G.B. Zevina ( ), O.P. Poltarukha. KEY WORDS: deep-sea fauna, European seas, Cirripedia.

Invertebrate Zoology, 2014, 11(1): 101 111 INVERTEBRATE ZOOLOGY, 2014 Deep-sea fauna of European seas: An annotated species check-list of benthic invertebrates living deeper than 2000 m in the seas bordering Europe. Cirripedia G.B. Zevina ( ), O.P. Poltarukha A.N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospect 33, Moscow, 119071, Russia. E-mail: poltarukha@rambler.ru ABSTRACT: An annotated check-list is given of Cirripedia species occurring deeper than 2000 m in the seas bordering Europe. The check-list is based on published data. The checklist includes 38 species. For each species synonymy, data on localities in European seas and general species distribution are provided. Station data are presented separately in the present thematic issue. How to cite this article: Zevina G.B., Poltarulha O.P. 2014. Deep-sea fauna of European seas: An annotated species check-list of benthic invertebrates living deeper than 2000 m in the seas bordering Europe. Cirripedia // Invertebrate Zoology. Vol.11. No.1. P.101 111. KEY WORDS: deep-sea fauna, European seas, Cirripedia. Глубоководная фауна европейских морей: аннотированный список видов донных беспозвоночных, обитающих глубже 2000 м в морях, окружающих Европу. Cirripedia Г.Б. Зевина ( ), О.П. Полтаруха A.N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospect 33, Moscow, 119071, Russia. E-mail: poltarukha@rambler.ru РЕЗЮМЕ: Приводится аннотированный список видов Cirripedia, обитающих глубже 2000 м в морях, окружающих Европу. Список основан на опубликованных данных. Список насчитывает 38 видов. Для каждого вида приведены синонимия, данные о нахождениях в европейских морях и сведения о распространении. Данные о станциях приводятся в отдельном разделе настоящего тематического выпуска. Как цитировать эту статью: Zevina G.B., Poltarulha O.P. 2014. Deep-sea fauna of European seas: An annotated species check-list of benthic invertebrates living deeper than 2000 m in the seas bordering Europe. Cirripedia // Invertebrate Zoology. Vol.11. No.1. P.101 111. КЛЮЧЕВЫЕ СЛОВА: глубоководная фауна, европейские моря, Cirripedia.

102 G.B. Zevina, O.P. Poltarukha Phylum Crustacea Class Thecostraca Subclass Cirripedia Superorder Thoracica Order Pedunculata Suborder Scalpellomorpha Family Calanticidae Genus Newmanilepas Zevina et Yakhontova, 1987 COMPOSITION: 1 species. TYPE SPECIES: Newmanilepas mirifica (Zevina, 1976). DISTRIBUTION: North Atlantic 1 (Zevina, Yakhontova, 1987 ). Newmanilepas mirifica (Zevina, 1976) Scillaelepas mirifica Zevina, 1976: 1152, fig. 2; Zevina, 1981a: 67, fig. 40. LOCALITIES: Akademik Kurchatov, St. 443 (Zevina, 1976). DISTRIBUTION: North Atlantic, DEPTH RANGE: 3120 3460 m (Zevina, Yakhontova, 1987). Genus Smilium Gray, 1825 COMPOSITION: 11 species. TYPE SPECIES: Smilium peronii Gray, 1825. DISTRIBUTION: Indo West Pacific 7, Indo West Pacific, subtropical Atlantic 1, South Atlantic 1, South Pacific 1, west Pacific 1 (Zevina, 1981a). CENTRE OF DIVERSITY: Indo West Pacific. Smilium acutum (Hoek, 1883) Scalpellum acutum Hoek, 1883: 80, pl. 3, fig. 19, pl. 8, fig. 12; Hoek, 1907a: 64, pl. VII, fig. 1; Gruvel, 1920: 12, pl. II, fig. 7; Scalpellum longirostrum Gruvel, 1902a: 70, 124, pl. 2, figs. 4, 5, pl. 3, figs. 17 20; Weltner, 1922: 70; Scalpellum hastatum Weltner, 1922: 68, Taf. II, fig. 3; (?) Scalpellum bengalense Annandale, 1906: 395; Annandale, 1916: 128, pl. VI, figs. 3, 4, pl. VII, fig. 3, pl. VIII, figs. 1 5; Scalpellum (Smilium) acutum Annandale, 1910: 154, fig. 2; Annandale, 1916: 128, pl. VIII, fig. 4; Calman, 1918: 101; Nilsson-Cantell (1921): 170, textfig. 23. LOCALITIES: Princesse-Alice and Hirondelle, St. 1116, 3119, (Gruvel, 1920); ABYPLANE St. CP 68, CP 69 (Foster, Buckeridge, 1995a); Cryos, St. BIOGAS V CP 07, EP I I CP 38, CP 39, PL 18 (Foster, Buckeridge, 1995a). DISTRIBUTION: Indo West Pacific, subtropical Atlantic, 61 2480 m (Zevina, 1981a). Family Scalpellidae Subfamily Meroscalpellinae Genus Gymnoscalpellum Newman and Ross, 1971 COMPOSITION: 5 species. TYPE SPECIES: Gymnoscalpellum tarasovi Newman and Ross, 1971. DISTRIBUTION: Antarctic Ocean (Atlantic sector) 2, northeast Atlantic 1, southeast Pacific 1, Central Pacific 1 (Zevina 1981a). Gymnoscalpellum insigne (Hoek, 1883) Scalpellum insigne Hoek, 1883: 68, pl. II, figs. 3, 4. LOCALITIES: Challenger, St. 6 (Hoek, 1883), 2775 m (single locality). DISTRIBUTION: Northeast Atlantic (Zevina, 1981a). Genus Litoscalpellum Newman, Ross, 1971 COMPOSITION: 23 species. TYPE SPECIES: Litoscalpellum korotkevitschae (Zevina 1968). DISTRIBUTION: Indo West Pacific 2; South Pacific 4; East Pacific 1; tropical West Pacific 1; North Pacific 3; Antarctic Ocean (Atlantic + Pacific sectors) 2; Indian Ocean 3; South Atlantic 2; tropical Atlantic 3; North Atlantic 2 (Zevina, 1981a, 1983; Young, Leta, 1996; Young, 1998b, 2004). Litoscalpellum anceps (Aurivillius, 1898) Scalpellum anceps Aurivillius, 1898: 191, figs. 1a c; Gruvel, 1920: 25, pl. V, figs. 1 3; Teloscalpellum anceps Zevina, 1978b: 1350. St. 652 (Aurivillius, 1898; Gruvel, 1920), 4261 m. Litoscalpellum meteoria Young, 1998b Litoscalpellum meteoria Young, 1898b: 16, fig. 12. LOCALITIES: Meteor, the following localities without station numbers: 42 55.4 N, 14 07.9 W (depth 5250 m); 42 44.5 N, 13 29.8 W (5315 m); 42 38.5 N, 13 31.0 W (5318 m); 42 04.1 N, 14 55.6 W (5275 m) (Young, 1998b). DEPTH RANGE: 5260 5318 m (only 5 localities) (Young, 1998b). Genus Neoscalpellum Pilsbry, 1907a COMPOSITION: 5 species. TYPE SPECIES: Neoscalpellum debile (Aurivillius, 1898). DISTRIBUTION: boreal Atlantic 1, Antarctic Ocean (Pacific sector) 1, Indo West Pacific 1, tropical Pacific 2 (Zevina, 1981a). Neoscalpellum debile (Aurivillius, 1898) Scalpellum debile Aurivillius, 1898: 189; Gruvel, 1905: 27; Gruvel, 1920:31, pl. V, figs. 13 15; Nilsson- Cantell, 1955: 217; Scalpellum edwardsi Gruvel, 1900a:

