Result Demonstration Report

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Result Demonstration Report 2014 Texas Quail Index Texas A&M AgriLife Extension Service Archer County Cooperator: Brad Mitchell- Mitchell and Parkey Ranches Justin B Gilliam, County Extension Agent for Archer County Becky Ruzicka, Extension Associate, Wildlife and Fisheries Department Dr. Dale Rollins, Statewide Coordinator, Reversing the Quail Decline Initiative Background and Objective The decline of bobwhite and scaled quail across their historic ranges has been an ongoing and pervasive problem for the last 20 years. Texas, which was once thought of as being the last stronghold for excellent quail populations, has experienced the same declines as the rest of the southeast in the last 20 years. The last 4 years have been some of the lowest on record (Figure 1). The Texas Quail Index (TQI) is a large-scale Texas A&M AgriLife Extension demonstration effort designed to raise awareness of quail decline in Texas by increasing community involvement and giving landowners the tools they need to successfully monitor quail on their property. Engaging in an active quail monitoring program allows landowners to objectively assess the effectiveness of their land management actions, make educated decisions on harvest quotas, and identify weak links in their habitat that could Figure 1. Texas Parks and Wildlife bobwhite quail forecast since 1978 shows the overall decline of quail. The last 4 years have been some of the lowest on record. http://www.tpwd.state.tx.us/huntwild/hunt/planning/ quail_forecast/forecast

Page 2 of 12 be improved to benefit quail. Statewide there are 36 counties participating in TQI (Figure 2) and a variety of different stakeholders, including: private land owners, Texas Parks and Wildlife (TPW) personnel, Natural Resource Conservation Service (NRCS) personnel, U. S. Fish and Wildlife Service (USFWS) personnel, and Texas Master Naturalists (TMN). Methods At the outset of the program, teams established 7-mile long permanent transects with 8 permanent mile marker locations from which to collect data for the duration of the project (Figures 3, 4). Teams were responsible for collecting data on 7 different metrics to assess the quail population level and habitat factors that affect quail populations on their site: spring call counts, dummy nests, predator surveys, habitat evaluations, roadside counts, fall covey counts, and rainfall totals. Figure 2. Active TQI participating counties shaded in black. There are 36 total statewide representing the Rolling Plains, High Plains, Edwards Plateau, South Texas Plains, Trans Pecos, and Gulf Coast Prairies ecoregions. Figure 3. Google Earth image of transect location on study in Archer County. Figure 4. Mile marker placard used on the TQI. Permanent monitoring locations are essential for comparing data across years.

Page 3 of 12 Spring Call Counts Both bobwhite and scaled (blue) quail males produce distinctive calls during the breeding season (Figure 5). The number of birds calling in a particular year is a good indication of the reproductive capital expected for that year both in terms of breeding effort and (less so) population size. Three spring call counts were conducted during May June. Participants recorded the number of calling males they heard at each mile marker over a 5-minute period. These data were then averaged to obtain an estimate of the average number of calling birds per mile marker for comparison across the transect and among sites statewide. Dummy Nests Nesting success is of critical importance to the long-term viability of quail populations. As a shortlived species with a high mortality rate, quail populations depend on a high number of new individuals being added each year in order to increase. However, suitable nesting habitat is often the weakest link in quail habitat statewide. Dummy nests are an excellent tool to evaluate the predator and nesting habitat context on their property as it pertains to quail during breeding season. Dummy nests are simply chicken eggs used to mimic an actual quail s nest placed in locations that quail would select as nest sites (Figure 6). Teams set out 4 lines of 6 dummy nests during the month of June and monitored Figure 7. Estimating the amount of nesting cover by walking a transect and counting the number of suitable nesting structures rooted within arm span. Figure 5. Calling male bobwhite quail. Photo courtesy of Russel Graves. Figure 6. Dummy nest concealed in prickly pear. Prickly pear makes an excellent nesting substrate; nests in prickly pear typically survive at a higher rate than those in grass. the nests at 14- and 28-days. Nests were recorded as intact or depredated. If the nests were depredated, teams described the eggshell remains and determined the most likely predator species based on the eggshell evidence (and related sign) left at the nest site. The dummy nest transects were also used to estimate the amount of suitable nesting cover per acre by walking a straight

