Typhlodromus) rhenanus, Narayanan & Ghai, Proe. natn. Inst. Sci. India, 29B(S) : 539-S41. (New Delhi: on mango malformation) Typhlodromus rhenanus, Gupta, Ent. Ree., 86: Typhlodromus rhenanus, Gupta & Dhooria, Abs/r. lndian Sei. Congr., 1974, p. 69. (Punjab & Himachal Pradesh) Typhlodromus rhenanus, Gupta, Indian J. Acar., 1: 12. (Punjab: mango) Typhlodromus (Amblydromella) rhenanus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield imbricate with 18 pairs of setae. Ventrianal shield triangular with 4 pairs of preanal setae and a pair of preanal pores. Macroseta present only on basitarsus IV.? India (New Delhi, Punjab, Himachal Pradesh), Germany Typhlodromus (Amblydromella) rhododendroni Gupta Typhlodromlls rhododendron; Gupta, Bull. zool. Surv. India, 1(1) : SO. (Holotype ~, Paratypes 2 ~ ~, India: Karnataka, Gundelpet, on Rhododendron arboreum, in ZSI) Typhlodromus (Amblydromella) rhododendron;, Gupta, ~ auna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 272 long, 160 wide, reticulate, with 18 pairs of setae. Measurments of setae: jl-16, j4-16, j5-18, j6-20, J a-25, 10-8, i s -15, za-16, zs-20, z4-20, 84:-24,86-25, S2-26, S4-32, S5-44, Z5-44, zo-20, Z4:-36. Ventrianal shield 84 long, 60 wide, lateral margins concave, with 4 pairs of preanal setae. Macrosetae on leg IV-25 long. India (Karnataka).

127 GUPTA: ORIBNTAL PHYTOSElIDAE Typblodromus (Amblydromella) ryokyuensis Ehara Typhlodromus (Anthoseius) ryukyuensis Ehara, Mushi, 40 : (Holotype ~, Paratypes 7 ~ ~, 10, Japan: Okinawa lsi., on citrus, in Zoot Inst. Fac. Sci. Hokk. Univ.) Typhlodromus (Anthoseius) ryukyuensis, Ebara, Mushi, 43 : (Taiwan : Calamus margaritiae ; Reevesia formosane). Diagnosis: Dorsal shield with conspicuous imbrication, with 18 pairs of satae. Measurments of setae: i l -13, ;,,-11, irs-ii, i e -13, , J rs -8, j s -12, Z a -12, Z s -13, z 4-13, s , s 6-17, S 2-1 7, S 4-15, S rs -10, Z IS -38 (stout, strongly serrate). Ventrianal shield pentagonal, longer than wide with 4 pairs of preanal setae and a pair of small transverse pores. Macroseta on leg IV : genu-ii, tibia-13, basitarsus-15, mostly capitate or subcapitate. Ebara (1967) described it. Japan, Taiwan Typhlodromus (Amblydromella) serrulatus Ehara Typhlodromus (Anthoseius) serrulatus Ehara, Mushi, 46: (Holotype ~, Japan: Shiroyama, on Zelkova serratae. Paratypes 2 ~ ~, Hokkaido, Sapporo, Jugu/alls aitanthi/olia; 1 ~ Mt. Hyonosen, Totton, on Edgeworthia papyri/era; 1 ~, Mt. Hyonosen, on willow; 5 ~ ~, same data as for holotype, all in BioI. Inst. Fac. Ed. Tott. Univ.) Typhlodromu.s (Anthoseius) serrulatus, Ebara & Bhandhufalck, J. Fac. Ed. Tottori Univ. Nat. Sci., 17(2) : 45. (Thailand: Casuarina, Paederia /oetlda, Pinus kesiya). Diagnosis: Dorsal shield imbricate, with 18 pairs of setae and 5 pairs of pores. Measurements of setae: il-16, j4-13, j5-13, i 6-16, J 9-16, J 5-9, r s -18, R 1-15, i s -1?, za-15, zs-16, z4:-17, s419, se-21, , S&-15, Z IS -42, Z I) -15 I Z 4: -23. Metasternal setae on plates. V en trianal shield longer than wide, lateral margins concave, with 4 pairs of preanal setae and a pair of crescentic pores. Fixed digit of chelicera with teeth, movable digit with 3 teeth, genu II 2 a B- 1, genu III 1 i a 1. Leg IV with macrosetae on genu-12, tibia-14, basitarsus-26. Japan, Thailand Typhlodromus (A~blydromella) sica (Chaudhri, Akbar & Rasool)-New Comb Amblydromella sica Chaudhri, Akbar & Rasool, Rept. PL. 480 Project, UAL, p (Holotype ~, Pakistan: Charrapani, 4000', Murree Hills, Lespedeza juncea, in UAL. Paratype 1 ~, Mansehra, on Mentha longlfolia, repository not men tioned). 15

128 114 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 Diagnosis: Dorsal shield partly and weakly reticulate, 286 long, 153 wide, concave near R1 with 18 pairs of setae and 4 pairs of pores. Setae js-za and Za-Zs shorter than setae next in line. Measurements of setae: j1-16, i4 = i 5-16 each, i 6-26, J a-31, J f)-10, is-zs-23 each, za-16, z4-26, s4-31, s6-34, S2==S4-39 each, S5-8, Z5-44, z5-18, Z4-39, r s -18, R I -21. Metasternal setae present on plates. Ventrianal shield 91 long, 55 wide, pentagonal, with 4 pairs of setae. Spermatheca sacular. Macroseta on leg IV : basitarsus-23, pointed. Unknown.. Pakistan Typhlodromus (Amblydromella) umbratus (Chaudhri, Akbar & Rasool)-New Comb Anlblydromella umbratus Chaudhri, Akbar & Rasool, Rept. PL. 480 Project, UAL. p (Holotype ~, Pakistan: 3 miles north of Abbotabad, on Punica granatun, in UAL. Paratypes 3 ~ ~, same data as for holotype, on Conyza stricta, Cupressus semperverens, Prunus communis, exemplary deposited in USNM). Diagnosis: Dorsal shield reticulate, 316 long, 184 wide, concave near S4 and R 1, with 18 pairs of setae and 6 pairs of visible pores. Measurements of setae: j1-11, j4=i 5-16 each, i6-21, , J 5-10, js-18, zs-16, Zs = Z4-21 each, S4-23, s8-26, Sa = S4-29 each, S5-18, Z5-49 (serrate with bulbous tip), z5-18, Z4-34, (serrate, pointed). Ventrianal shield 107 long, 78 wide, with 4 pairs of preanal setae and a pair of pores. Metasternal plates present. Spermatheca sacular with umbrellalike process. Macrosetae on leg IV : genu-16, tibia-16, basitarsus-36, all with bulbous tip. Pakistan Typhlodromus (AmblydromeUa) zafari Chaudhti Typhlodromus zajari Chaudhri, Acarologia, 7(4) : (Holotype ~, Pakistan: Lyallpur, on mulberry, in USNM. Paratype 1 ~, same data as for holotype, in UAL). Diagnosis: Dorsal shield 343 long, 202 wide, reticulate, with 18 pairs of setae. Measurrments of setae: jl-20, j4-10, j5-13, j6-15, 1a- 19, J 5-10, ja-16, za-14, zs-16, z4-22, s4-24, s6-25, S2-29, , S5-27, Z5-56 (weakly serrate), z5-14, Z4-32. Ventrianal shield longer (103)

129 GUPTA: ORlBNTAL PHYTOSBUDAB 115 than wide (78) with 4 pairs of preanal setae; 2 pairs of meta podal plates. Leg IV with macrosetae on genu-20, tibia-20, basitarsus-34, all spatulate. Fixed digit of chelicera with 6 subapical teeth and pilus dentilis. Spermatheca with bell-shaped cervix and long neck. Pakistan. In addition to the above species, some more species are known but could not be included in key for the same reasons as mentioned for Amblyseius (Amb/yseius) spp. on page Typhlodromus (Amblydromella) arunachalensis Gupta Typhlodromus (Amblydromel/a) arunachalensis Gupta, Oriental Ins Typhlodromus (Amblydromella) arunachalensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield reticulate, 302 long, 146 wide with 18 pairs of setae. Measurements of setae: jl.-14, j4-9, j~-ii, j6-14, l a -15, J 6-4, ja-l~, za-10, zs-11, Z4-13, , s6-22, S2-22, S4-24, S5-12, Zf)-47, z5-13, Z4-34, r a, Rl.-13, each. Ventrianal shield 78 long, 78 wide, with 4 pairs of preanal setae. Macrosetae on leg IV : basitarsus-35. Gupta (1986)described it. India (Arunachal Pradesh) Typhlodromus (AmblydromeUa) husaini Khan Typhlodromus husaini Khan, Sci. Ind., 7 : (Holotype ~,Pakistan : on mango, repository unknown). Diagnosis: Due to non availability of original literature, no information in this regard can be given. Pakistan Typhlodromus (AmbJydromella) incasus (Chaudhri) Amblydromella incasus Chaudhri, Pak. J. Agr. Sci., 12(3-6) : 206. (Holotype ~, Pakistan: Lyallpur, on Datura innoxia, in UAF; other coljections : 3 ~ ~, Pakistan, Cheechawatni forest, on Aerila javanica; 1 ~, Rajana forest, on Achyranthes aspera). Diagnosis: Dorsal shield 344 long, 196 wide, with 18 pairs of setae and 3 pairs of pores. Mesurements of setae: il-i8, j..-18, j~-20, j6-22,

130 116 REC. ZOOL. SURV. INOlA, OCC. PAPER No ,1 6-8, is = z"a = 22 each.. zs:x: Z4 = 25 each, s4-27, s6-29, Sa-3D, S4-31, S&-21, Z&-47, z&-zo, Z4-31, rs-22, R l -24. Ventrianal shield with 4 pairs of preanal setae. Leg IV with macroseta on basitarsus with bulbo~s tip_ Not known. Pakistan Typhlodromus (Amblydromella) jackmickleyi Typhlodromusjackmickleyi, Oomen, In Veenman & Zonen, Wagenengen, p.89. Indonesia Typhlodromus (Amblydromella) libitus (Chaudhri) Amblydromella libitus Chaudhri, Pak. J. Agr. Sci., 12(3-4) : 212. (Holotype ~,Pakistan : UAF campus, on wheat, in UAF). Diagnosis; Dorsal shield 332 long, 160 wide with 18 pairs of setae, all setae thorn-like except Z5 and Z4 which are serrate. Measurem~nts of setae: jl-24, j4-19, j5-21, j6-34, J 2-39, , js-24, za-22, za-27, z4-34, s4-41, s8-44, Sa-51, S S5-8, Z6-58, zl)-22, Z4-59, r s -24, R l -27. Ventrianal shield 100 long, 60 wide. Macroseta on basitarsus IV-24. Remarks: Pakistan. Chaudhri et ale (1979) re-described it Typhlodromus (AmblydromeUa) limitatus (Chaudhri, Akbar & Rasool)-New Comb Amblydromella limitatus Chaudhri, Akbar & Rasool, Rept. PL 480 Project, UAF, p. 35. (Holotype ~,Pakistan : Bhalwal, on Gossypium hirsutum, in UAF). Diagnosis: Dorsal shield without reticulation, 260 long, 135 wide, with 18 pairs of setae; ZI) and Z4 with bulbous tip and serrate; others smooth. Measurements of setae: jl-1j, j4.-j6-13 each, j6-12, la-11, J 1)-10, is = Zs = Zs = z4z-13 each, , s8-35, S9-19, S4-23, SI)-13, Z'5"-31, zl)-13, Z4-31, r s' R l -18 each. Ventrianal shield not clear. Macrosetae on leg IV ; genu-10, tibia-12, basitarsus-30, all with bulbous tip. Pakistan.