Deep-sea fauna of European seas. Cirripedia 103 189; Gruvel, 1902a: 63, pl. 2, figs. 3B, 16a, b; Gruvel, 1905: 28, fig. 27; Scalpellum alboranense Gruvel, 1920: 33, pl. 5, figs. 4 6; Scalpellum dicheloplax Pilsbry, 1907a: 70, fig. 28; Hoek, 1914: 4; Scalpellum dicheloplax bentophila Pilsbry, 1907a: 73, fig. 28d; (?) Scalpellum (Arcoscalpellum) dicheloplax bentophila Weltner, 1922: 67; Meroscalpellum bifurcatum Foster, Buckeridge, 1995a: 173, fig. 7A B. LOCALITIES: Talisman, St. 136 (Gruvel, 1900a), Pincesse-Alice and Hirondelle, Sts. 650, 652, 749 (Aurivillius, 1898; Gruvel, 1920); Vityaz, St. 7943 (Poltarukha, Zevina, 2006b); Meteor, St. 3 29 and localities without station numbers: 42 55.4 N, 14 07.9 W (depth 5250 m); 42 44.5 N, 13 29.8 W (5315 m); 42 38.5 N, 13 31.0 W (5318 m); 42 04.1 N, 14 55.6 W (5275 m) (Young, 1998b); Jean Charcot, Sts. 131, 202, 245, 249, 250 (Young, 1998a); BIOGAS XI CP 35 (Poltarukha, Zevina, 2006a, b); MAR-ECO, St. 54/377, 64/381 (Poltarukha, 2007). DISTRIBUTION: boreal Atlantic. DEPTH RANGE: 2120 5318 m (Zevina, 1981a; Young, 1998b). Subfamily Arcoscalpellinae Genus Amigdoscalpellum Zevina, 1978b COMPOSITION: 20 species. TYPE SPECIES: Amigdoscalpellum manum (Zevina, 1973). DISTRIBUTION: Cosmopolitan (exclusion of Arctic Ocean) 1, Indian Ocean, southwest Pacific, South Atlantic 1, Indo West Pacific 3, Indo West Pacific, North Atlantic 1, Indian Ocean 4, southeast Pacific 1, North Pacific 2, southwest Pacific 2, Atlantic Ocean 1, North Atlantic 2, South Atlantic 1, west Atlantic 1 (Zevina, 1981a; Foster, Buckeridge 1995a; Young, 2001, 2002). Amigdoscalpellum praeceps (Hoek, 1907) Scalpellum praeceps Hoek, 1907a: 114, pl. VIII, fig. 13. LOCALITIES: BIOGAS VIII, CP 30, CP 35 (Foster, Buckeridge, 1995a). DISTRIBUTION: Indo West Pacific, North Atlantic. DEPTH RANGE: 411 4720 m (Zevina, 1981a; Foster, Buckeridge, 1995a). Amigdoscalpellum rigidum (Aurivillius, 1888) Scalpellum rigidum Aurivillius, 1898: 189; Gruvel, 1905: 86; Zevina, 1976: 1155; Scalpellum striatum Gruvel, 1900a: 191; Gruvel, 1902a: 77, 133, pl. 2, fig. 3I, pl. 4, figs. 3, 5, 7, 8; Gruvel, 1905: 72, fig. 81; Gruvel, 1920: 23, pl. 2, figs. 4 6, 9 11, pl. 7, fig. 11; Nilsson- Cantell, 1955: 219; Belloc, 1959: 3; Scalpellum talismani Gruvel, 1900a: 193; Gruvel, 1902a: 86, pl. 2, figs. 3D, 6, 7; Gruvel, 1905: 86, fig. 96; Gruvel, 1920: 23; Broch, 1953: 8, fig. 4; Nilsson-Cantell, 1955: 219; Amigdoscalpellum talismani Zevina, 1978b: 1349; Zevina, 1981a: 269, fig. 203. LOCALITIES: Talisman, St. 131, 136 (Young, 2002); Pincesse-Alice and Hirondelle, St. 527, 650, 652, 753 (Aurivillius,1898; Gruvel, 1920); Meteor, St. 3 29 and localities without station numbers: 42 55.4 N, 14 07.9 W (depth 5250 m); 42 04.1 N, 14 55.6 W (5275 m); 42 38.5 N, 13 31.0 W (5318 m) (Young, 1998b); Ingolf, St. 20 (Broch, 1953; Zevina, 1981a); Jean Charcot, St. 174, 176, 202, 206, 227, 245, 251 (Young, 1998a); BENGAL 2, St. 13078/27 (Young, 2001). DEPTH RANGE: 1265 4400 m (Zevina, 1981a). Amigdoscalpellum vitreum (Hoek, 1883) Scalpellum vitreum Hoek, 1883: 115, pl. 5, fig. 14; Weltner, 1897: 251; Gruvel, 1902a: 54; Gruvel, 1905: 84, fig. 94; Nilsson-Cantell, 1955: 219; Zevina, 1973a: 137; Scalpellum formosum Hoek, 1907a: 110, pl. 8, fig. II, IIa; Stubbings, 1936: 55 (part); Nilsson-Cantell, 1938: 21; Nilsson-Cantell, 1955: 219; Tarasov, Zevina, 1957: 140, pl. 1, fig. 1; Arcoscalpellum formosum Newman, Ross, 1971: 60, pl. VIIIG, textfig. 26; Arcoscalpellum vitreum Newman, Ross, 1971: 87, pl. VIIIE, F, text-fig. 44 47; Foster (1978): 58, pl. 7D, fig. 34. LOCALITIES: Akademik Kurchatov, St. 432, (Zevina, 1976). DISTRIBUTION: cosmopolitan, excluding Arctic Ocean. DEPTH RANGE: 558 6096 m (Zevina, 1981a). Genus Arcoscalpellum Hoek, 1907 COMPOSITION: 39 species. TYPE SPECIES: Arcoscalpellum michelottianum (Seguenza, 1876). DISTRIBUTION: Pacific Ocean 10, North Atlantic 8, South Atlantic 7, North Atlantic, Indian Ocean, Indo West Pacific 1, Atlantic and Pacific Oceans 1, tropical Atlantic 3, Indian Ocean 3, Indo West Pacific 4, South Atlantic, South Indian Ocean 2 (Zevina, 1981a, 1993; Young, 1992, 1998a, 2001; Foster, Buckeridge, 1995a). Arcoscalpellum atlanticum (Gruvel, 1900) Scalpellum atlanticum Gruvel, 1900a: 190; Gruvel, 1902a: 74, pl. 2, figs. 3F, 17, 18; Gruvel, 1902b: 246; Gruvel, 1902c: 523; Gruvel, 1905: 68, fig. 76; Gruvel, 1920: 26, pl. 7, fig. 5; Hoek, 1914: 4; Belloc, 1959: 3; Teloscalpellum atlanticum Zevina, 1978b: 1350; Zevina, 1981a: 377, fig. 294; Arcoscalpellum crenulatum Foster, Buckeridge, 1995a: 170, fig. 5a f; Arcoscalpellum tritonis Young, 1998a: 19, figs. 15, 16; Young, 1998b: 36, fig. 1 [non Arcoscalpellum tritonis (Hoek, 1883)]. LOCALITIES: BIOGAS VIII CP 30 (Foster, Buckeridge, 1995a); J. Charcot, St. 251 (Young, 1998a). DEPTH RANGE: 960 3600 m (Young, 2002).

