TWO NEW RACES OF PASSERINE

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SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME «9, NUMBER IR TWO NEW RACES OF PASSERINE BIRDS FROM THAILAND BY H. G. DEIGN AN Division o{ liirds, U. S. National ^Jus^ln lafe'sf^ ^J>.^^vsi?*^'^^ (Publication 3605) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION DECEMBER 11, l<m(i

SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 99, NUMBER 18 TWO NEW RACES OF PASSERINE BIRDS FROM THAILAND BY H. G. DEIGNAN Division of Birds, U. S. National Museum (Publication 3605) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION DECEMBER 11, 1940

BALTIMORE, MD., C. 8. A.

TWO NEW RACES OF PASSERINE BIRDS FROM THAILAND By H. G. DEIGNAN ' Division of Birds, U. S. National Museum The orange-gorgeted flycatcher, Siphia strophiata Hodgson, is a regular and rather common visitor in western North Siam. winter to the higher peaks of Twenty-four specimens from Doi Angka (December), Doi Khun Tan (November), Doi Suthep (December, January, February, March), Doi Chiengdao (January), and Loi Mwe, Kengtung State (February, March) are inseparable in either sex from a series of 28 birds from the western Himalayas, Assam, Yunnan, '^ Szechuan, and Shensi, and must be called Siphia strophiata strophiata. De Schauensee reported the capture of a male on Kyu Loi, Kengtung State (February) and another on Doi Suthep (December), which had "an almost entirely black throat, in the center of which is a small concealed patch of white formed by the bases of the feathers." Since then he has received a third example from Doi Pha Hom Pok (February). In the meantime, I myself took a female on Doi Suthep (November), which differs from the female of the nominate race much as De Schauensee's three specimens differ from the male of that form. I am convinced that these birds belong to a hitherto unrecognized subspecies, which may be found to breed somewhere in the Southern Shan States. For it I propose the name SIPHIA STROPHIATA ASEMA, n. subsp. Type A female, U.S.N.M. No. 336545, taken at 5,500 feet on Doi Suthep, Chiengmai Province, western North Siam, November 7, 1936, by H. G. Deignan. The type specimen diff'ers most strikingly from the corresponding sex of both strophiata and fuscogularis (Annam) in wholly lacking a gorget. The chin, throat, and center of the upper breast are uniformly of a color which lies between buckthorn brown (Ridgway) and Isabella color (Ridgway) and which changes imperceptibly into the olivaceous-brown of the sides of the breast and the flanks ; all the ' Published with permission of the Secretary of the Smithsonian Institution. ' Proc. Acad. Nat. Sci. Philadelphia, 1934, p. 213. Smithsonian Miscellaneous Collections, Vol. 99, No. 18 I

2 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 99 feathers of the throat and breast have the concealed bases dark slategray and the concealed portion of the shaft white ; the lores, supercilia, ear-coverts, and sides of the neck are dark brownish slate, not pure slate-gray as in the other races. Three examples (two males, one female) of what I take to be strophiata in the first winter plumage are nearest to my specimen in coloration of the underparts, but all have the usual well-defined pale orange gorget with white bases of the feathers showing through. The three adult males which presumably belong here differ from the corresponding sex of strophiata and fuscogularis in having the gorget so much reduced in area, both the orange and the white portions, but especially the former, that it is almost or entirely invisible until the feathers of the breast are raised. Stresemann and Heinrich * have remarked on the fact that of seven females from Mount Victoria, two (one with an oviduct egg) wore the plumage of the male. Of my series of 45 sexed adults, 35 are labeled as males, 10 as females, and of these latter 5 are in male plumage. So great is the disproportion of the sexes in this series that I am inclined to believe that many of the "males" have been thus la1)eled by native collectors merely because of external appearances. In fuscogularis, 5 females are distinguishable from 2 males only by the slightly paler gray of the throat and breast. The allegation of the "Fauna of British India, Birds" that fuscogularis occurs in the Northern Shan States need be taken no more seriously than the inclusion of North Borneo within the range of strophiata. Fuscogularis may readily be known from strophiata by its having the upperparts, especially olivaceous-brown. the head, more rufous-brown, less A single specimen of Ixos mcclellandii, McClelland's bulbul, II from the mountains of northeastern Nan Province, is sufficiently distinct from any known form to justify erection of yet another local race of this plastic species. For it I propose the name IXOS MCCLELLANDII LOQUAX, n. subsp. Type. An adult female, U.S.N.M. No. 350105, collected on Phu Kha, 4,500 feet, Nan Province, eastern North Siam, April 14, 1936, by H. G. Deignan. Mitteil. Zool. Mus. Berlin, 1940, p. 187.

NO. l8 TWO NEW RACES OF PASSERINE BIRDS DEIGNAN 3 The brown back separates this bird from all described forms of the species except holtii, similis, and binghaml. Both geographically and in plumage it lies between binghami and holtii. From holtii it is separable by the much paler buity color of the underparts and by having the bend of the wing and the under tail-coverts buffy yellow, not buff. From binghami it is separable by the slightly deeper buffy color of the underparts, by having the entire underparts (including the center of the abdomen) suffused with this color, and by having the bend of the wing and the under tail-coverts buffy yellow, not pale yellow. Three forms of this species are now known from the higher mountains of northern Thailand. The green-backed tickcui has been taken on Doi Angka, Doi Khun Tan, Doi Suthep, Doi Rangka, and Doi Chiengdao. The brown-backed loqimx has been found only on Phu Kha, The brown-backed binghami barely enters Siam on Doi Pha Horn Pok. I have found no trace of inosculation between the brown-backed and green-backed races. Tickelli and binghami seem to be separable by no external character except color of back and each individual is definitely of one or the other form. Eight specimens from Doi Chiengdao are tickelli; from Doi Pha Hom Pok, the next locality to the north, De Schauensee has four examples, three of them binghami, the other tickelli. This last might represent an off-season wanderer but more likely indicates that, at the periphery of range, brownbacked parents tend to produce an occasional green-backed offspring. What seems to be an analogous case is offered by Porphyria albus in Siam : where the breeding-ranges of poliocephalns and viridis meet we find, nesting in juxtaposition, blue-backed individuals which must be called by the former name, others with green backs which must be given the latter name, and apparently no examples which are not clearly of one or the other form, so far as can be judged by external characters. Messrs. Delacour and Jabouille * have recently stated that similis is a synonym of holtii; this conclusion is not borne out by my material, in which seven specimens of similis are easily distinguishable from five of holtii by the deeper rufous-buff of the underparts and the darker gray of the throat. Of the previously named forms of this bulbul, I have been able to examine mcclellandii, tickelli, canescens, griseiventer, binghami, similis, and holtii, all of which I consider valid. Peracensis and the recently described centralis have not been available, but both differ * L'Oiseau et la Revue franqaise d'ornithoiogie, 1940, p. 191.

4 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 99 widely from luqua.v in having the back green. Ventralis, incidentally, was apparently compared only with mcclellandii; nothing in the description indicates how it diifers from tickelli, if at all. My thanks are due to J. H. Riley, who first brought to my attention the peculiarities of the specimen that becomes the type of Siphia strophiata asema, and to R. M. De Schauensee and the Philadelphia Academy of Natural Sciences, who have generously sent me their valuable series of the species here discussed.