131 GUPTA: ORIENTAL PHYTOSEIIDAE Typhlodromus (AmhlydromeUa) maracus (Chaudhri) Amblydromella maracus Chaudhri, Pak. J. Agr. Sci., 12(3-4) : 208. (Holotype ~,Pakistan : UAF campus, on mango, in UAF) Amblydromella maracus, Chaudhri, Akbar & Rasool, Rept. PL 480 Project, UAF, p Diagnosis: Dorsal shield 364 long, 200 wide, all dorsal setae simple except Ze and Z4 which are serrate; Z5 with bulbous tip. Measurements of setae: i I -20, i4 -h~-14 each, js-20, J 9-24, J 5-10, ja-24, zs-14, za'=z4=s4-24 each, so-27, Sa-S4-31 each, S(S-24, Z5-41, z(s-14' Z4-34, ra-20, R Ventrianal shield rectangular, 122 long, 75 wide, with 4 pairs of preanal setae. Macroseta on basitarsus IV -22 (bulbous). Pakistan Typhlodromus (AmblydromelIa) nidibu Pritchard & Baker Typhlodromus (Neoseiulus) nidibu Pritchard & Baker, Hilgardia, 33(7) : 221. (Holotype ~, Belgian Congo: Mulunga, on Australian pine, in USNM. Paratypes 9 ~ ~, Belgian Congo, Kysenyi, on Acacia sp. ; 5 ~ ~, Slanleyville, on Bridelia sp. ; 1 C, Mulunga, on Australian pine; 1 ~, Leopoldville, on raffia palm; 2 ~ ~, Leopoldville, on Combretum sp. ; 1 ~,Leopoldville, on tree; 1 ~ J Nioka, on Vernonia sp. ; 10 J Ruanda Urundi, on Acacia cyanophylla. all in USNM) Typhlodromus nidibu, Oomen, In Veenman & Zonen, Wageneng(n, p. 89. Diagnosis; Dorsal shield irregularly octiculate, with 18 pairs of setae. ProIa teral setae slightly bent, progressively increasing in length. Ventrianal shield with lateral margins concave. Spermatheca \vith elongate triangulate cervix; slender atrium. Genu IV with capitate macroseta ; tibia IV with also a capitate macroseta ; basitarsus IV with long capitate macroseta. It was described by Pritchard & Baker (1962). Africa (Belgian Congo), Indonesia Typblodromus (Amblydromella) nilgiriensis Gupta Typhlodromus (Amblydromel/a) nilgiriensis Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p (Holotype ~, India: Tamil Nadu, Top slip, on undet. tree, in ZSI). Diagnosis: Dorsal shield 355 long, 212 wide, smooth, well sclero F tized with 18 pairs of setae and 6 pairs of pores. Measurements of setae; il-22, i4-22, i 15-22, ;6-29, J 2-40, J 15-5, i 3-35, za-11, zs-44, z4-22,

132 118 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 S4-49, s6-49, Sa-50, S4-33, S5-29, Z5-78 (serrate), z5-26, Z4:-49 (serrate), r s, R l -2S-26. Ventrianal shield 112 long, 100 wide, with 4 pairs of preanal setae. Macrosetae on leg IV : genu-39, tibia-33, basitarsu.s-62, all knobbed. India (Tamil Nadu) Typhlodromus (Amblydromella) ornatulus (Chaudhri) Amblydromel/a ornatulus Chaudhri, Pak. J. Agr. Sci., 12(3-4) : 201. (Holotype ~, Pakiitan : UAF campus, on mango, in UAF). Diagnosis: Dorsal shield 306 long, 160 wide, deeply concave near Sa, with 18 pairs of setae; mostly slightly re-curved. Measurements of setae: jl-22, j4-j5-25 each, J a -37, J5-10, js-32, za-24, zs-31, z4-34, s4-57, s6-44, Saf"t55, S4-51, S5-10, Z~-51, z~-26, Z4-57, rs=rl=27 each. Spermathecal atrium hammer-like, cervix bell-shaped. Macroseta on basitarsus IV -31. Pakistan Typhlodromus (Amblydromella) pjatycladus Jie-liu, Lai-rong & Li-sheng Typhlodromus platycladus Jie-liu, Lai-rong & Li-sheng, Fudan J. (Nat. Sci.), 19(4): 472. (Holotype ~, South China: Jiangsu Zhenjiang, on Platyc:ladus orientaiis). Diagnosis: Dorsal shield with 18 pairs of setae. Measurements of setae: j1-19, j4-17, j~-17, j6-22, J a-24, J ~-11, js-24, za-19, zs-22, z4-24, s4-24, s6-28, Sa-28, S4-35, S~-30, ZD-48, z5-19, Z4-33, rs -19, R l -22, JV &-33.? South China Typhlodromus (Amblydromella) serratus (Chaudhri) Amblydromella serratus Chaudhri, Pak. J. Agr. Sci., 12(3-6) : 204. (Holotype ~, Pakistan: Dapher Forest Plantation, on Withallia somni/era, in UAF) Amblydronlella serratus, Chaudhri, Akbar & Rasool, Rept. PL 480 Project, UAF, p

133 OUPTA: ORIENTAL PHYTOSEIlDAE 119 Diagnosis: Dorsal shield 347 long, 210 wide, with 18 pairs of setae. All dorsal setae shorter than setae next in line. Measurements of setae: jl-18, j4-j5-10 each, js-13, J a -16, J 5-10, ja-za=13 each, z4=s4-16 each, S8 =S2-18 each, S4 = S5-29 each, , z5-10, , r.-13, R I -18. Ventrianal shield 120 long, 94 wide, with 4 pairs of preanal setae. Spermatheca sacular. Macroseta on basitarsus IV-23 (knobbed). Male; Pakistan Typhlodromus (Amhlydromella) sonprayagensis Gupta Typhlodromus (Amhlydromel/a) sonprayagensis Gupta, Entomon, 10(3) : (Holotype ~, Paratype 1 ~, India: Uttar Pradesh, Sonprayag, on pear, in ZSI) Typhlodromus (Amhlydrome/la) sonprayagensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield well sclerotized, reticulate, 360 long, 213 wide, with 18 pairs of setae. Measurements of setae: jl-14, j4-14, j5-16, J a-20, J 5-6, js-22, za-16 zs-20, Z4-18, S4-22, 5s-22, Sa-27, S4t -27, S&-20, Z5-45, z5-16, Z4-25, ra-22, R I -1S. Ventrianal shield 112 long, 85 wide, with 4 pairs of preanal setae and a pair of preanal pores. Macrosetae on leg IV: genu-14, tibia-16, basitarsus-35, all with bulbous tip. India (Uttar Pradesh). Subgenus Anthoseius De Leon 244. TyphIodromus (Anthoseius) majumderi Gupta Typhlodromus (Anthoseius) majumderi Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p (Holotype ~, Paratype 1 ~, India: West Bengal, Kalyani, on Heliolropilllll indicum, in ZSI). Diagnosis: Dorsal shield gently reticulate, deeply incised at the level of RI with 18 pairs of setae, all being smooth and simple. Measurements of setae: i I -10, i4-11, j5-12, j6-12, J a -12, J 5-9, ja-12, z2-10, zs-12, z4-15, s4-10, s6-14, Sa-15, S4-12, S5-14, Z5-16 (slightly thicker, smooth), Ventrianal shield much longer (123) than wide (90) with 3

134 120 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 95 pairs of preanal setae and a pair of preanal pores. on leg IV. Distribution; India (West Bengal). Macroseta absent Subgenus Bretbria Tuttle & Muma Key to the species of Brethria 1. Ventrianal shield very long, almost 2 times as long as wide, lateral margins straight Ventrianal shield only slightly longer than wide, lateral margins slightly concave roshanlali con/usus 245. Typhlodromus (Bretbria) confusus Narayanan, Kaur & Ghai Typhlodromus (Typhlodromus) con/usus Narayanan, Kaur & Ghai, Proc. natn. Inst. Sci. India,26B: (Holotype ~. India: Delhi, sunflower, in NPC) Typhlodromus (Brethria) con/usus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p ' 4. Diagnosis: Dorsal shield faintly reticulate, with 17 pairs of setae, all setae smooth except Z5 and Z4 which are thick and serrate. Setae is, Zs' Z4 almost equal; Sa and S5 almost equal; Za shorter. Ventrianal shield longer than broad. No macroseta on leg IV. India (Delhi) Typhlodromus (Brethria) roshanlali Narayanan & Ghai Typhlodromus (Typhlodromus) roshanlali Narayanan & Ghai, Proc. natn. Inst. Sci. India, 29 B(5): 539. (Holotype not designated; described from 4 ~ ~, 3d' d', collected from New Delhi, on malformed mango tree, in NPC) Typhlodromus (Brethria) roshaniali, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield reticulate with 17 pairs of setae, all being smooth. Measurements of setae: jl-24, j4, j5-20, i 6-32, Ja-44, J 5-8, is-16, za-23, zs-31, z4-31, 54:-33, s6-45, Sa-50, S4-42, Z5-50, z5-20, Z4-56 Ventrianal shield longer (124) than broad, with 4 pairs of preanal setae and a pair of pores. Macroseta absent on leg IV. Narayanan & Ghai (1964) described it. India (New Delhi).

135 GUPTA: ORIENTAL PHYTOSEIIDAE 121 Subgenus Calvidromus Muma 247. Typhlodromus (Clavidromus) neotransvaalensis Gupta Typhlodromus neotransvaalensis Gupta, Bull. zool. Surv. India, 1(1) : (Holotype ~, India: Tamil Nadu, Ooty, Casuarina. Paratype 1 ~, Aruvankadu, on tea, all in ZSI) Typhlodromus (Clavidromus) neotransvaalensis, Gupta, Fauna of India (Acari : Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield heavily sculptured, 328 long, 190 wide, with 18 pairs of setae, all being weakly serrate. Measurements of setae: i 1 -i6' 1 a each, l5-12., js, Za, Z4-18 each, S4: -20, s6-24, Sa-20, S4-24, S5-25, Z5-36, z5-18, Z4-28 Ventrianal shield 112 long, 92 wide, smooth with 4 pairs of preanal setae. Spermatheca with flask-shaped cervix. Macrosetae on leg IV : genu-16, tibia-20, basitarsus-26, all with flattened tip. ' India (Tamil Nadu). Subgenus Chiliseios Gonzalez & Schuster 248. Typhlodromus (Chilisetus) Iicini (Tseng) Chiliseius (Chiliseius) ficin; Tseng, J. Agr. Ass. China,94: Diagnosis: Due to non-availability of original literature, these particulars could not be provided~ Taiwan Typhlodromus (Chiliseius) separalus Wei-nan & Zhao-Quan Typhlodromus (Chi/iseius) separatus Wei-nan & Zhao-Quan, Acta Zootaxonomica Sinica, 10(4) : (Holotype ~, Paratypes 3 ~ ~, South China: Jinafenglang, Hainan lsi., on Rhus sp.). Diagnosis: Dorsal shield long, wide, with 18 pairs of setae. Measurements of setae: j , j4-5-7, j6-6-7, j6-7-o, la-8-to, J 6-5-6, ja-21-26, za-5, za-10, Z , S4-12, , Sa- lo. S4-10, S6-5, Z , z6-7, Z , ra, R1-7 each. 16 South China.