104 G.B. Zevina, O.P. Poltarukha Arcoscalpellum eponkos Young, 1998a Arcoscalpellum eponkos Young, 1998a: 36, fig. 3, 4. LOCALITIES: Jean Charcot, St. 249 (Young, 1998a). DEPTH RANGE: 4620 4690 m (single locality) (Young, 1998a). Arcoscalpellum mamillatum (Aurivillius, 1892) Scalpellum mamillatum Aurivillius, 1892: 191; Gruvel, 1905: 69; Gruvel, 1920: 21, pl. 5, figs. 19 21; Nilsson-Cantell, 1955: 218; Belloc, 1959: 3; Zevina, 1976: 1155; Amigdoscalpellum mamillatum Zevina, 1978b: 1349; Zevina, 1981a: 269, in part, not fig. 202. St. 527 (Aurivillius, 1898; Gruvel, 1920); St. 652 (Aurivillius, 1898); ECOFER 1, St. CP 01 (Young, 2001); Vityaz, St. 7943, 45 35 N, 12 35 W (Poltarukha, Zevina, 2006b). DEPTH RANGE: 3017 4900 m (Zevina, 1981a; Poltarukha, Zevina, 2006b). Arcoscalpellum michelottianum (Seguenza, 1876) Scalpellum michelottianum Seguenza, 1876: 381, pl. 6, figs. 15 25, pl. 10, fig. 26; Withers, 1953: 225; Scalpellum velutinum Hoek, 1883: 96, pl. 4, figs. 10 11, pl. 9, figs. 7 9; Weltner, 1897: 251; Weltner, 1922: 75; Gruvel, 1902a: 56, pl. 2, figs. 3c, 10a, 10b, 14, pl. 3, figs. 1, 27 31, pl. 4, figs. 6, 11 22; Gruvel, 1905: 73, fig. 83; Gruvel, 1912: 2; Hoek, 1907a: 59, 79, 87; Pilsbry, 1907a: 26, pl. 3, figs. 2, 3; Annandale, 1908: pl. 4, fig. 7; Annandale, 1911: 588; Annandale, 1913: 229; Calman, 1918: 108; Barnard, 1925: 1; Nilsson-Cantell, 1927: 743, fig. 1; Nilsson-Cantell, 1928: 4; Nilsson-Cantell, 1931: 7; Broch, 1931: 18; Stubbings, 1936: 28, fig. 12; Stubbings, 1967: 234; Tarasov, Zevina, 1957: 24; Zevina, 1972: 43, fig. 2; Scalpellum eximium Hoek, 1883: 100, pl. 4, figs. 6, 7, pl. 9, fig. 10; Weltner, 1897: 247; Scalpellum sordidum Aurivillius, 1898: 190; Scalpellum erectum Aurivillius, 1898: 192; Gruvel, 1905: 74; Scalpellum alatum Gruvel, 1900a: 192; Gruvel, 1902a: 57. St. 515 (Aurivillius, 1898; Gruvel, 1920); Akademik Kurchatov, St. 444, (Zevina, 1976); ABYPLANE St.CP 68, Jean Charcot, St. KG 229 (Foster, Buckeridge, 1995a). DISTRIBUTION: North Atlantic (72 N 34 S), Indian Ocean, Pacific Ocean. DEPTH RANGE: 64 3010 m (Zevina,1981a). Arcoscalpellum pentagonum (Nilsson-Cantell, 1955) Scalpellum pentagonum Nilsson-Cantell, 1955: 215, fig. 1. LOCALITIES: Albatross, 40 33 N, 35 24 W (Nilsson-Cantell, 1955). DISTRIBUTION: North Atlantic (single locality) (Zevina, 1981a). DEPTH RANGE: 4540 4600 m (Zevina, 1981a). Genus Catherinum Zevina, 1978b COMPOSITION: 15 species. TYPE SPECIES: Catherinum recurvitergum (Gruvel, 1902a). DISTRIBUTION: Pacific Ocean 5, Indo West Pacific 2, Indian Ocean 4, South Indian Ocean 1, North Atlantic Ocean, Indian Ocean 1, North Atlantic Ocean 2 (Zevina, 1981a; Zevina, Galkin, 1993). CENTRE OF DIVERSITY: Pacific Ocean, Indo West Pacific. Catherinum albatrossianum (Pilsbry, 1907) Scalpellum albatrossianum Pilsbry, 1907a: 54, fig. 19; Annandale, 1908: pl. VIII, fig. 10; Annandale, 1913: 232; Annandale, 1916: pl. VI, fig. 9 (non Nilsson- Cantell, 1925); Stubbings, 1936: 66; Broch, 1953: 6, fig. 3; Zevina, 1973a: 847. LOCALITIES: not exactly known; presence in the deep north-east Atlantic mentioned in (Zevina, 1981a). DISTRIBUTION: North Atlantic, Indian Ocean. DEPTH RANGE: 760 4180 m (Zevina, 1981a). Catherinum recurvitergum (Gruvel, 1900) Scalpellum recurvitergum Gruvel, 1900a: 190; Gruvel, 1902a: 67, pl. 2, figs. 3H, 21, 22; Gruvel, 1905: 49, fig. 54; Weltner, 1922: 72; Zevina, 1976: 1155. LOCALITIES: Talisman, St. 118 (Gruvel, 1900a), Jean Charcot, Sts. 202, 245 (Young, 1998a); ECOFER 1, St. CP 01, CP 02 (Young, 2001); Vityaz, St. 7943 (Poltarukha, Zevina, 2006b); Akademik Mstislav Keldysh, St. 500 (Poltarukha, Zevina, 2006b); MAR-ECO, St. 52/374, 64/381, 72/386 (Poltarukha, 2007). DISTRIBUTION: North Atlantic, west Indian Ocean. DEPTH RANGE: 1289 4297 m (Zevina, 1981a; Young, 1998a). Genus Planoscalpellum Zevina, 1978 COMPOSITION: 5 species. TYPE SPECIES: Planoscalpellum planum (Hoek, 1883). DISTRIBUTION: South Indian Ocean 1, Indo West Pacific 1, North Atlantic 1, Pacific Ocean 1, west Indian Ocean 1 (Zevina, 1981a; Shreider, 1994). Planoscalpellum limpidus (Zevina, 1976) Scalpellum limpidus Zevina, 1976: 1152, fig. 2. LOCALITIES: Meteor, St. 3-29 and localities without station numbers: 42 55.4 N, 14 07.9 W