Page 4 of 12 line along the transect and counting the number of suitable nesting structures rooted within one person s arm span (Figure 5). Predator Surveys In addition to dummy nests, game cameras were also used to monitor the relative abundance of predators and determine which nest predator species were present on the property. Predators can account for upwards of 80% of nest failures in ground-nesting birds. The most common types of nest predators are raccoons, skunks, and other mesomammal predators. Feral hogs can also attribute to nest failures in quail. 2 Bushnell Trophy Cam game cameras (Model 119436, Bushnell Outdoor Products, Overland, KS) were deployed for 15 days during the month of July. Cameras were set up approximately 24 inches off the ground on posts along ranch roads at a 45 degree angle to the road to provide the best field of view and to capture common nest predators as they were moving from one area to another (Figure 8). The number of separate predator observations was recorded as well as the species of predator observed. Figure 8. Camera set up at a 45 degree angle to the road to provide the best field of view and to capture common nest predators. Habitat Evaluations Teams evaluated the overall quality of habitat along the transect using a formal habitat evaluation (Bobwhite quail version available at http://wildlife.tamu.edu/files/2013/12/habitat-evaluation-bobwhite.pdf or Bobwhite Habitat Evaluation in the itunes app store; Scaled quail version available at http://wildlife.tamu.edu/files/2013/12/habitat-evaluation- Scaled.pdf). This habitat evaluation is designed to address the four main needs of quail habitat: diversity and percent of woody plants, availability and diversity of food, percent cover of suitable nesting habitat, and the interspersion of those items on the landscape. The habitat scores for each mile marker were compared to measures of quail population abundance. The limiting habitat factors highlighted by the evaluations were also quantified.

Page 5 of 12 Roadside Counts Teams conducted 3 roadside counts in September. Roadside counts are used as a measure of relative abundance for the quail population. Texas Parks and Wildlife conducts roadside counts statewide every year to prepare their annual quail forecast (see figure 1). Counts for TQI were conducted during the morning and late-afternoon hours by driving a 83 mile route at less than 20 mph on the dirt ranch roads and simply counting the number of quail observed (Figure 9). From these data, the number of quail observed per mile was calculated. This rate was compared among sites in the TQI program and could be compared among multiple years of data on the same site (provided the route remained the same). Figure 9. Male bobwhite quail observed during a roadside count. Photo by Becky Ruzicka. Fall Covey Call Counts Once bobwhite quail have concluded breeding season and returned to coveys, each covey will make a distinctive call at sunrise that can be used to count the number of coveys on the landscape. These fall covey call counts can be used as a measure of relative abundance for bobwhite quail (scaled quail do not make a covey call). Teams conducted counts by arriving at the mile marker post 40 minutes before official sunrise and listening for calling coveys until 20 mins after the last call was heard. Each mile marker was counted one time. The number of coveys calling at each mile marker was compared across the transect to the habitat evaluation score at that mile marker. Additionally, the average number of calling coveys per mile marker was compared among sites in the study. Rainfall The total amount of precipitation received during the study period was recorded from a range gage on site Quail populations, even on the highest quality habitat, are boom and bust from year to year. Much of that change in population numbers is driven by rainfall, as such it is important to take into account.

Average Roosters per Stop Page 6 of 12 Results Spring Call Counts The statewide average number of bobwhite quail roosters counted per stop was 2.6 with individual county s values ranging from 0 to 12 roosters per stop (Figure 9). The statewide average number of scaled quail counted per stop was 0.1 with values ranging from 0 to 3 roosters per stop. It is important to note that scaled quail call less frequently than bobwhites and therefore the call count index is not comparable across species. However, overall scaled quail were less prevalent than bobwhites across our study sites. The average number of bobwhite roosters per stop on the Archer County site was 0.95 with values ranging from 0 to 4 per stop (Figure 10). This puts Archer County in the 26 th percentile statewide for bobwhites. When judging the response of spring call counts, typically counts from 0-3 are considered poor, 3-6 are considered fair, and 6-9 are considered good. Counts above 9 are excellent, but it is difficult to distinguish individual quail above 9 roosters per stop. Figure 9. Statewide averages of bobwhite quail counted per stop during spring call counts. 2.5 2 1.5 1 0.5 Figure 10. Variation in bobwhite quail roosters counted per mile marker during spring call counts. 0 0 1 2 3 4 5 6 7 Mile Markers

Count Page 7 of 12 Dummy Nests Overall dummy nest survival was 50% statewide; Archer County was above the statewide average at 100% dummy nest survival (Table 1). This ranks the site in 100th percentile compared to other participating sites statewide. A good rule of thumb is that dummy nest survival over 40% indicates that the combination of nesting cover and predation pressure is not a limiting factor for success of quail nests at that location. The average number of nesting clumps per acre at the dummy nest locations in Archer County was 271. This is below the recommended threshold of 300 suitable nest sites per acre. Table 1. Dummy nest survival and potential nesting clumps per acre on the Archer county site. Dummy Nest "Survival" Nesting Clumps per Acre Mile Marker 0 100% 322 Mile Marker 1 100% 333 Mile Marker 2 100% 189 Mile Marker 4 100% 241 Overall Average 100% 271 Predator Surveys Two different species of nest predators were identified on the Archer County site (Figures 11, 12, 13). The most commonly identified predators were feral hogs. 16 14 12 10 8 6 4 2 0 Coyote Predator Species Observed Hog Figure 11. Predator species identified using game cameras.