136 122 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 95' 250. Typhlodromus (Chiliseius) tengi Wei-nan & Zhao-Quan Typhlod"omus (Chiliseius) tengi Wei-nan & Zhao-Quan, Acta Zootaxonomica Sinica, 10(4) : (Holotype ~, Paratypes 2 ~ ~, South China: Jinafengling, Hainan lsi., Guan ~ dong Province, on Sophora sp.). Diagnosis: Dorsal shield long, wide. Measurements of setae: jl-5, j4-5-7, j5-5-7, j6-8, Ja-l0, J5-7-10, js-20-22, za-6-7, zs-lo, z4-10, 54-52, 56-10, Sa-10, S4-8-10, S5-7 -8, Z5-150, z5-11, 'Z , rs-7-8, JV Male; South China. Subgenus Metaseiulus Muma 251. Typbolodromus (Metaseiulus) co~tiguus Chant Typhlodomus (Typhlodromus) contiguus Chant, Can. EIlI., 91 : (Holotype ~, Hong Kong: Taipai, on citrus. in Canadian National Collection. Paratype-? in USNM and Citrus Expt. Sta., Riverside, California) Typhlodromlls (Typhlodromus) contiguus, Ebara, Mushi, 39 : 20. (Philippines). Diagnosis: Dorsal shield with 16 pairs of setae, 8 pairs in lateral rows. Setae ji' js' zs' 54 s6' SIS and Z4 long, other setae minute. Measurements of setae: jl-23, j4-5, j5-5, j6-8, J a-9, J 1S-7, rs-13, R 1-8, js-42, za-6, zs-59, Z4-7, 54-73, s6-69, Sa-9, Z5-130, z5-6, Z4-73 Ventrianal shield with 4 pairs of preanal setae. Macrosetae on leg IV : genu-29, tibia-31, basitarsus-35. Dorsal chaetotaxy similar as in female. Hong Kong, Japan, Philippines. Subgenus Orientiseius Muma & Denmark Key to the species of Orientiseius 1. Dorsal shield incised near Rl Dorsal shield not incised near R 1 2. Z2 longer than Z2-ZS Z2 shorter than Z2-Za 3. Macrosetae capitate Macrosetae simple 2 6 samiiensis 3 thailandicus 4

137 GUPTA: ORIENTAL PHYTOSEIIDAE 4. j61i times as long as ja ja nearly i or less than ~ of j6 S. Z4, slightly longer than Z 2 Z4, 2 times as long as Z2 6. Zs only 2 times as long as Z2 Zs over 4 times as long as Z2 7. Macroseta present only on basitarsus IV Macrosetae present on genu, tibia and basitarsus of leg IV 8. ja only slightly longer than j6 ja much longer, 2 times of j6 9. Z4, much shorter than z,-s,) Z4 reaches only upto the base of S6 Z, reaches upto base of S4:, Z, extends beyond the base of orissaellsi~ 5 rickeri hadii manipurensis 7 channabasavannai 8 pruni 9 lalazariensis agilis 252. Typhlodromus (Orientiseius) agilis (Chaudhri)-New Comb Orientiseius agilis Chaudhri, Pak. J. Zool., 7(2): (Holotype ~, Pakistan: 2 miles north of Havelian, on Rubus sp., in UAL). Diagnosis: Dorsal shield 316 long, 153 wide, smooth, without concavity near Rl with 18 pairs of setae and 5 pairs of pores. Measurements of setae: jl-23, i4-23, j5-21, js-29, J 2-36, J 5.. 5, ia-47, za-16, zs-53, z4-18, S4-60, s6-65, Sa-60, S4-36, S5-16, Z5-57, z5-5, rs-29 each, R l -18, Z4 extends beyond base of Z5. Ventrianal shield 104 long, 73 wide, with 4 pairs of preanal setae. Leg IV macrosetae on genu-34, tibia-26, basitarsus-52. Pakistan Typhlodromus (Orientiseius) channabasavannai Gupta Typhlodromus channabasavanni Gupta, Bull. zool. Surv. India, 1(1) : SO-52. (Holotype ~, India: Tamil Nadu) Aruvankadu, on tea, in ZSI) Typhlodromus (Olientiseius) channabasavannai, Gupta, Fauna of India (Acari : Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 300 long, 160 wide, with 18 pairs of setae, all being smooth and pointed. Measurements of setae: jl-16, j4-j~-8 each, i 6 -l a -18 each, 1 5-6, ja-36, za-8, zs-36, Z4-8, s4-43, s6-48, S2-48, S -32, S~-12, Z~-70, z5-8, Z4-56. Ventrianal shield much longer

138 124 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 (IOO) than wide (72) with 4 pairs of preanal setae. Spermatheca with funnel-shaped cervix. Macroseta on basitarsus IV-56 long. India (Tamil Nadu) Typblodromus (Orientiseius) hadii Chaudhri Typhlodromus hadii Chaudhri, Acar%gja, 7 : (Holotype ~, Pakistan, Behrain (Swat State), on fig. in UAL) Typhlodromus kashmiricus Gupta, Indian J. Acar., 5 : Typhlodromus (Orienliseius) hadii, Gupta, Fauna of India (Acari: Mesostgmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 343 long, 174 wide with 18 pairs of setae and 6 pairs of pores. Measurements of setae: jl-23, j4-6, i 5-8, i8-16, J a -16, J a -6, j~-43, Z2-8, zs-55, Z4-20, s4-55, s8-61, Sa-53, S4-43, Sli-35, Z5-74 (weakly serrate), z5-6, Z4.-59, ra-27, R l -31. Ventrianal shield 109 long, 74 wide, with 4 pairs of preanal setae. Leg IV with macrosetae on genu-39, tibia-35 and basitarsus-61. Fixed digit of chelicera with 2 subapical teeth. Macrosetae on leg IV: genu-24-30, tibia-32-35, basitarsus Pakistan Typblodromus (Orientiseius) lalazariensis (Chaudhri)-New Comb Orienti.'eius lalazariensis Chaudhri, Pakistan J. Zool., 7(2): (Holotype ~, Pakistan: Lalazar, 1000' (Kaghan Valley), on Abies pindrow, in UAL). Diagnosis: Dorsal shield 347 long, 194 wide, with 18 pairs of setae and 5 pairs of pores. Measurements of setae: jl-5, j4-5, jt5-10, j6-16, 1 a -26, J IS -5, js-42, za-13, zs-39, z4.-13, , s6-34, Sa-34, S,,-29, S5-16, 2&-55 (weakly serrate), zls-5, Z,,-36, (nearer to S4 than to SIS)' rs-26, R Ventrianal shield 117 long, 99 wide, with 4 pairs of preanal setae, concave medioanteriorly. Leg IV with macro setae on genu-26, tibia-36 and basitarsus-70. Distribution; Pakistan.

139 GUPTA: ORIENTAL PHYTOSEllDAE Typblodromus (Orientiseius) manipurensis Gupta Typhlodromus manipurensis Gupta, Indian J. Acar., 2: 1-2. (Holotype ~,India: Manipur, Mao, on undet. plant. Paratypes 3 ~ ~,same data as for holotype, all in ZSI) Typhlodromus (Orientiseius) manipurensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 288 long, 160 wide, with 18 pairs of 'setae, all being smooth and simple; Z5 longest 52, other setae measure: ii-20, ;4-8, j5-12, j6-16, J a -16, J 5-6, js-12, za-8, zs-18, z4-18, s4-24, s6-32, , S4-24, S5-20, z5-12, Z4-32. Ventrianal shield longer (88) than wide (69) with 4 pairs of preanal setae and a pair of preanal pores. Spermatheca with thick broad cervix. Macrosetae on leg IV : genu-25, tibia- 28, basitarsus-56. India (Manipur) Typhlodromus (Orientiseius) orisssaensis Gupta Typhlodromus orissaensis Gupta: Indian J. Acar., 2 : 4-6. (Holotype ~,India : Orissa, Angul, on undet. plant, in ZSI) Typhlodromus (Orientiseius) orissaensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 280 long, 148 wide, with 18 pairs of setae, all being smooth and slender. Measurements of setae: jl-24, j4-26, i 5-33, j6-48, J a -54, J 5-5, is-44, za-6, zs-53, z4-24, s.j.-53, s6-64, S2-62, , S5-16, Z5-64, z5-16, Z4, -56. Ventrianal shield 88 long, 76 wide, lateral margins deeply concave with 4 pairs of preanal setae. Spermatheca with thick long cervix. Macrosetae on leg IV : genu-lo, tibia-14, basitarsus-36. Distribution; India (Orissa) Typhlodromus (Orientiseius) pruni Gupta Typhlodromus pruni Gupta, Oriental Ins., 4(2): (Holotype S, Paratype 1 ~, India: West Bengal, Kalimpong, on PrUllliS persica, in ZSI) Typhlodromus (Orientiseius) pruni, Gupta, Fauna of India (Acari: Mesostig.. mata) Family Phytoseiidae, p Diagnosis: pairs of setae. Dorsal shield reticulate, 328 long, 170 wide, with 18 Measurements of setae: jl-27, j4-4, jo-5, j6-7, J 2-91,

140 126 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 J 5-4, js-36, z2-5, zs-4s, Z4-6, s4-45, s6-48, S2-27, S4-14, 51)-9, Z5-76, z5-5, Z4-36. Ventrianal shield 100 long, 72 wide, with 4 pairs of preanal setae and a pair of preanal pores. Macrosetae on leg IV : genu 40, tibia-27, basitarsus-46. India (West Bengal) Typhlodromus (Orientiseius) rickeri Chant Typhlodrolnus (Typhlodronzus) rickeri Chant, Can. Ent., 92 : (Holotype ~, Pa ratypes 3 ~ ~, 10, 2 Proto, 2 Deuto, India : Shillong, on citrus, in Canadian National Collection, Belleville. Paratypes 4 ~ ~, Proto, India: Bangalore, on citrus, in Citrus, Exp. Sta., Riverside. California) Typhlodromus rickeri, Gupta, Indian J. Acar., 2 : 6. (Nagaland : Undet. plant) Typhlodromus (Orientiseius) rickeri, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 280 long, 160 wide, with 18 pairs of setae. Measurements of setae: j1-20, j4-12, j5-24, j6.. 24, J 2-32, Jrs-6, js-44, z2-8, zs-52, Z4-10, S4-60, s6-66, , S4-36, S5-16, Z5-76, zrs-8, Z4-60. Ventrianal shield 92 long, 64 wide, pentagonal, with 4 pairs of preanal setae. Spermatheca with funnel-shaped cervix. Macroseta on leg IV: genu-32, tibia-28, basitarsus-48. Chant (1960) gave a brief description of male. India (Meghalaya, Karnataka, Nagaland), U. S. A. (introduced) Typhlodromus (Orientiseius) samliensis (Chaudhri)-New Comb Orientiseius safnliellsis Chaudhri, Pakistan J. Zool., 7(2): (Holotype ~, Pakistan: Samli, 4000' (Murree Hills), on Pru'}us persica, in UAL. Paratypes 1 ~, A bbotabad, on Zizyphus sp. ; 1 ~, Nathiagali, on Abies pindrow ; 2 ~ ~,Madyan, on Dryopleris sp. and Spiraea lindleyana; examplary deposited in USNM). Diagnosis: Dorsal shield 357 long, ~ 94 wide, deeply concave near R 1, with 18 pairs of setae and 8 pairs of pores. Measurements of setae: jl-5, i 4-5, it~-16, i 6 -l8, J 2-29, J 5-5, js-49, z2-13, zs 49, 24-13, 84-57, s6-60, Sa-55, S4-39, S5-31, Z5-75 (weakly serrate), Z5-S, Z4-51, (nearer to S4 than to S5)' r s-34, R l -29. Ventrianal shield 120 long, 86 wide, with 4 pairs of preanal setae, constricted laterally.