Deep-sea fauna of European seas. Cirripedia 105 (depth 5250 m); 42 04.1 N, 14 55.6 W (5275 m) (Young, 1998b); Akademik Kurchatov, St. 431 (Zevina, 1976); Jean Charcot, Sts. 245, 249 (Young, 1998a); Akademik Mstislav Keldysh, St. 390-3, (Poltarukha, Zevina, 2006a). DEPTH RANGE: 4270 5900 m (Zevina, 1981a; Young, 1998a). Genus Teloscalpellum Zevina, 1978b COMPOSITION: 20 species. TYPE SPECIES: Teloscalpellum spicatum (Zevina, 1975). DISTRIBUTION: Indo West Pacific 6, Indian Ocean 2, Pacific Ocean 1; South Pacific 1, West Pacific 1; East Pacific 2; tropical Pacific 1; North Atlantic, Indo West Pacific 1; east Atlantic 2; west Atlantic 3 (Zevina, 1981a; Young, 2001; 2004). Teloscalpellum luteum (Gruvel, 1900) Scalpellum luteum Gruvel, 1900a: 192; Gruvel, 1902a: 80, 135, pl. 2, figs. 3E, 11, pl. 4, figs. 4, 9, 10; Gruvel, 1905: 84, fig. 93. LOCALITIES: Talisman, St. 118 (Gruvel, 1900a), Jean Charcot, St. 129 (Young, 1998a). DEPTH RANGE: 3000 3175 m (Zevina, 1981a; Young, 1998a). Genus Trianguloscalpellum Zevina, 1978b COMPOSITION: 23 species. TYPE SPECIES: Trianguloscalpellum balanoides (Hoek, 1883) DISTRIBUTION: Indo West Pacific 9, Pacific 1, West Pacific 2; East Pacific 1; boreal Pacific 1; South Indian Ocean 2; North Atlantic 2; South Atlantic 2; west Atlantic 1; Tropical Atlantic 1; Antarctic Ocean (Atlantic and Indian sectors) 1 (Zevina, 1981a; Young, Leta, 1996; Young, 1998a, b). CENTRE OF DIVERSITY: Indo West Pacific. Trianguloscalpellum ovale (Hoek, 1883) Scalpellum regium Wyvile Thomson, 1873: 347 (in part, only figs. 1, 2); Wyvile Thomson, 1877: (in part, only figs, 2, 3); Pilsbry, 1907a: (in part, only pl. III, fig. 4); Scalpellum regium var. ovale Hoek, 1883: 109, pl. V, figs. 5, 6; Scalpellum gigas Gruvel, 1902a: 52, 126, pl. II, fig. 3A, pl. III, figs. 22 26 [non Scalpellum gigas Hoek, 1883]; Trianguloscalpellum regium ovale Zevina, 1981a: 311, fig. 235; Trianguloscalpellum gigas Foster, Buckeridge, 1995a: 169, fig. 2E [non Scalpellum gigas Hoek, 1883]. LOCALITIES: Meteor, localities without station numbers: 45 48.6 N, 17 15.8 W (depth 4740 m); 45 49.8 N, 17 14.2 W (depth 4730 m); 42 55.4 N, 14 07.9 W (depth 5250 m); 42 44.5 N, 13 29.8 W (5315 m); 42 38.5 N, 13 31.0 W (5318 m); 42 04.1 N, 14 55.6 W (5275 m) (Young, 1998b); Cryos, St. CP 42 (Foster, Buckeridge, 1995a); Jean Charcot, St. 249, (Young, 1998a); Discovery St. 13368#53 (Young, 2001); Akademik Mstislav Keldysh, St. 503 (Poltarukha, Zevina, 2006b). DEPTH RANGE: 4620 5211 m (Zevina, 1981a; Young, 1998a). Trianguloscalpellum regium (Wyvile Thomson, 1873) Scalpellum regium regium Wyvile Thomson, 1873: 347 [in part, not figs. 1, 2 = Trianguloscalpellum ovale (Hoek, 1883)]; Wyvile Thomson, 1877: 4 [in part, not figs. 2, 3 = Trianguloscalpellum ovale (Hoek, 1883)]; Hoek, 1883: 106: pl. 4 figs. 3 5; Hoek, 1884: 10; Pilsbry, 1907a: 28, pl. 3 figs. 4, 5 [in part, not pl. III, fig. 4 = Trianguloscalpellum ovale (Hoek, 1883)]; Gruvel, 1912: 2; Gruvel, 1920: 30, pl. 1 fig. 7; Scalpellum molle Aurivillius, 1898: 191; Gruvel, 1905: 77; Gruvel, 1920: 29, pl. V, figs. 10 12; Nilsson-Cantell, 1955: 218; Belloc, 1959: 3; Zevina, 1976: 1155; Trianguloscalpellum regium regium Zevina, 1981a: 309, fig. 234; non Scalpellum regium Zevina, 1970: 263, pls. 10, 11; Zevina, 1973ñ: 136 [=Trianguloscalpellum gigas (Hoek, 1883)]; not Trianguloscalpellum regium Zevina, 1981b: 86; Foster, Buckeridge, 1995b: 355, fig. 6K [=Trianguloscalpellum? mollucanum (Hoek, 1883)]. St. 527, 3006 (Aurivillius, 1898; Gruvel, 1920); Meteor, St. 3-29 (Young, 1998b); BIO- GAS VII CP 28, CP 30 (Foster, Buckeridge, 1995a); Discovery St. 13078#27, 13200/93, 13627#11 (Young, 2001); Vityaz, St. 7943 (Poltarukha, Zevina, 2006b); Jean Charcot, St. 129 (Young, 1998a); Jean Charcot, St. 202, 245, 251 (Young, 1998a); Akademik Mstislav Keldysh, St. 500 (Poltarukha, Zevina, 2006b); MAR-ECO, St. 40/367 (Poltarukha, 2007). DISTRIBUTION: boreal and tropical Atlantic, Pacific, Indian Ocean. DEPTH RANGE: 1507 6135 m (Zevina, 1981a). Genus Verum Zevina, 1978 COMPOSITION: 25 species. TYPE SPECIES: Verum zenkevitchi (Zevina, 1972). DISTRIBUTION: Arctic Ocean, North Atlantic 1, southeast Atlantic, southwest Indian Ocean 4, North Atlantic Ocean 3, Indo West Pacific 6, South Pacific, west Indian Ocean, North Atlantic 4, North Pacific 1, All tropical seas 1, southeast Pacific 1, Pacific Ocean 1, Indian Ocean 2, unknown 1 (Zevina, 1981a; Foster, Buckeridge, 1995a; Young, 1998b). CENTRE OF DIVERSITY: Indo West Pacific.