Habitat Evaluation Score Page 8 of 12 Figure 12. Coyote captured using game cameras. Figure 13. Feral Hogs captured using game camera. Habitat Evaluations Statewide the average habitat evaluation score was 0.67 which equates to a rating of good habitat. Habitat evaluation scores from 0.01 0.24 describe poor habitat, 0.25-0.49 describe fair habitat, 0.50 0.74 describe good habitat, and 0.75 1.00 describe excellent habitat. Statewide scores ranged the full distribution from 0 to 1. In Archer County the average habitat score was 0.64 and the range of values was 0.62-0.68 (Figure 14). 1.00 0.75 0.50 0.25 0.00 0 2 5 Mile Marker Figure 14. Habitat evaluation score at each mile marker.

Frequency Page 9 of 12 In Archer County, the most commonly identified limiting habitat factors were nesting cover and woody interspersion diversity (Figure 15, 16). This is a method to identify the weak links on a property based on your habitat evaluations. The more points a factor has relative to the factors (i.e. the number of times you have identified it as a limiting factor), the more of a weakness it is on that site. 3 2 1 0 Limiting Habitat Factors Figure 15. Habitat evaluation score at each mile marker. Figure 16. Photo of Archer county site that shows the typical habitat across the site with recovering grass and a lack of woody diversity. Although, grass was present it was below the minimum size threshold for quality quail nesting cover.

Number of Coveys Average Bobwhites/ Mile Page 10 of 12 Roadside Counts The statewide average roadside count was 2 bobwhite quail per mile. The average count on the Archer County site was.38 quail per mile. This ranks the site in 26 th percentile compared to other participating sites statewide (Figure 17). 12.00 10.00 8.00 6.00 4.00 2.00 0.00 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 Figure 17. Bobwhite quail observed/mile statewide by county. Archer County ranking denoted by arrow. Fall Covey Call Counts The statewide average for fall (covey) call counts was 3.2 bobwhite coveys per mile marker. The Archer County site averaged 1.5 bobwhite coveys per mile marker and the number of coveys counted varied by mile marker (Figure 18). This ranks the site in the 18 th percentile compared to the other participating sites statewide. 4 3 2 1 0 0 1 2 3 4 5 6 7 Mile Marker Figure 18. Number of coveys counted per mile marker.

Page 11 of 12 Rainfall The total rainfall for the year as of November 22, 2014 was 21.15 inches. This was below an average of 30 inches. Overall, the Archer County study site is in Extreme drought conditions as of 11 November 2014 (Figure 19). Figure 19. U.S. Drought Monitor results for the state of Texas. Pulled on November 18, 2014.

Page 12 of 12 Discussion and Conclusions According to NOAA.GOV the five year average rainfall is 20 inches with a low in 2011 of 13 inches and a high of 2010 29 inches. This is an average of 10 inches short per year or 50 inches short on the past five years from normal. This has negatively affected the forage, cover, and water availability not only for quail but for all wildlife on site. The forcasted El Nino conditions should bring increased rainfall which will hopefully address these problems. In terms of quail habitat, the several years of below normal rainfall coupled with the extreme drought conditions we are facing has hurt nesting cover on site most. It will need time and ample rainfall to recover; grasslands are just starting to return this year. While nesting cover is very low there was still a 100% survival rate of dummy nests. This is more than likely due to low population of predators in the area: only 15 hogs and 1 coyote were caught on camera in these areas. Visually there were also very few predators during the time of the trial. We did not see any indications of predators such as raccoons or skunks in the area and only observed a handful of feral hogs and coyotes during the day while collecting the data. While quail populations seem to be up from past years there is still not an abundance of quail in the pastures. Spring call counts this year indicated renewed breeding effort from the quail on site following the drought, but numbers were still low compared to the statewide values. The ranch owner has spent time out working bird dogs and has found a few coveys with the majority of the coveys being comprised of juvenile birds confirming the breeding effort and showing that some hatches were successful this spring. However, the fall population is still low compared to the quality hunting years in the past. Given the good reproductive effort this year, timely rainfall on this site will likely set the population up for a rebound next hunting season. Acknowledgments I would like to thank Brad Mitchell, with Mitchell ranch for his time in helping to conduct the study and helping to collect data throughout the year. I would also like to thank Todd Herring and Ann Parkey for allowing us to use part of the Parkey Ranch for the demonstration. Trade names of commercial products used in this report is included only for better understanding and clarity. Reference to commercial products or trade names is made with the understanding that no discrimination is intended and no endorsement by Texas AgriLife Extension Service and the Texas A&M University System is implied. Readers should realize that results from one experiment do not represent conclusive evidence that the same response would occur where conditions vary.