141 GUPTA: ORIENTAL PHYTOSEIIDAE 127 Metapodal plates 2 paired. Spermatheca with bell-shaped cervix. Chelicera with 4 teeth on fixed digit and 2 teeth on movable digit. Leg IV with macrosetae on genu-42, tibia-39, basitarsus-62. Pakistan Typhlodromus (Orientiseius) thailandicus Ehara & Bhandhufalck Typhlodromus (Anthoseius) thailalldicus Ehara & Bhandhufalck, J. Fac. Ed. Tottori Univ. Nat. Sci., 27(2) : (Holotype ~,Thailand: Mang, on persimmon. Paratypes 5 ~ ~, same data as for holotype; 1 ~, Fang, on citrus; 1 ~, Fang, on peach, all in BioI. lost. Fac. Ed., Tott. Univ). Diagnosis: Dorsal shield 340 long, 230 wide, reticulate laterally, with 5 pairs of pores and 18 pairs of setae. Measurements of setae: j1-22, j4-17, j5-19, j6-25, J , J 5-6, js-37, z2-16, zs-34, z4-32, s..-43, s8-50, S2-51, S4-31, S5-29, Z5-73, z5-19, Z4-53, rs-23, R1-18. Metasternal plate with seta present. Ventrianal shield pentagonal, longer than wide, transversely striated with 4 pairs of setae and a pair of pores. Genu II 2 g. g. 1, Genu III 1 a g 1. Macrosetae on leg IV : genu-42, tibia-27, basitarsus-47, all capitate. Thailand. Subgenus Paraseiulus Muma Key to the species of Paraseiulus 1. All dorsal setae serrate Dorsal setae not serrate 2. Dorsal shield deeply concave near So and Rl Dorsal shield deeply concave only near Rl 3. Macroseta absent on leg IV Macroseta present on basi tarsus IV 4. Distal tip of peritreme slightly recurved Distal tip of peritreme not recurved 5. Ventrianal shield much narrower than width at the anal level Anterior margin of ventrianal shield as wide as anal region COlnptlls 2 vitreus operantis neoso/eiger trinlediosetus igllavlls

142 128 REC. ZOOL. SURV. INDIA, Occ. PAPER No Typblodromus (Paraseiulus) comptus Corpuz Typhlodromus eonlptus Corpuz, Philipp. Agrie., 50 : (Holotype ~, Paratype 1 ~, Philippines : Laguna, Los Banos, College campus, on Casllorina equisetifolia, in Dept. Ento., Univ. Philip.). Diagnosis: Dorsal shield slender, heavily imbricate, 320 long, 168 wide, with 18 pairs of setae; r s also on dorsal shield. All the setae except J 5 short, thickened and pilose. Sternal shield with 2 pairs of setae; 3rd pair on membrane, 4th pair on metasternal plates. Ventrianal shield triangular, slightly longer than wide, fine reticulations; with 4 pairs of preanal setae. Fixed digit of chelicera with 4 teeth and a pilus dentilis, movable digit with 1 tooth. Philippines Typblodromus (Paraseiulus) ignavus (Chaudhri, Akbar & Rasool)-New Comb Paraseiulus ignavus Chaudhri, Akbar & Rasool, Rept. PL 480 Project, UAL, p (Holotype ~, Pakistan: 2 miles north of Ughi, on Prunus armen;ca, in UAL). Diagnosis: Dorsal shield, reticulate, 357 long, 224 wide, deeply concave near Rl with 19 pairs of setae; setae shorter than distance of setae next in line. Measurements of setae: ji-10, j4 -j5-13 each, je-16, J a-1s, J 5-10, z5 = ze-13 each, Z" -23, is = Za = 84 = each, za-13, z4-10, Sa21, S4-26, S5-23, Z5-36, rs-21, R l -16. Metasternal setae on plates. Ventrianal shield with deeply concave lateral margins, with 4 pairs or preanal setae and a pair of pores. Leg IV with no macr08eta. Pakistan Typhlodromus (Paraseiulus) neosoleiger Gupta Typhlodromus (Paraseiulus) neosoleiger Gupta, Indian J. Acar., 5 : (Holotype ~,India: Jammu & Kashmir, Srinagar, on chrysanthemum. in ZSI) Typhlodromus (Paraseiu/us) neosoieiger, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield reticulate, 240 long, 210 wide, with 19 pairs of setae. Measurements of setae: i l - 23, j4-14, j5-16, i6-16, J I -16, J 5-z6, is-20, z2-16, zs-18, Z4-20, S4-20, 86-20, Sa-20, , S~-20, Z~-36,

143 GUPTA: ORIENTAL PHYTOSEIlDAE 129 Z&-36, zls-16, z6-16, Z4-24, rs 24, R l -20. Ventrianal shield 116 long, 68 wide, with 4 pairs of preanal setae. Macrosetae on leg IV: basitarsus-24. India (Jammu & Kashmir) Typhlodromus (Paraseiulus) operantis (Chaudhri, Akbar & Rasool)-New Comb Paraseiulus operantis Chaudhri, Akbar & Rasool, Rept. PL. 480 Project, VAL, p.228. (Holotype ~, Pakistan: Samli, 4000' (Murree Hills), on Desmodium sp., in UAL). Diagnosis: Dorsal shield weakly reticulate, 408 long, 234 wide, deeply concave near R 1, with 19 pairs of setae. All setae shorter than setae next in line. Measurements of setae: jl-16, j4 = j&-10 each, ;6-13, J~-16, J 5-10, zrs- 13, z6-16, Z4-21, js = za = zs-13 each, z4-16, S4.==56-21 each, Sa-26, S4-29, Srs-31, Z5-42, ra=rl-18 each. Metasternal setae on metasternal plates. Ventrianal shield vaseshaped, concave near 3rd and 4th preanal setae, with 4 pairs of preanal setae and a pair of pores. Macrosetae on leg IV : basitarsus-29. Pakistan Typhlodromus (Paraseiulus) trimediosetus lie-liu, Lai-rong & Li-sheng Typhlodromus trimediosetus Jie-liu, Lai-rong & Li-sheng, Fudan J. (Nat. Sci.), 19(4) : (Holotype ~, South China: Shangha, Jiangwan, on Humulus scandens, 2 Paratype ~ ~, same data as for holotype; 1 ~, Jingsu on Plerncary stenoptera ; Repository-?). Diagnosis: Dorsal shield reticulate, with 19 pairs of setae. Measurements o~ setae: j1-10, j4-11, ;5-11, j6-14, la-15, 15-7, zf)-10, z6-14, Z4-24, ;3-17, z2-18, zs-18, z4-19, s4,-19, s6-21, Si-21, S4-21, SIS-21, Z5-26 (serrate), r a -l?, R l -16. Sternal shield with 2 pairs of setae, 3rd pair on interscutal membrane, 4th pair on platelets. Ventrianal shield elongate, lateral margins deeply concave with 2 pairs of preanal setae.? 17 South China.

144 130 REC. ZOOL. SURV. INDIA, OCC. PAPER No Typhlodromus (Paraseiulus) vitreus (Chaudhri, Akbar & Rasool)-New Comb Paraseiulus vitreus Chaudhri, Akbar & Rasool, Rept. PL. 480 Project, UA~., p (Holotype ~, Pakistan : Barian, 6000' (Murree Hills) on Daphne popyracea, in UAL. Paratype 1 ~, Sunny Bank, 7500' (Murree Hills), on Cornus sp., (repository not mentioned). Diagnosis: Dorsal shield reticulate, 357 long, 214 wide, deeply concave near S6 and R1 with 17 pairs of setae, all being shorter than the setae next in line. Measurements of setae: j1-16, j4 = j~-8 each, j8-10, 1s-13, 1~-5 z5=z6-10 each, Z4-18, js=zg=z4-10 each, zs-13, S4 = S6 = Sa-16 each, S4 = S5-18 each, Z5-29, rs-16, R1-13. Metasternal setae on metasternal plates. Ventrianal shield with strongly concave lateral margins, with 4 pairs of preanal setae and a pair of pores. Fixed digit of chelicera with 2 teeth, movable digit with one tooth. Macroseta on leg IV : basitarsus-2.3. Pakistan. Subgenus Typhloctonus Muma Key to the species of Typhloctonus 1. Ventrianal shield with 3 pairs of preanal setae Ventrianal shield with 4 pairs of preanal setae 2. Z~ serrate, knobbed; Ventrianal shield considerably longer than wide Za serrate, not knobbed; Ventrianal shield only slightly longer than wide nesbitti 2 transitans vollsella 268. Typhlodromus (Typhloctonus) nesbitti Womersley Typhlodromus nesbitti Womersley, Aust. J. Zool., 2: (Holotype ~,Australia: New South Wales, on galls of lucerne, in South Australian Museum) Typhlodromus (Typhlodronlus) nesbitti, Narayanan & Ghai, Proc. natn. Inst. Sci India, 29B (5) : (Delhi: Mango malfomation) Typhlodromus (Typhloctonus) nesbitti, Gupta, Fauna of India (Ac~ri: Meso$tjgmata) Famil)' Ph)'toseiidae, p ,

145 GUPTA: ORIBNTAL PHYTOSEIIDAE Diagnosis: Dorsal shield with 19 pairs of setae. Measurements of setae: i l -28, j4-17, i5-1?, j6-17, 12-20, 1{)-11, is-20, z2-20, z,-22, z4:-22, 54-25, ss-25, Z1-20, S228, S4-22, S5-22, Z{)-56, zl)-17, Z4-36 Ventrianal shield 143 long, 110 wide, with 3 pairs of preanal setae and a pair of preanal pores. Macroseta present only on basitarsus IV. India (Delhi), Australia Typhlodromus (Typhloctonus) transitans Gupta Typhlodromus transitans Gupta, Indian J. Acar., 5 : (Holotype ~ t India: Jammu & Kashmir, Anantnag, on pear. Paratypes 2 ~ ~, Srinagar, on mulberry, all in ZSI) Typhlodromus (Typhloctonus) transitans, Gupta, Fauna of India (Acari : Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield lightly sclerotized, reticulate, 320 long, 176 wide, with 19 pairs of setae. Measurements of setae: jl-16, i 4-16, i 5-16, i s -18, la-20, J 5-8, i s -21, za-16. zs-21, z4-24, , s8-24, Z1-20, Sa-28, S4-28, S~-20, ZI)-48, z5-l6, Z4-28, rs-21, R l -20. Ventrianal shield smooth, 92 long, 64 wide, with 4 pairs of preanal setae. Macrosetae on leg IV: genu-20, tibia-18, basitarsus-36. India (Jammu & Kashmir) Typhlodromus (Typhloctonus) vollsella (Chaudhri, Akbar & Rasool)-New Comb Typhloctonus vol/sella Chaudhri, Akbar & Rasool~ Rept. PL 480 Project OAL, p (Holotype ~,Pakistan: 2 miles east of Murree hills, 7500', on Pinus sp., in UAL). Diagnosis: Dorsal shield reticulate, 326 long, 286 wide, deeply concave near R 1, with 19 pairs of setae. All the setae shorter than distance of seta next in line, except js which is longer than js-zs. Measurements of setae: j1-18, j4-13, j5-16, i6-18, l a -18, 1 5-8, zg-16, Z4-26, is-21, za-18, zs-2l, z4-21, 54-23, s6-26, Zl-18, Sa-26, S4-26, S&-18, Z5-39, rs-23, R l -18. Metasternal setae on platelets. Ventrianal shield 101 long, 75 wide, with lateral margins nearly parallel, with 4 pairs of preanal setae and a pair of pores. Leg IV with 3 macrosetae, genu-16, tibia-16, basitarsus-31, with blunt tip, that on basitarsus IV with hyaline margin.