106 G.B. Zevina, O.P. Poltarukha Verum carinatum (Hoek, 1883) Scalpellum carinatum Hoek, 1883: 76, pl. III, figs. 7, 8; Pilsbry, 1907a:53, fig. 18; Gruvel, 1920: 20; Barnard, 1925: 3; Broch, 1953: 7; Scalpellum imperfectum Pilsbry, 1907a: 75, fig. 30, pl. IV, figs. 15 18; Barnard, 1924: 47; Broch, 1953: 9. St. 515 (Aurivillius, 1898; Gruvel, 1920). DISTRIBUTION: Atlantic Ocean, 37 N 34 S, 73 W 17 E. DEPTH RANGE: 1102 2234 m (Zevina, 1981a). Verum frillosum Foster et Buckeridge, 1995 Verum frillosum Foster, Buckeridge, 1995a: 174, fig. 8 LOCALITIES: BIOGAS VII CP 28, (Foster, Buckeridge, 1995a). DISTRIBUTION: central Atlantic region, 3380 m (single locality) (Foster, Buckeridge, 1995a). Verum minutum (Hoek, 1883) Scalpellum minutum Hoek, 1883: 113, pl. V, fig. 12; Stubbings, 1936: 28. LOCALITIES: ABYPLANE St. CP 68, CP 69 (Foster, Buckeridge, 1995a). DISTRIBUTION: South Pacific, west Indian Ocean, subtropical east Atlantic Ocean. DEPTH RANGE: 802 2639 m (Zevina, 1981a; Foster, Buckeridge, 1995a). Verum parazelandiae Young, 1998 Scalpellum novae-zelandiae Gruvel, 1902a: 54, pl. II, fig. 12, 13, 15; Verum novaezelandiae Foster, Buckeridge, 1995a: 174, fig. 7C E (in part: specimens from Atlantic). LOCALITIES: ABYPLANE St. CP 38, CP 39 (Foster, Buckeridge, 1995a). DEPTH RANGE: 882 4850 m (Young, 1998b). Verum striolatum (G.O. Sars, 1877) Scalpellum striolatum G. Sars, 1877: 364; Tarasov, 1936: 46; Tarasov, 1937: 45; Tarasov, 1945: 54, pl. 1, fig. 3; Broch, 1953: 5, fig. 2; Tarasov, Zevina, 1957: 138, figs. 10, 42, 43; Zevina, Tarasov, 1964: 232, figs. 4 6; Scalpellum albatrossianum Nilsson-Cantell, 1925: 7, fig. 2; Nilsson-Cantell, 1926: 2. LOCALITIES: Ingolf, St. 113 (Broch, 1953); North Pole-4, 76 47 N, 173 17 E, (Zevina, 1981a); North Pole-22, St. 58, (Zevina, 1981a); Marvel, St. PL 1196 (Young, 2001). DISTRIBUTION: Arctic Ocean, North Atlantic. DEPTH RANGE: 348 3941 m (Zevina, 1981a). Genus Weltnerium Zevina, 1978 COMPOSITION: 13 species. TYPE SPECIES: Weltnerium nymphocola (Hoek, 1883). DISTRIBUTION: Arctic Ocean, Subarctic 1, Antarctic Ocean (Atlantic, Indian, Pacific sectors) 1, South Atlantic 5, North Atlantic 2, Indo West Pacific 2, Sea of Okhotsk 1, unknown 1 (Zevina, 1981a; Young, 2000, 2002). CENTRE OF DIVERSITY: not determined. Weltnerium pusillum (Aurivillius, 1898) Scalpellum pusillum Aurivillius, 1898: 194; Gruvel, 1920: 24, pl. 6, fig. 10. St. 188 (Aurivillius, 1898; Gruvel, 1920). DISTRIBUTION: North Atlantic Ocean. DEPTH RANGE: 1267 2000 m (Zevina, 1981a). Suborder Lepadomorpha Family Poecilasmatidae Genus Dichelaspis Darwin, 1852 COMPOSITION: 5 species. TYPE SPECIES: Dichelaspis orthogonia Darwin, 1852. DISTRIBUTION: Indo West Pacific 1, North Atlantic, Indian Ocean 1, North Atlantic 2, Pacific Ocean 1 (Zevina, 1982; Young, 1998b). Dichelaspis thieli Young, 1998 Dichelaspis thieli Young, 1998b: 7, figs. 5, 6. LOCALITIES: SEAMOUNT 2, St. TS 267 (Young, 2001). DEPTH RANGE: 1300 2235 m (Young, 2001). Dichelaspis sp. LOCALITIES: SEAMOUNT 2, St. CP 268 (Young, 2001). DEPTH RANGE: 2145 2205 m (Young, 2001). Genus Glyptelasma Pilsbry, 1907a COMPOSITION: 12 species. TYPE SPECIES: Glyptelasma subcarinatum (Pilsbry, 1907 a). DISTRIBUTION: Cosmopolitan 1, tropical Pacific, Atlantic and Indian Oceans 1, Indo West Pacific 3, Indo West Pacific, northwest Atlantic 1; Indian Ocean 2, South Pacific 1, northwest Atlantic 2, northwest Atlantic, Indian Ocean 1 (Zevina, 1982, 1990; Zevina, Shreider, 1992). CENTRE OF DIVERSITY: Indo West Pacific. Glyptelasm carinatum (Hoek, 1883) Poecilasma carinatum Hoek, 1883 : 44, pl. 1, figs. 8 10, pl. 2, fig. 1, pl. 7, figs. 6 7; Poecilasma (Glyptelasma) carinatum Nilsson-Cantell, 1921: 258; Glyptelasma carinatum Broch, 1931: 32, fig. 11; Megalasma (Glyptelasma) carinatum Pilsbry, 1907a: 93; Pilsbry, 1907b: 416; Calman, 1918: 401, figs. 1 3; Barnard, 1924: 54; Zevina, 1982: 92, fig. 82.

Deep-sea fauna of European seas. Cirripedia 107 St. 1116 (Gruvel, 1920); Seamount 2, St. TS 267 (Young, 2001). DISTRIBUTION: cosmopolitan. DEPTH RANGE: 60 2865 m (Zevina, 1982). Glyptelasm hamatum (Calman, 1919) Megalasma (Glyptelasma) hamatum Calman, 1919: 370, figs. 5 7; Nilsson-Cantell, 1927: 770, fig. 12; Nilsson-Cantell, 1928: 23, fig. 11; Nilsson-Cantell, 1931: 10; Nilsson-Cantell, 1934: 49; Nilsson-Cantell, 1955: 219; Weisbord, 1979: 48, pl. 5, figs. 1, 2, pl. 14, fig. 3; Zevina, 1982: 93, fig. 83; Megalasma carinatum Foster, 1978: 26, pl. 3b, fig. 12 [not Megalasma carinatum (Hoek, 1883)]. LOCALITIES: Jean Charcot, St. 202, 227 (Young, 1998a); SEAMOUNT 2, St. TS 267 (Young, 2001). DISTRIBUTION: tropical Pacific, Atlantic and Indian Oceans. DEPTH RANGE: 366 3660 m (Zevina, 1982). Suborder Heteralepadomorpha Family Heteralepadidae Genus Heteralepas Pilsbry, 1907 COMPOSITION: 23 species. TYPE SPECIES: Heteralepas rex (Pilsbry, 1907a). DISTRIBUTION: Indo West Pacific 5, west Pacific 4, central Pacific 1, east Pacific 3, South Pacific 2, Indian Ocean 1, Atlantic Ocean, Pacific Ocean 1, Atlantic Ocean 6 (Zevina, Kolbasov, 2000; Young, 2001). CENTRE OF DIVERSITY: Indo West Pacific. Heteralepas segonzaci Young, 2001 Heteralepas segonzaci Young, 2001: 711 715, figs. 4 5. LOCALITIES: SEAMOUNT 2, St. TS 267, 2235 m (single locality) (Young, 2001). Order Sessilia Suborder Verrucomorpha Family Verrucidae Genus Altiverruca Pilsbry, 1916 COMPOSITION: 34 species. TYPE SPECIES: Altiverruca hoeki (Pilsbry, 1907). DISTRIBUTION: cosmopolitan 2, Indo West Pacific 5, west Pacific 9, central Pacific 3, east Pacific 2, boreal Pacific 1, South Atlantic 3, northwest Atlantic 3, North Atlantic 6 (Aurivillius, 1898; Hoek, 1913; Pilsbry, 1916; Nilsson-Cantell, 1929, 1955; Rao, Newman, 1972; Zevina, 1975, 1987a, b, 1988, 1990; Rosell, 1991; Zevina, Galkin, 1992; Buckeridge, 1994, 1997; Young, 1998a). CENTRE OF DIVERSITY: west Pacific. Altiverruca erecta (Gruvel, 1900) Verruca erecta Gruvel, 1900b: 243. LOCALITIES: Talisman, St. 118 (Gruvel, 1900b); Marvel, St. PL 1196 (Young, 2001). DEPTH RANGE: 2295 3175 m (Gruvel, 1900b; Nilsson-Cantell, 1955; Young, 2001). Altiverruca longicarinata (Gruvel, 1900) Verruca longicarinata Gruvel, 1900b: 242. LOCALITIES: Akademik Mstislav Keldysh, St. 3452, 3956, 3959, 3982 (Poltarukha, Zevina, 2006b). DEPTH RANGE: 2400 3432 m (Nilsson-Cantell, 1955; Poltarukha, Zevina, 2006b). Altiverruca obliqua (Hoek, 1883) Verruca obliqua Hoek, 1883: 143, pl. 12, figs. 15 17; Hoek, 1907b: 9; Weltner, 1897: 274; Gruvel, 1905: 173, fig. 191; Verruca (Altiverruca) obliqua Pilsbry, 1916: 40; Altiverruca vertica Foster, Buckeridge, 1995a: 180, fig. 14. LOCALITIES: Challenger, St. 6 (Hoek, 1883). DISTRIBUTION: Northeast Atlantic. DEPTH RANGE: 1182 2775 m (Hoek, 1883; Foster, Buckeridge, 1995a; Young, 1998a). Altiverruca quadrangularis (Hoek, 1883) Verruca quadrangularis Hoek, 1883: 140, pl. 11, figs. 10, 11, pl. 12, figs. 8 12. St. 3119 (Gruvel, 1920); Marvel, St. PL 1196 (Young, 2001). DISTRIBUTION: Cosmopolitan. DEPTH RANGE: 2380 3475 m (Young, 1998a). Genus Costatoverruca Young, 1998a COMPOSITION: 13 species. TYPE SPECIES: Costatoverruca alba (Pilsbry, 1907). DISTRIBUTION: northwest Atlantic 6, northwest Atlantic, Indo West Pacific 1, west Atlantic 1, northeast Atlantic 2, east Indian Ocean 1, west Indian Ocean 1, west Pacific 1 (Gruvel, 1912; Nilsson-Cantell, 1929; Weisbord, 1979; Foster, Buckeridge, 1995b; Buckeridge, 1997; Young, 1998a, 2000). CENTRE OF DIVERSITY: northwest Atlantic. Costatoverruca grimaldi (Gruvel, 1912) Verruca grimaldi Gruvel, 1912: 5; Gruvel, 1920: 48, pl. 2, figs. 14, 15, pl. 3, figs. 5, 6.