146 132 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 95 Distribution; Pakistan. In addition to the above, the following species are also known but those could not be included in key for the reasons as given for Amblyseius (Amblyseius) spp. on p Typhlodromus (Typhloctonus) prunus (Denmark & Rather)-New Comb Typhloctonus prunus Denmark & Rather, Internal. J. Acarol., 10 (3): (Holotype ~,India : Srinagar, on Prunus, in USNM '1). Diagnosis: For want of original literature, this information could not be furnished.? Remarks: Gupta. India (Kashmir). In all probability this species is same as T. (T.) prunus Subgenus Typhlodromus Scheuten Key to the species of Typhlodromos 1. Ventrianal shield much longer than broad... Ventrianal shield almost as long as broad.. 2. SI) very small, nearly 1/2 as long as S" and much shorter than its distance between its base and that of Z IS 8 5 subequal to S", Z6 almost touches base of Z5 garhwalicus 2 communis neorhenanus 272. Typhlodromus (Typhlodromus) communis Gupta Typhlodromlls communis Gupta, Entomologists' mon. Mag., 115 : 209. (Holotype ~, Paratypes 2 ~ ~, India: West Bengal) Darjeeling Dist., on pear, Pyrlls comlnunis, in Z8I) Typhlodromus (Typhlodrulnus) communis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 280 long, 160 wide, reticulate, with 18 pairs of setae. Measurements of setae: i l -13, i4-13, j~-10, j6-12, -1a-14, J 5-5, js-12, Z2-6, zs-12, z4-12, s4,-13. s6-16, Sa-13, S4-21, S~-11, Zfi-47 (smooth), z~-12, (smooth). Ventrianal shield as long as 'wide,

147 GuPTA: ORIENTAL PHYTOSEIlDAE 133 with 3 pairs of preanal setae. Macroseta present only on basitarsus IV-35. India (West Bengal) Typhlodromus (Typhlodromus) garhwalicus Gupta Typhlodromus garhwalicus Gupta, Indian J. A car., 6 : (Holotype ~,India : Uttar Pradesh, Almora, Kausani, on Casuarina sp., in ZST) Typhlodromus (Typhlodromus) garhwalicus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shied 387 long, 228 wide, with 18 pairs of setae; mostly small except Z5 which is longer. Measurements of setae: jl-13, j.. -j6' 1 a each, J 5-8, js-22, za-17, za-17, z4-17, s4-13, s6-22, , S4-30, S5-21, Z5-39, Z4-26, z5-14, ra-29. Ventrianal shield 129 long, 69 wide, with 4 pairs of preanal setae. Leg IV with macroseta only on basitarsus. India (Uttar Pradesh) Typhlodromus (Typhlodromus) neorhenanus Gupta Typhlodromus neorhenanlls Gupta, Indian J. Acar., 2(2) : 4. (Holotype ~,Paratypes 3 ~ ~, 1 0, India: Meghalaya, Tura, on fern, in ZSI) Typhlodromus (Typhlodromus) neorhenanus, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 296 long, 156 wide, with 18 pairs of setae. Measurements of setae: jl-16, j4-10, j5-16, j6-20, J a -24, J 5-8, j s -13, Z a -8, z 3-16, z 4-22, s 4-25, s 6-2 9, Sa -33, S 4-36, S 5-32, Z 5-56, z5-16, Z4-44. Ventrianal shield broader anteriorly, lateral margins concave, with 4 pairs of preanal setae. Spermatheca with funnelshaped cervix. India (Meghalaya). In addition to the above species, some more species are known which could not be included in key for the reasons as mentioned for Amblyseius (Amb/yseius) spp. on p. 14.

148 134 : tee. ZOOL. SURV. INDIA, Oee. PAPER No Typblodromus (Typblodromus) disparis (Chaudhri, Akbar & Rasool)-New Comb Denmarkia disparis Chaudhri, Akbar & Rasool, Rept. PL 480 Project, UAF, p.46. (Holotype ~,Pakistan : Mansehera, in UAF). Diagnosis: Dorsal shield reticulate posteriorly, 418 long, 214 wide. Measurements of setae: jl-31, j4-j5-16 each, j6-19, J 2-23, J 5-16, js-42, za-21, Z3 = S6 = 26 each, Z4-29, S4 = Sa = S4-23 each, S5-23, Z5-78, z5-10, Z4-44, r 3-23, R Macroseta on basitarsus IV-34. Pakistan Typblodromus (Typblodromus) barkeri (Hughes) Neoseiulus barker; Hughes, Min. Agr. Fish. Lond., H. M. Stationary office, p (Holotype ~, London Docks, OD germinating barley, in BM (NH) Amblyseius (Amblyseius) barker;, Wei-nan, Acarology, 6(1) p Diagnosis: Dorsal shield imbricate with 18 pairs of setae. Ventrianal shield slightly longer than wide with 3 pairs of preanal setae. One pair of metapodal plate present. Spermatophoral process hammer-shaped. South China, England, Taiwan Typhlodromus (Typhlodromus) sijiensis Gupta Typhlodromus (Typhlodromus) sijiensis Gupta, Oriental Ins Typhlodromus (Typhlodromus) sijiensis, Gupta, Fauna of India (Acari: Mesostigmata) Family Phytoseiidae, p Diagnosis: Dorsal shield 370 long, 262 wide, with 18 pairs of setae. Measurements of setae: jl-26, j4-20, j5-20, j6-22, J a-24, J 5-7, js-44, zj-41, zs-35, z4-44, s4-44, s6-42, Sa-37, S4-34, S5-26, Z5-56, zc;-20, Z4-38, r s 40, R l -26. Ventrianal shield 125 long, 109 wide, with 4 pairs of preanal setae. Macroseta present only on basitarsus IV-56. India (Arunachal Pradesh).

149 GUPTA: ORIENTAL PHYTOSElIDAE 135 Subfamily GNORlMINAE Chaudhri Gnoriminae Chaudhri, Pak. J. Agril. Sci., 12(1-2): 100. Diagnosis: Dorsal shield light to heavily sclerotized with 12 pairs of lateral setae of diverse length, 3 pairs of sublateral setae, 2 pairs of setae around ventrianal shield, ventrianal shield large, robust, reticulate, with 6-7 pairs of preanal setae, a pair of preanal and a long serrate post anal seta present; a fold like structure present on ventrianal shield little anterior to anal region. Type genus: Gnorimus Chaudhri, Key to the genera of Gnoriminae 1. Ventrianal shield with 7 pairs of preanal setae, metapodal and metasternal plates absent Ventrianal shield with 6 pairs of preanal setae, 2 pairs of metapodal plates present... Garhwalicus Gnorimus Genus Garhwalicus Gupta & Ray Garhwalicus Gupta & Ray, Bull. zool. Surv. India, 4(3) : 268. Type: Garhwalicus himalayensis Gupta & Ray, Garhwalicus himalayensis Gupta & Ray Garhwalicus hima/ayensis Gupta & Ray, Bull. zool. Surv. India, 4(3) : (Holotype ~,India : Uttar Pradesh, Garhwal region, Siror vill., on undet. plant., in ZSI). Diagnosis: Dorsal shield 370 long, 278 wide, heavily sclerotized with 12 pairs of lateral setae, 6 pairs of dorsocentral setae, 2 pairs of median setae and 3 pairs of sublateral setae. Most of the dorsal setae rod-like, a few with widened tip. Measurements of setae: jl-18. j4-31 (broken), h~-29, j8-34, J 2-7, J~-4, js-38, za-5, zs-44, z4-36 (broken), S4-5, s6-44, Zl-6, S2-44 (broken), Ss-5, S4-5, S~-5, Z~-60, z~ broken, Z4-6, fs-38, R1-small, R4, -small. Ventrianal shield 190 long, 210 wide, reticulate, with 7 pairs of preanal setae, postanal seta rod-like. Leg IV with 2 macrosetae on genu (27 long, 18 long), 2 macrosetae on tibia (18 long and 20 long), basitarsus with one macroseta-22, all being spatulate. India (Uttar Pradesh).

150 136 REC. ZOOL. SURV. INDlA, OCC. PAPER No. 95 Genus Gnorimus Chaudhri Type: GnDrimus tabella Chaudhri, GDorimus tabella Chaudhri Gnorimus tabella Chaudhri, Pak. J. Agric. Sci., 12(1-2): (Holotype ~, Pakistan: 2miles north of Shalamar Rd., Lahore, on Cynodon dactyion, in U AL). Diagnosis: Dorsal shield reticulate, 370 long, 220 wide, with 25 pairs of setae and 8 pairs of pores. Measurements of setae: jl-34, j4-18, j5-23, j6-26, J a -18 each, J 5-8, ja-42, za-39, za-36, z4-18, s4-52, s6-42, Z1-55, 8 a -52, 8 a -47, , , Z5-65, z5-31, Z~ (Ma)-60, Z4 (Ms)-60. Ventrianal shield reticulate, 161 long, 172 wide, almost oval with 6 pairs of preanal setae. Metapodal plates 2 paired. Spermatheca with cervix funnel-shaped. Macrosetae on leg IV: genu-44, basitarsus-62, distitarsus-81. Pakistan. SUMMARY The present work concerns the phytoseiid fauna of Oriental region and deals with 279 species belonging to 12 genera under 2 subfamilies. Besides diagnosis and distribution, the relevant references along with locality and habitat records, particulars about type specimens with repositories, etc. in respect of all the species are provided. Keys to subfamilies, genera, subgenera and also for most of the species are included. Several generic transferances are proposed and a subgeneric concept is followed. The number of species from different parts of Oriental region are : India-142, Pakistan-65, Thailand.. 29, Taiwan-26, Philippines-21, Indonesia-15, Malayasia-6 and South China-40. So far nothing was known from Nepal, Bhutan, Burma and Bangladesh and, hence, a preliminary list of species from those regions basing on author's own recent study is provided in Appendix-I and that reports-7, 11, 7 and 9 species, respectively from those regions. Appendix II represents the country-wise distribution of species in Oriental region. The present account reveals that Oriental region represents a little less than 30% of the known world phytoseud faqna.

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156 142 REC. ZOOL. SURV. INDiA, OCC. PAPER No. 95 GUPTA, S. K. & NAHAR, S. C Plant mites (Acari) of agricultural importance in Bihar. In Contrib. to Acar. in India, pp (Ed. G. P. ChannaBasavanna). GUPTA, S. K. & RAY, S A new genus and one new species under subfamily Gnoriminae Chaudhri from India (Acarina: Phytoseiidae). Bull. zool. Surv. India, 4: GUPTA, s. K. & RAY, S. 1981a. Species of the subgenera Paraphytoseius and Asperoseius from India, with description of a new species of Paraphytoseius from India. Bull. zool. Surv. India, 4 : GUPTA, S. K., SIDHU, A. S., DHOORIA, M. S. & SINGH, G Preliminary note on the phytophagous and predatory mite fauna of the Punjab and Himachal Pradesh. Sci. & Cult., 37 : HUFFAKER, C. B. & SPITZER, C. H Data on the natural control of cyclamen mite on strawberries. J. econ. Ent., 44 : HUFFAKER, C. B., VAN DE VRIE, M. & McMurtry, J. A Ecology of tetranychid mites and their natural enemies: A review. II. Tetranychid populations and their possible control by predators: An evaluation. Hilgardia, 40 : JAGADISH, P. S. & NAGESHCHANDRA, B. K Biology of Typhlodromips tetranychivorus (Acari: Phytoseiidae) on red palm mite Raoiella indica (Acari: Tenuipalpidae). Abstr. I All India Symp. Acar., pp. 49. KAMATH, S. M Studies on the feeding habits, development and reproduction of the predaceous mite, Phytoseiulus persimilis A. H. (Acarina: Phytoseiidae) on some phytophagous mites in India. Comm. Inst. Bioi. Contr. Tech. Bull., -10; KAZIMI, S. K. & GHANI, M. A Some phytoseiid mites of west Pakistan. Comm. Inst. Bioi. Contr. Tech. Bull. 13. JIE-LIEU, XIN., LAI-RONG, LIANG. & LI-SHENG, K Three new species of the genus Typhlodromus Scheuten (Acarina: Phytoseiidae). Fudall J. (Nat. SCi.), 19 : LAI-RONG, LIANG. & Ll-SHENG, K Two new species of the genus Amblyseius Berlese (Acarina: Phytoseiidae). Fudan J. (Nat. Sci.), 20 : LAI-RONG, LIANG & LI-SHENG, K. 1981a. Two new species of the genus Phytoseius from China (Acari: Phytoseiidae). Entomotaxonomia 3 :

157 GUPTA: ORIENTAL PHYTOSEIlDAE 143 Lo, P. K. C Phytoseiid mites from Taiwan (Acari: Mesostigmata). Bull. Sun Yat-sen Cult. Found., No.5: MALLIK, B Biology of Amhlyseius /ongispinosus (Evans) (Acarina: Phytoseiidae) and Tetranychus ludeni Zacher (Acarina: Tetranychidae) and interaction between them. M. Sc. thesis. Univ. Agri. Sciences., Bangalore, 72 pp. MALLIK, B. & CHANNABASAVANNA, G. P Life history and life table of Tetranychus ludeni and its predator Amhlyseius lortgispinosus (Acari: Tetranychidae, Phytoseiidae). Indian J. Acar., 8: McMURTRY, J. A., HUFFAKER, C. B. & VAN DE VRIE. M Ecology of tetranychid mites and their natural enemies: A review. I. Tetranychid enemies, their biological characters and the impact of spray practices. Hilgardia, 40 : McMURTRY, J. A. & SCRIVEN, G. T Studies on the feeding, reproduction and development of Amblyseius hibisci (Acarina : Phytoseiidae) on various food substances. Ann. ent. Soc. Al'n., 57 : McMURTRY, J. A. & SCRIVEN, O. T. 1964a. Biology of the predaceous mite Typhlodromus rickeri (Acarina: Phytoseiidae). Ann. ent. Soc. Am., 57 : MUMA, M. H New Phytoseiidae (Acarina: Mesostigmata) from southern Asia. Fla. Ent., SO : MTJMA, M. H. & DENMARK, H. A Phytoseiidae of Florida. Arthropods of Florida and neighbouring land areas: 6 : NARAYANAN, E. S. & GHAI, S Some new records and ne"w species of mites associated with malformation of mango trees in India. Proc. natn. Inst. Sci. India, 29(B) : NARAYANAN, E. S. & KAUR, R. B Two new species of the genus Typhlodromus Scheuten from India (Acarina: Phytoseiidae). Proc. Indian Acad. Sci., 51 (8) : 1-8. NARAYANAN, E. S., KAUR, R. B. & OHAI, S Importance of some taxonomic characters in the family Phytoseiidae Berl., 1916 (predatory mites) with new records and descriptions of species. Proc. natn. Inst. Sci. India., 26 (B) ;

158 144 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 NARAYANAN, E. S. & KHOT, N. S Studies on some entomophagous mites and their possible use in biological control. Abstr. Indian Sci. Congr., 46 : 411. OOMEN, P. A Studies on population dynamics of the scarlet mite, Brevipalpus phoenicis, a pest of tea in Indonesia. Pub. H. Veenman & Zonen, B. V, Wagenangen, 89 pp. PARROTT, P. J., HODGKISS, H. E. & SCHOENE, W. J The Eriophyidae. Part I. The apple and pear mites. N. Y. Agr. Exp. Stat Bull., 283 : PRASAD, V A catalogue of mites of India. Indira Acarology Publising House, Ludhiana, 320 pp. PUTTASWAMY Studies on the ecology of Tetranychus ludeni Zacher (Acari: Tetranychidae) and its interaction with the predator, Typhlodromips tetranychivorus Gupta (Acari: Phytoseiidae). Ph. D. thesis Univ. Agr. Sciences., Bangalore. RAO, V. P. & RAO, V. S Two new records of predaceous mites from India. Comm. Inst. Bioi. Contr. Tech. Bull., 4 : RAO, V. P., DUTTA, B. & RAMASESHIAH, G Natural enemy complex of flushworm and phytophagous mites on tea in India. Tea Board Sci. Pub. Sere No.5: RATHER, A. Q New species and new records of PhytDseius Ribaga (Acari: Phytoseiidae) from India. Acarologia 26(1): RAY, S. & GUPTA, S. K A new species of Platyseiella Muma (Acari: Phytoseiidae) with collection records of twelve other species. Bull. zoo!. Surv. India, 4(3) : RAY, S. & GUPTA, S. K Some new records of Phytoseiidae from India (Acarina: Mesostigmata). Rec. zool. Surv. India, 80 : SANDHU, G. S., KAUSHAL, K. K. & GUPTA, S. K Mites associated with maize and their predators in the Punjab. Sci. & Cult., 39 : SCHICHA, E, & CORPUZ-RAROS, L. A Contribution to the knowledge of the genus Paraphytoseius Swirski & Shechter (Acarina: Phytoseiidae). Internat. J. Acarol., 11(2) : SHIH, T CHAIN-lNG, Field evaluation of five species of Amblyseius on Tetranychus kanzawai. Acaro!o8Y~ 6 : ,

159 GUPTA: ORIENTAL PHYTOSEIIDAE 145 SOMCHOUDHURY, A. K Seasonal fluctuation of population of Amblyseius delhiensis, a predator on eggs of cotton jassid. Abstr. I All India Symp. Acar., pp SOMCHOUDHURY, A. K Seasonal fluctuation of Amblyseius delhiensis (Acari: Phytoseiidae), a predator on eggs of cotton jassid. In Contrib. to Acar. in India, pp SWIRSKI, E. & GOLAN, Y On some phytoseiid Inites (Acarina) from Luzon Island (Philippines). Israel J. agric. Res., 17 : (1967). TJYING, I. S Two new species of the mite family Phytoseiidae (Acarina: Mesostigmata) from Taiwan. Pl. Prot. Bull. Taiwan 14: 1-7 (1972). TSENG, Y. -H Systematics of the mite family Phytoseiidae from Taiwan with a revised key to the genera of the world. II. J. Agr. Assoc. China New Ser., 94 : TSENG, Y. -H Two new predatory mites from Taiwan (Acarina: Cheyletidae, Phytoseiidae). Pl. Prot. Bull. Taiwan, 15(2): (1973). TSENG, Y. -H Mites associated with weeds, paddy, rice and upland rice fields in Taiwan. In: Griffiths, D. A. & Bowman, C. E. (eds.), Acarology, 6, Ellis Horwood Ltd. Chichester London, 2 : VAN DE VRIE, M., McMURTRY, J. A. & HUFFAKER, C. B Ecology of tetranychid mites and their natural enemies. A review. III Biology, ecology and pest status and host plant relations of tetranychids. Hilgardia, 41 : WEI-NAN, W Two new species of Amblyseius from Yunnan Province, China (Acarina: Phytoseiidae). Acta Zootaxonomica flinica, 9: WEI-NAN, W 1984a. Notes on the genus Amblyseius Berlese with descriptions of two new species from citrus orchards in South China. In: Griffiths, D. A. & Bowman, C. E. (eds.), Acar%gia, 6, Ellis Horwood Ltd. Chichester London, 1 : WEI-NAN, W. & ZHAO-QUAN, L Three new species of the genus Phytoseius from South China. Acta Entomologica Sinica, 27(4) :

160 146 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 WEI-NAN, W. & ZHAO-QUAN, L Three new species of the genus Amblyseius Berlese from South China (Acarina: Phytoseiidae). Acta Zootaxonomica Sinica, 10(3) ; WEI-NAN, W. & ZHAO-QUAN, L. 1985a. Four new species of the phytoseiid mites from Hainen Island, Guangdong Province (Acarina: Phytoseiidae). Acta Zootaxonomica Sinica, 10(4): WEI-NAN, W. & XING, Q Two new species of the genus Amblyseius Berlese (Acari: Phytoseiidae). En tomo taxonomia, 5(3): WEI-NAN, W. & XING, Q. 1983a. A new species of Okiseius Ehara and the male of o. subtropicus Ehara (Acarina: Phytoseiidae). Entomotaxonomia, 5(1) :

161 APPENDlx-t List of phytoseiid species known from Nepal, Bhutan, Burma and Bangladesh (basing on author's recent study) Name of species Countries Nepal Bhutan Burma Bangladesh A. (A.) largo ens is A. (A.) herbicolus + + A. (A.) kulini + A. (A.) sp. + + A. (E.) coccineae A. (E.) ovalis + A. (E.) pruni A. (P.) multidentatus + + A. (N.) sp. + A. (T.) kalimpongensis + + A. (T.) crotalariae + A. (T.) suknaensis A. (T.) syzygii + P. (P.) roseus + + T. (A.) darjeeiingensis + + T. (0.) pruni + T. (T.) communis + T. (T.) sp. + + Total

162 A~PENDIX-li Distribution of Phytoseiidae in different countries of the Oriental region [+ indicates record of that species] Name of species Countries en ~.~ Q) cd,.q t\s.6.~ c= ""d -.-I en () en (\i p. s:: ~ Q) m ~ A (\i cd d ~ -.-I -~ -.-I ~ ~ ~ 0 ~..s '"8 6 c; ~ ~ ~ ~ ~... C/) ~ (\i en Y-4.-1,...c.,-4.,..c cd "'C 1. Amhlyseius (Amhlyseius) adhatodae A. (A.) aeria/is A. (A.) bajulus A. (A.) brevieervix A. (A.) channabasavannai A. (A.) cinetus A. (A.) deductus A. (A.) excelsus A. (A.) hainanensis A. (A.) hapoliensis A. (A.) herbicolus A. (A.) impaetus A. (A.) indirae A. (A.) kulini A. (A.) largoensis A. (A.) maai A. (A.) mcmurtryi A. (At) monomaeroseta A. (A.) muraleedharani A. (A.) neorykei A. (A.) obtuserellus A. (A.) obtusus A. (A.) orientalis A. (A.) paraaerialis A. (A.) pubes A. (A.) shoreae A. (A.) spineus A. (A.) tamtavensis ,- 29. A. (A.) theae (\i.. -

163 ApPENDix-Ii 149 Name of species Countries rn ro d v... d ro ro.....d ~ d U rn ro ~ d d rn v ro ~ ~ ro ro d..d ~ ro rn ~.-4 ~ ro ~ ~ ro ~ =' ro d ro..d ro..d d ~ ~ ~ ~ ~ ~ f-c f-4 ~ en 30. A. (A.) vestificus A. (Asperoseius) heveae A. (A.) hevearum A. (A.) hyauliangensis A. (A.) nucifera A. (Euseius) aizawai A. (E.) alangii A. (E.) alstoniae A. (E.) bambusae A. (E.) coccineae A. (E.) coccosocius A. (E.) delhiensis A. (E.) densus A. (Eo) eucalypti A. (E.) finlandicus A. (E.) guangxiensis A. (E.) irregularis A. (E.) macrospatulatus ~ A. (E.) mediocris A. (E.) neococcineae A. (E.) nicholsi A. (E.) notatus A. (E.) okumae A. (E.) ovalis A. (E.) pruni A. (E.) relictus A. (E.) rhododendronis A. (E.) ruiliensis A. (E.) sacchari A. (E.) septicus A. (Eo) womersleyi A. (E.) yunnanensis A. (N eoseiulus) aceriae A. (N.) aequipilus ~

164 150 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 95 Nan1e of species Countries en Q.~ v Cd ~ Q ~.~ 'U ~.~.,...c en () en Cd -0.. Q ~ Q) ~.IoJ 0.. ~ ~ ~ Cd en,... ~.,...c C'CS.~.~.,...c ~ 0.IoJ,....~.~ 'U ~ 'U ==' C'CS C'CS Q ~ ~ C'CS ~... p... p... f-4 f-4... en 64. A. (N.) amantis A. (N.) anuwati A. (N.) assamensis A. (N.) bakeri A. (N.) bheraensis A. (N.) calorai A. (N.) carancula A. (N.) curvus A. (N.) cynodonae A. (N.) eremicus A. (N.) Jallacis A. (N.) Jraterculus A. (N.) imbricatus A. (N.) indicus A. (N.) longispinosus A. (N.) myrtia A. (N.) paspalivorus A. (N.) rangatensis A. (N.) reductus A. (N.) siamensis A. (N.) stolidus A. (N.) tabularis A. (N.) taiwanicus A. (N.) venustus A. (paraphytoseius) cracentis A. (P.) multidentatus A. (P.) scleroticus A. (P hytoscutella) salebrosus A. (P.) wongsirii A. (proprioseiopsis) arunachalensis A. (P.) beatus A. (P.) bulga >6. A. (P.) dentatus A. (P.) peltatus ~ ~ 0 ~ ~