108 G.B. Zevina, O.P. Poltarukha St. 3119 (Gruvel, 1920), 2380 m. DISTRIBUTION: northeast Atlantic (Gruvel, 1920). Genus Metaverruca Pilsbry, 1916 COMPOSITION: 17 species. TYPE SPECIES: Metaverruca coraliophila (Pilsbry, 1916). DISTRIBUTION: Indo West Pacific 2, southwest Pacific 6, Indo West Pacific, northeast Atlantic, northwest Atlantic 1, tropical Indian Ocean 3, tropical and subtropical Atlantic 5 (Aurivillius, 1898; Gruvel, 1911, 1920; Broch, 1931; Zevina, 1971, 1987a, 1990; Buckeridge, 1994, 1997; Foster, Buckeridge, 1995a, 1995b; Young, 1998a, 1998b). CENTRE OF DIVERSITY: Indo West Pacific. Metaverruca aequalis (Aurivillius, 1898) Verruca aequalis Aurivillius, 1898: 196; Gruvel, 1905: 176; Gruvel, 1920: 42, pl. 5, figs. 28 29, pl. 6, figs. 6 7; Hoek, 1907b: 9; Belloc, 1959: 4. LOCALITIES: Jean Charcot, St. 171, 174 (Young, 1998a). DISTRIBUTION: northeast Atlantic. DEPTH RANGE: 1022 3215 m (Aurivillius, 1898; Gruvel, 1920; Young, 1998a). Metaverruca recta (Aurivillius, 1898) Verruca recta Aurivillius, 1898: 195; Verruca sculpta Aurivillius, 1898: 197; Verruca linearis Gruvel, 1900b: 243; Gruvel, 1902a: 107, pl. 5, figs. 11, 12; Verruca magna Gruvel, 1901: 261; Verruca halotheca Pilsbry, 1907b: 188, pl. 12 figs. 9, 10; Verruca capsula Hoek, 1907a: 130, pl. 12, figs. 1-3, pl. 13, figs. 1 4; Verruca coraliophila Pilsbry, 1916: 21, pl. 1, figs. 1 5; Verruca cookei Rosell, 1991: 299, pl. 11 figs. r, s, u, v (non Verruca cookei Pilsbry, 1927). LOCALITIES: ABYPLANE St. CP 38 (Foster, Buckeridge, 1995a). DISTRIBUTION: Indo West Pacific, North Atlantic. DEPTH RANGE: 240 2100 m (Foster, Buckeridge, 1995a; Young, 1998a). References Annandale N. 1905. Malaysian barnacles in the Indian Mus. with a list of Indian Pedunculata // Mem. Asiatic Soc. Bengal. Vol.1. No.5. P.73 84. Annandale N. 1906. Nat. Hist. Notes from the R.I.M.S. Ship Investigator. Ser. III. 12. Preliminary Rep. on the Indian Stalked Barnacles // Ann. Mag. Nat. Hist. Vol.7. No.17(50). P.389 400. Annandale N. 1908. Crustacea Entomostraca. Illustrations of the Zoology of R.I.M.S. Investigator. Government Printer. Calcutta. Annandale N. 1909. An account of the Indian Cirripedia Pedunculata. Pt. I. Family Lepadidae // Mem. Indian Mus. Vol.2. No.2. P.60 138. Annandale N. 1910. The Indian Barnacles of the subgenus Smilium with remarks on the classification of the genus Scalpellum // Rec. Indian Mus. Vol.5. P.145 155. Annandale N. 1911. Some barnacles of the genus Scalpellum from Irish Seas // Ann. Mag. Nat. Hist. Vol.8. No.7. P.588 590. Annandale N. 1913. The Indian Barnacles of Subgenus Scalpellum // Rec. Indian Mus. Vol.9. No.4. P.227 236. Annandale N. 1916. Three plates to illustrate the Scalpellidae and Iblidae of Indian Seas with synonymy and notes // Mem. Indian Mus. Vol.6. No.2. P.127 131. Aurivillius C.W.S. 1892. Neue Cirripeden aus dem Atlantischen, Indischen und Stillen Ocean // Öfvers. Öfv. Kongl. Sv. Vetenskapsakademiens Förhand. N.S. Bd.3. S.123 134. Aurivillius C.W.S. 1898. Cirripèdes nouveaux provenant des Campagnes scientifique de S.A.S. le Prince de Monaco // Bull. Soc. Zool. France. Vol.23. P.189 198. Barnard K.H. 1924. Contributions to the Crustacean fauna of South Africa ¹ 7 Cirripedia // Ann. South African Mus. Vol.20. No.1. P.1 103. Barnard K.H. 1925. Rep. on a collection of Cirripedia (barnacles) from South African waters // Rep. Fish. Mar. Biol. Surv. Union of South Africa. Vol.4. No.6. P.1 5. Belloc G. 1959. Catalogue des types de Cirrhipèdes du Musée océanographique de Monaco // Bull. Inst. Oceanogr. Monaco. Vol.1157. P.1 7. Broch H. 1931. Papers from Dr. Th. Mortensen s Pacific Expedition 1914 16. LVI. Indomalayan Cirripedia // Vidensk. Medd. Dansk Naturhist. Foren. Vol.91. P.1 146. Broch. H. 1953. Cirripedia Thoracica // Dan. Ing. Exped. Vol.3. No.14. P.1 17. Buckeridge J.S. 1994. Cirripedia Thoracica: Verrucomorpha of New Caledonia, Indonesia Wallis and Futuna Islands // Crosnier A. (ed.). Résultats Campagnes MUSORSTOM. T.12. Mémoires. Mus. Nat. Hist. Nat. Vol.161. P.87 125. Buckeridge J.S. 1997. Cirripedia Thoracica: new ranges and species of Verrucomorpha from the Indian and Southwest Pacific Oceans // Crosnier A. (ed.). Resultats des Campagnes MUSORSTOM. T.18. Mémoires. Mus. Nat. Hist. Nat. Vol.176. P.125 149. Calman W.T. 1918. On Barnacles of the Genus Scalpellum from Deep-sea Telegraph-Cables // Ann. Mag. Nat. Hist. Vol.9. No. 1(1). P.96 124. Calman W.T. 1919. On Barnacles of the Genus Megalasma from Deep-sea Telegraph-Cables // Ann. Mag. Nat. Hist. Vol.9. No.4(24). P.361 374. Darwin C.A. 1852. Monograph on the Subclass Cirripedia with figures of all the species. The Lepadidae; or pedunculate cirripedes. London. 400 p. Darwin C.A. 1854. Monograph on the Subclass Cirripedia with figures of all the species. The Balanidae, the Verrucidae etc. London. 684 p. Foster B.A. 1978. The Marine fauna of New Zealand: Barnacles (Cirripedia: Thoracica) // Mem. New Zealand Oceanogr. Inst. Vol.69. 160 p.