165 APPENDIX-II 151 Name of species Countries ~ C/.) d Q)...-t ~ C'\S d...-t "'d..d...-t a...-t C/.) () en C'\S ~ ~ ~ Q) C'\S ~ 04 C'\S C'\S d ~ en..d ~...-t.-4,...c...-t 0.~...-t ~ ~.! "'tj ::s ~ a C'\S..d ~ d 0 ~...-t '"C...-t C'\S...-4 P-4 ~ E-t... V) 98. A. (P.) synachattiensis A. (Proprioseius) kumaonensis A. (Typhlodromalus) chikmagalurensis A. (T.) chitradurgae A. (T.) eucalypticus A. (T.) ficusi A. (T.) jarooa A. (T.) kalimpongensis A. (T.) laaensis A. (T.) lablabi A. (T.) manipurensis A. (T.) newsami A. (T.) sorghumae A. (Typhlodromips) areeae A. (T.) asiatiew A. (T.) baiyunensis A. (T.) crotalariae A. (T.) draconis A. (T.) eujeniae A. (T.) ghanii A. (T.) guajavae A. (T.) hidakai A. (T.) johoreae A. (T.) lenis A. (T.) lab is A. (T.) linearis A. (T.) mangiferae A. (T.) meghalayensis A. (T.) neoc rotalariae A. (T.) neoghanii A. (T.) okinawanus A. (T.) officinaria , A. (T.) ogurai

166 152 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 Name of species Countries Cd.~ d Cd Cd (I.) G).9 Cd ('d. c:l "ts... c3... (I.) (I.) ~ Q) Cd.f"'4.. 8: ~,... P"'"C... 0 ~ as (I.) ~..d ~ ~ ~.~,... ~ ('d ::s 0 ~ ~ ~ '1j ~ -.-4.,... "ts Cd ~ Cd..a cts...-.c ~ ~ f-c A. (T.) perfectus A. (T.) polyantheae A. (T.) potentillae A. (T.) repletus A. (T.) sapienticola A. (T.) similis A. (T.) sijiensis A. (T.) suknaensis A. (T.) syzygii A. (T.) tetranychivorus + _ A. (T.) vignae Amathia rancidus Indoseiulus eharai ricini Iphiseius (lphiseius) andamanicus (I.) hapoli I. (Trorhoseius) bakeri Paraamblyseius formosanus P. fragariae P. mumai Platyseiella formosanus P. mumai Okiseius chinensis o. himalayana -' O. sikkimensis + _ O. subtropicus O. yazuliensis lndodromus meerutensis Phytoseius (Pennaseius) amba P. (P.) curvatus P. (P.) diutius P. (P.) hongkongensis P. (P.) improcerus P. (P.) incisus

167 APPENDIX-II 153 Name of species Countries ~ CIl ~ (1)...c ~ ~ ~ ro ~...c c:1....",", CIl CIl cd ~ U ~ (1) cd ~ A cd cd ~ r:n cd 0 ~ ~ cd g cd a ~ cd ~ ~ :>-...c ~ "'d...c... ~ "'d cd... c:1 ~ ~ (-4 ~... rj) ~ 165. P. (P.) kapuri P. (P.) minutus P. (P.) namdaphaensis P. (P.) taianensis Phytoseius (Phytoseius) bandipurensis P. (P.) brevicrinis P. (P.) coheni P. (P.) corniger P. (P.) crinitus P. (P.) domesticus P. (P.) glareosus P. (P.) hawaiiensis P. (P.) huangi P. (P.) indicus P. (P.) intermedius P. (P.) jujuba P. (P.) longus P. (P.) macropilis P. (P.) macrosetosus P. (P.) mansehraensis P. (P.) meyerae P. (P.) mixtus P. (P.) neocorniger P. (P.) neoferox P. (P.) nipponicus P. (P.) petentis P. (P.) punjabensis P. (P.) qianshanensis P. (P.) rachelae P. (P.) rasilis P. (P.) rimandoi P. (P.) roseus P. (P.) rugatus _.. '- 20 +

168 154 REC. ZOOL. SURV. INDIA, OCC. PAPBR No. 95 Name of species Countries ('j CIl ~ Q)... ~ ~ Q ~ Q "'d (J.,.-4 CIl ~ 0. Q d Q) ('j ~ ('j P4 ~ Q ('j en.,.-4.,.-4 t;;:::....s ~...-4.~ 0 ~.,.-4 ('j '"ts ::s ~ '"CS ~... ~ ~ ~ ~... en \~ ~ ~...cl c:= P. (P.) rugosus P. (P.) ruidus P. (P.) silvaticus P. (P.) spatulatus , P. (P.) swirskii P. (P.) wainsteini Typhlodromus (Amblydromella) acacia T. (A.) arunachalensis T. (A.) balakotiensis T. (A.) bambusicolus T. (A.) cordiae T. (A.) chrysanthemi T (A.) dactylifera T. (A.) divergentis T. (A.) darjeelingensis T. (A.) eremitidis T. (A.) fleschneri T. (A.) galummatus T. (A.) ghanii T. (A.) gopali T. (A.) himalayensis T. (A.) homalii _ T (A.) husaini T. (A.) incasus T. (A.) jackmickleyi T. (A.) kodaikanalensis T. (A.) libitus T. (A.) Iimitatus T. (A.) loralaiana T. (A.) maracus T. (A.) mori T. (A.) nidibu T. (A.) nilgiriensis _.. r en ~

169 APPENDIx-it 155 Name of species Countries c'tj.p04 ""d ~ c'tj en.9 Q) c'tj c:: c::.,...c ""d.j::.p04.p04 c:: en U en c'tj ~ Q) c'tj c'tj ~ c::..d >-- ~ en 0. ~ ~ c'tj ~ c'tj..d c'tj.j:: ~.,...c.,...c.,...c c'tj 0 ~ ~.,...c.,...c ""d c'tj =' ~ 0 ~ ~ ~ f-4 f-4 ~ (J) 231. T. (A.) octavus T. (A.) ornatulus T. (A.) platycladus T. (A.) philippinus T. (A.) rhenanus T. (A.) rhododendroni T. (A.) ryukyuensis T. (A.) serratus T. (A.) serrulatus T. (A.) sica T. (A.) sonprayagensis T. (A.) umbratus T. (A.) zafari T. (Anthoseius) majumderi T. (Brethria) confusus T. (B.) roshanlali T. (Chiliseius) lieni T. (C.) se paratus T. (C.) tengi T. (Clavidromus) neotransvaalensis T. (Metaseiulus) contiguus T. (Orientiseius) agilis T. (0.) c hannabasavannai T. (0.) hadii T. (0.) ialazariensis T. (0.) manipurensis T. (0.) orissaensis T. (0.) pruni T. (0.) rickeri T. (0.) samliensis T. (0.) thailandicus T. (Paraseiulus) comptus T. (P.) ignavus c'tj ~

170 156 REC. ZOOL. SURV. INDIA, OCC. PAPER No. 95 Name of species Countries as U) d Cl)..c as as d.e.~ "'C..d..c U) U) () ~ 0. Q d Cl) as as 1;; ~ t'\s t'\s d ~...-f.s..c..c.,..c.~ ~...-c 0.-4 ~.,..c.~ t'\s ::s "'C d t'\s..d t'\s..d "CJ ~ d 0... ~ ~ f-4 f-4 ~ en 264. T. (P.) neosoleiger T. (P.) operantis -/ T. (P.) trimediosetus T. (P.) vitreus T (Typhloctonus) nesbitti T. (T.) pruni T. (T.) transitans T. (T.) vol/sella Typhlodromus (Typhlodromus) barkeri T. (T.) communis T. (T.) disparis T. (T.) garhwalicus + - -' 276. T. (T.) neorhenanus T. (T.) sijiensis Garhwalicus himalayensis + - -,, Gnorimus tabella Total No~ of species ~

171 ADDENDA I. On page No.4, while giving key to the genera of Amblyseiinae, the inclusion of the genus Platyseie/[a was inadvertently omitted and, hence, the revised key is provided here to include that genus. Key to the genera of Amblyseiinae 1. Ventrianal shield absent or indistinctly demarcated, only preanal setae distinctly present Ventrianal shield distinctly present 2. Sublateral seta II (R 1 ) absent Sublateral seta II (R 1 ) present 3. Rl on dorsal shield Rl on lateral integument 4. Lateral integument sclerotized so that ra and Rl though on usual lateral position appear to be on lateroven tral extension of dorsal shield Lateral integument not sclerotized as above 5. Metapodal plates large, single paired, triangular, genital shield very broad and punctate; ventrianal shield massive ; genu III with 6 setae... Metapodal plates 2 paired, slender, elongate, genital and ventrianal shields usually narrow ; genu III with 7 setae Indoseiulus 2 Platyseiella 3 Okiseius 4 Iphiseius 5 P araamblyseius Amblyseius II. After the manuscript of this review had gone to the press, additional information on distribution in respect of some species c ame in hand and those are given below :

172 158 REc. ZOOL. SURV. INDIA, Oce. PAPER No. 95 Species Name of species Distribution Reference No. 5. Amhlyseius (Amhlyseius) herbicolus Taiwan Shih, 1984 South China Wei-nan, A. (A.) lorgoensis South China Wei-nan, 1984 Indonesia Oomen, A. (A.) mcmurtryi South China Wei-nan, A. (A.) orientalis South China Wei-nan, A. (A.) vestificus Taiwan Tseng, A. (A.) nichols; South China Wei-nan, A. (Euseius) ovalis South China Wei-nan, A. (N eoseiulus) anuwati South China Wei-nan, A. (N.) longispinosus Pakistan Chaudhri et al., South China A. (P araphytoseius) South China Wei-nan, 1984 multidentatus Taiwan Wei-nan, 1984 Tseng, A. (Proprioseiopsis) peltatus Taiwan Tseng, A. (Typhlodromalus) newsami South China Wei-nan, A. (Typhlodromips) hidakai Thailand Ehara & Bhandhufalck, A. (T.) okinawanus South China Wei-nan, Okiseius subtropicus South China Wei-nan & Xing, 1983a 164. Phytoseius (Pennaseius) minutus Taiwan Tseng, Phytoseius (Phytoseius) coheni Philippines Swirski & Golan, P. (P.) crinitus Indonesia o omen, 1984

173 ADDENDA 159 III. In case of species Nos. 34, 109, 144, 146, 157, 167,231 and 277, the original references were incompletely given because of nonavailability of the concerned publication at the time of sending the manuscript to the press. However, the publication reached the author when printing of this review was almost complete and, hence, those complete references are given below: Species No Amblyseius (Asperoseius) hyauliangensis Gupta, Oriental Ins., 21 : 112. Amblyseius (Typhlodroma/us) ficusi Gupta, Oriental Ins., 21 : 114. Indoseiulus eharai Gupta, Oriental Ins., 21 : 115. Iphiseius (Iphiseius) hapoli Gupta, Oriental Ins., 21 : Okiseius yazuliensis Gupta, Oriental Ins., 21: 116. P hytaseius (P ennaseius) namdaphaensis Gupta, Oriental Ins., 21: 117. Typhlodromus (AmblYdromella) arunachalensis Gupta, Oriental Ins., 21: Typhlodromus (Typhlodromus) sijiensis Gupta, Oriental Ins., 21: 119. IV. According to Lo (1984) [Acarology 6 (2) : ]-28 spp. while as per Shih (1984) [Acarology 6 (2) : ]-40 spp. of Phytoseiidae are known from Taiwan. Recently, Tseng (1982) (Bureau Cammod. Insp. Quar. Ministry of Econom. Affairs, 164 pp.) and Tseng (1983) (Chinese J. Entomol., 3 : 33-74) made further contributions on Taiwan phytoseiid fauna. Unfortunately, Tseng's above cited works were not available to the author and nor did Lo (1984) and Shih (1984) provided any list of phytoseiid spp. known from Taiwan. Hence, in all probability, the total species much exceeds the number (= 26) that has been included in this review. V. Denmarkia Chaudhri, Akbar & Rasool, 1979 has been treated here as synonym for Typhlodromus Scheuten as the position of s4

174 160 REC. ZOOL. SURV. INDIA, Oce. PAPBR No. 95 and S6 almost at same level in a single species (the type species of this genus) based on a lone specimen, does not appear to be an adequate reason for erecting this genus. The genera Ansaria and Kashmiricus both erected by the same authors in 1979 are considered here.as synonyms for Amblyseius Berlese, because the characters on which those two genera were erected are not uncommon in Amblyseius.

175 INDEX A acacia (Typhlodromus)-100, 102 aceriae (Amblyseius)-33, 34 adhatodae (Amblyseius)-6, 7 aequipi/us (Amblyseius)-33, 34 aerialis (Amblyseius)-7 agilis (Amblydromella)-123 agilis (Typhlodromus)-123 aizawai (Amblyseius)-20, 21 alangii (Amblyseius)-3'0 alstoniae (Amblyseius)-20, 21 amantis (Amblyseius)-43 amantis (Ansaria)-43 amba (Phytoseius)-83 Amblydromella-l00, 101 Amblydromus-l00 Amblyseiella-4 Amblyseiinae-4 Amblyseio psis-4 Amblyseiulella-4 Amblyseiulus-5 Amblyseius-4, 5, 6 Amblyseius (Amblyseiopsis)-4 Amblyseius (Amblyseius)-6 Amblyseius (Ptenoseius)-5 Amblyseius (Seiopsis)-4 Amblyscutus-5 Amathia-71 andamanicus (lphiseius)-73 Ansaria-5 Anthoseius-l 00, 101, 119 anuwati (Amhlyseius)-33, 34 arecae (Amblyseius)-58, 59 arunachalensis (Amblyseius)-50 arunachalensis (Typhlodromus)-115 asiaticus (Amhlyseius)-58, 59 asia tic us (Typhlodromus)-59 Asperoseius-4, 5 assamensis (Amblyseius)-33, 35 assamensis (Typhlodromus)-35 At hiaseius-5 Australiseiulus-l00 B baiyunensis (Amblyseius)-69 bajulus (Amblyseius)-14 bakeri (lphiseius)-73 balakotiensis (AmblYdromella)-103 balakotiensis (Typhlodromus)- 102, 103 bambusae (Amblyseius)-20, 22 bambusae (Euseius)-22 bambusicolus (Typhlodromus)- 101, 103 bandipurensis (Phytoseius)-85 baraki (Amblyseius)-33, 35 barkeri (Typhlodromus)-134 beatus (Amblyseius)-50 behraensis (Amblyseius)-43 behraensis (Neoseiulus)-43 Brethria-l01, 120 brevicervix-15 brevicrinis (P hytoseius)-85, 86 bulga (Amblyseius)-51 C calorai (Amblyseius)-34, 36 caruncula (Amblyseius)-44 caruncula (Neoseiulus)-44 caudiglans (Typhlodromus)-100 channabasavannai (Amblyseius)-7, 8 channabasavannai (Typhlodromus)- 123, 124 channabasavanni (Amblyseius)-8 channabasavanni (Typhlodromus)- 123 Chantiseius

176 162 REC. ZOOL. SURV. INDIA, OCC. PAPBR No. 95 chikmagalurensis (Amblyseius)-56 Chiliseius-100, 121 chinensis (Okiseius)-77 chitradurgae (Amb/yseius)-56 chrysanthemi (Typhlodromus)-102, 104 cinctus (Amblyseius)-6, 18 Clavidromina-100 Clavidromus-100, 101, 121 coccineae (Amhlyseius)-20, 22 coccosocius (Amhlyseius)-20, 23 coheni (P hytoseius)-85, 86 communis (Typhlodromus)-132, 133 comptus (Typhlodromus)-127, 128 confusus (Typhlodromus)-120 contiguus (Typhlodromus)-122 cordiae (Cydnoseius)-103 cordiae (Typhlodromus)-102, 103 corniger (Phytoseius)-85, 87 cracentis (Amblyseius)-48 cracentis (Ptenoseius)-48 crinitus (Phytoseius)-84, 8"7 crotalariae (Amhlyseius)-58, 60 curvatus (Phytoseius)-79, 80 curvus (Amblyseius)-44 Cydnodromel/a-lOO Jentatus (Proprioseiopsis)-51 Jisparis (Denmarkia)-134 Jisparis (Typhlodromus)~-134 diutius (Phytoseius)-79, 80.iivergentis (AmblYdromella)-105 tiivergentis (Typhlodromus)-102, '105 domesticus (Phytoseius)-98 draconis (Amblyseius)-69 draconis (Typhlodromips)-69 E Echinoseius-lOO eharai (Indoseiulus)-72 eharai (Typhlodromus)-lOl, 106 eremetidis (Amblydromella)-106 eremetidis (Typhlodromus)-106 eremicus (Amblyseius)-44 eremicus (Neoseiulus)-44 eucalypti (Amhlyseius)-20, 24 eucalypticus (Amblyseius)-52, 53 eucalypticus (Typhlodromalus)-53 eujeniae (Amhlyseius)-58, 60 Euseius-6, 20 excelsus (Amblyseius)-15 Cydnodromus-5 fallacis (Amblyseius)-33, 37 Cydnoseius -100 fallacis (Iphidulus)-37 cynodonae (Amblyseius)-33, 36 fallacis (Neoseiulus)-37 D falfacis (Typhlodromus)-37 dactylifera (Amblydromella)-104 ficusi (Amblyseius)-57 dactyllfera (Typhlodromus)-lOl, finlandicus (Amblyseius)-20, 24, finlandicus (Seiulus)-24 darjeelingensis (Typhlodromus)-105 fleschneri (Typhlodromus)-101, 107 deductus (Amblyseius)-15 formosanus (Paraamblyseius)-74 degenerans (Sejus)-72 formosanus (Platyseiella)-76 deledni (Amblyseius).-9 fraterculus (Amblyseius)-33, 37 delhiensis (Amblyseius)-20, 23, 24 fragariae (Paraamblyseius)-74 densus (Amhlyseius)-30 dentatus (Amb/yseius)-51 F G Galendromimus-100

177 INDEX 163 Galendromus-100 galummatus (Amblydromella)-107 galummatus (Typhlodromus)-l 01, 107 Garhwalicus-135 garhwalicus (Typhlodromus)-132-, 133 ghanii (Am b lydo1n ella ) ghanii (Amblyseius)-58, 61 ghanii (Cydnodromus)-60 ghanii (Typhlodromus)-102, 108 glareosus (Phytoseius)-85, 88 Gnoriminae-3, 134 Gnorimus-135 gopali (Typhlodromus)-102, 108 guajavae (Amhlyseius)-58, 60 guangxiensis (Amhlyseius)-30 H hadii (Typhlodromus)-123, 124 hainanensis (Amblyseius)-16 hapoli (Iphiseius )-73 hapoliensis (Amhlyseius)-16 hawaiiensis (Phytoseius)-85, 88 herhicolus (Amblyseius)-6, 9 heveae (Amhlyseius)-18 heveae (Typhlodromus)-18 hevearum (Amhlyseius)-19 hevearum (Typhlodromus)-19 hidakai (Amhlyseius)-58, 61 himalayensis (Garhwalicus)-135 himalayana (Okiseius)-76, 77 himalayensis (Ty phlodromus )-101, 108, 109 hongkongensis (Phytoseius)-79, 80, 81 homalii (Typhlodromus)-101, 109 huangi (Phytoseius)-85, 89 husaini (Typhlodromus)-115 hyauliangensis (Amhlyseius)-19 I ignavus (P araseiulus)-128 ignavus (Typhlodromus)-127, 128 imbricatus (Amblyseius)-33, 38 impactus (Amhlyseius)-6, 9 improcerus (Phytoseius)-79, 81 incasus (Amblydromella)-115 incasus (Typhlodromus)-115, 116 incisus (P hytoseius)-83 indicus (Amblyseius)-34, 38 indicus (Phytoseius)-85, 92 indirae (Amblyseius)-16 Indodromus-78 Indoseiulus-4, 71 Indoseius-71 insularis (Typhlodromus)-105 intermedius (Phytoseius) -84, 89 Iphidulus-100 Iphiseiini-72 Iphiseius-4, 72 ipomoeae (Amblyseius)-7 irregularis (Amhlyseius)-20, 25 irregularis (Typhlodromus)-25 J jackmickleyi (Typhlodromus)-116 jarooa (Amblyseius)-52, 53 jaunpurensis (Phytoseius)-85, 94 johoreae (Amblyseius)-58, 61 jujuba (P hytoseius)-84, 90 K kalimpongensis (Amblyseius)-52, 53 Kampimodromus-4 kapuri (P hytoseius)-79, 81, 82 Kashmiricus-5 kodaikanalensis (Euseius)-29 kodaikanalensis (Typhlodromus)- 102, 109, 110 kulini (Amblyseius)-7, 10

178 164 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 95 kumaonensis (Amhlyseius)-S2 kumaonensis (Proprioseius)-52 L laaensis (Amblyseius)-S 7 lobis (Amblyseius)-58, 62 lablabi (Amblyseius)-52, 54 lalazariensis (Orientiseius)-124 lalazariensis (Typhlodromus)-123, 124 largoensis (Amblyseius)-6, 9, 10, 11 lenis (Amblyseius)-57, 62 libitus (Alnblydromella )-116 libitus (Typhlodromus)-116 lieni (Chiliseius)-121 lieni (Typhlodromus)-121 limitatus (Amblydromella)-116 Iimitatus (Typhlodromus)-116 linearis (Amhl yseius )-58, 62 longispinosus (Amblyseius)-33, 39,40 /ongispinosus (Neoseiulus)-39 /ongispinosus (Typhlodromus)-39 longus (Phytoseius)-98 loralaiana (AmblYdromella)-110 loralaiana (Typhlodromus)-102, 110 M maa; (Amblyseius)-16 macropilis (Phytoseius)-85, 90 macrosetosus (Phytoseius)-84, 91 macrospatulatus (Amblyseius)-31 majumderi (Typhlodromus)-119, 120 mangiferae (Amhlyseius)-S7, 63 manipurensis (Amhlyseius)-S2, 55 manipurensis (Typhlodromus)-123, 125 mansehraensis (Phytoseius)-84, 91 maracus (Amblydromella)-117 maraeus (Typhlodromus)-117 mcmtlrtryi (Amhlyseius)-6, 11, 12 meerutensis (Indodromus)-78, 79 medioeris (Amblyseius)-31 meghalayensis (Amblyseius)-58, 63 Melodromus-100 Metaseiulus-100, 101, 102 meyerae (Phytoseius)-84, 91 minutus (Phytoseius)-79, 82 mixtus (Phytoseius)-84, 92 monomacroseta (Amblyseius)-17 mori (Typhlodromus)-lOl, 111 multidentatus (Amhlyseius)-46 multidelltatus (Paraphytoseius)-46 mumai (Paraamblyseius)-74, 75 mumai (Platyseiella)-75 muraleedharani (Amblyseius)-7, 13 myrtia (Amblyseius)-4S myrtia (Neo~eiulus)-45 N namdaphaensis (Phytoseius)-83 neococcineae (Amblyseius)-20, 25 neocorniger (Phytoseius)-84, 93 neocrotalariae (Amblyseius)-58~ 64 lleoerotalariae (Typhlodromips)-64 neojerox (Phytoseius)-84, 93 neoghanii (Amblyseius)-70 neorhenanus (Typhlodromus)-132, 133 neorykei (Amblyseius)-6, 12 Neoseiulella-lOO Neoseiulus-6, 33, 100 neosoleiger (Typhlodromus)-127, 128 neotransvaalensis (Typhlodromus)- nesbitti (Typhlodromus)-130 newsami (Amblyseius)-S2, 55 nicholsi (Amhlyseius)-20, 26 nidibu (Typhlodromus)-117 nilgiriensis (Typhlodromus)