Deep-sea fauna of European seas. Cirripedia 109 Foster B.A. Buckeridge J.S. 1995a. Barnacles (Cirripedia: Thoracica) of seas off the Straits of Gibraltar // Bull. Mus. Nat. Hist. Nat. Paris. Vol.4. No.17. Ser.A. (1 2). P.163 192. Foster B.A., Buckeridge J.S. 1995b. Barnacles (Cirripedia: Thoracica) of seas off Réunion Island and East Indies // Bull. Mus. Nat. Hist. Nat. Paris. Vol.4. No.16.(A). P.345 382. Gray J.E. 1825. A synopsis of the genera of cirripedes arranged in natural families with a description of some new species // Ann. of Philosophy. Vol.2. P.97 107. Gruvel A. 1900a. Sur une espèces nouvelles du genre Scalpellum provenant de la collection du Muséum de Histoire Naturelle de Paris // Bull. Mus. Nat. Hist. Nat. Paris. T.6. P.189 194. Gruvel A. 1900b. Sur les espèces nouvelles appartenant au genre Verruca provenant de la campagne du Talisman // Bull. Mus. Nat. Hist. Nat. Paris. T.6. P.242 244. Gruvel A. 1901. Diagnoses de quelques espèces nouvelles de Cirrhipèdes // Bull. Mus. Nat. Hist. Nat. Paris. T.6. P.256 263. Gruvel A. 1902a. Cirrhipèdes // Expéditions Scientifiques du Travailleur et du Talisman. 1880 1883. Paris. P.1 178. Gruvel A. 1902b. Revision des Cirrhipèdes appurtenant a la collection du Muséum de History Naturelle // Nouv. Arch. Mus. Hist. Nat. T.4. No.4(2). P.215 312. Gruvel A. 1902c. Catalogue des Cirrhipèdes appurtenant a la collection du Muséum // Bull. Mus. Nat. Hist. Nat. Marseille. T.8. P.522 566. Gruvel A. 1905. Monographie des Cirrhipèdes ou Thecostraces. Paris. 472 p. Gruvel A. 1911. Sur deux espèces nouvelles de Cirrhipèdes appartenant a la collection du Mus. // Bull. Mus. Nat. Hist. Nat. Paris. T.17. No.5. P.290 291. Gruvel A. 1912. Note preliminaire sur les Cirrhipèdes recueillis pendant les campagnes de SAS le Prince de Monaco // Bull. Inst. Oceanogr. Paris. Vol.241. P.1 7. Gruvel A. 1920. Cirrhipèdes provenant des campagnes scientifiques de S.A.S. le Prince de Monaco 1885 1913 // Rés. Camp. Sci. acc. par le Prince Albert I er de Monaco. T.53. P.1 89. Hoek P.P.C. 1883. Report on the Cirripedia collected by H.M.S. Challenger during the years 1873 1876 // Rep. sci. res. of the voyage of H.M.S. Challenger during the years 1873 76. Vol.8. No.25. P.1 169. Hoek P.P.C. 1884. Report on the Cirripedia collected by H.M.S. Challenger during the years 1873 1876. Anatomical part // Rep. sci. res. of the voyage of H.M.S. Challenger during the years 1873 76. Vol.10. No.28. P.1 47. Hoek P.P.C. 1907a. The Cirripedia of the Siboga Expedition. A. Cirripedia Pedunculata // Siboga-Expeditie. Monogr. 31a. P.1 127. Hoek P.P.C. 1907b. Cirripedia. Resultats du Voyage S.Y. Belgica // Rap. Scie. Zoologie. P.3 9. Hoek P.P.C. 1913. The Cirripedia of the Siboga-Expedition. Cirripedia. Sessilia // Siboga-Expeditie. Monogr. 31b. P.129 275. Newman W.A. 1987. Evolution of cirripedes and their major groups // Barnacle Biology. Rotterdam. P.3 42. Newman W.A., Ross A. 1971. Antarctic Cirripedia // Ant. Res. Series. Vol.14. P.1 209. Nilsson-Cantell C.A. 1921. Cirripedien. Studien zur Kenntniss der Biologie. Anatomie und Systematik der Gruppe // Zool. Bidrag fran Uppsala. Bd.7. S.75 395. Nilsson-Cantell C.A. 1925. Neue und wenig bekannte Cirripedien aus dem Museum zu Stockholm und zu Upsala // Ark Zool. Stockholm. Bd.18. No.A(3). S.1 46. Nilsson-Cantell C.A. 1926. Antarktische und subantarktische Cirripedien // Ark. Zool. Stockholm. Bd.18. No.A(27). S.1 16. Nilsson-Cantell C.A. 1927. Some barnacles in the British Mus. (Nat. Hist.) // Proc. Zool. Soc. London. Vol.11. P.743 790. Nilsson-Cantell C.A. 1928. Studies on cirripeds in the British Mus. (Nat. Hist.) // Ann. Mag. Nat. Hist. London. Vol.10. No.2(7). P.1 99. Nilsson-Cantell C.A. 1929. Cirripedien des Genus Verruca der Deutschen Tiefsee-Expedition auf dem Dampfer Valdiva 1898 1899 // Zool. Jahrbücher. Vol.58. S.459 480. Nilsson-Cantell C.A. 1931. Cirripeds from the Indian Ocean and Malay Archipelago in the British Museum (Nat. Hist.) // Ark. Zool. Stockholm. Vol.23. No.A(18). P.1 12. Nilsson-Cantell C.A. 1934. Cirripeds from the Indian Ocean and Malay Archipelago in the Zoological Museum of Amsterdam // Zool. Mededelingen, Leiden. Vol.17. P.31 63. Nilsson-Cantell C.A. 1938. Cirripeds from Indian Ocean in the collection of the Indian Museum, Calcutta // Indian Mus. Mem. Vol.13. No.1. P.1 81. Nilsson-Cantell C.A. 1955. Cirripedia // Reports of the Swedish Deep-Sea Expedition. Vol.2. Zool. No.17. P.215 220. Pilsbry H.A. 1907a. The barnacles (Cirripedia) contained in the collections of the U.S. Nat. Mus. Washington // Bull. U.S. Nat. Mus. Vol.60. 122 p. Pilsbry H.A. 1907b. Hawaiian Cirripedia // Bull. Bureau of Fish. Washington. Vol.26. P.181 190. Pilsbry H.A. 1916. The sessile barnacles (Cirripedia) collected by the United States Natural Museum, including a monograph of the American species // Bull. U.S. Nat. Mus. Vol.93. P.1 366. Poltarukha O.P. 2007 [Deep-sea barnacles (Cirripedia, Thoracica) collected by MAR-ECO expedition in North Atlantic] // Bulleten Moskovskogo Obschestva Ispitateley Prirody. Otdel biol. Vol.112. No.4. P.18 22 [in Russian, with English summary]. Poltarukha O.P., Zevina G.B. 2006a. Barnacles (Cirripedia, Thoracica) of the Reykjanes Ridge // Biogeography of the North Atlantic seamounts. Moscow. P.152 161. Poltarukha O.P., Zevina G.B. 2006b. Barnacles (Cirripedia Thoracica) of the north-eastern Atlantic // Biogeography of the North Atlantic seamounts. Moscow. P.162 176. Rao L.M.V., Newman W.A. 1972. Thoracic Cirripedia from Guyots of the Mid-Pacific Mountains // Trans. San Diego Soc. Nat. Hist. Vol.17. No.6. P.69 94. Rosell N.C. 1991. Crustacea Cirripedia Thoracica: Musorstom 3 Philippines collection // Res. Campagnes Musorstom. Mém. Mus. Nat. Hist. Nat. T.9. No.A(152). P.9 61. Seguenza G. 1876. Ricerchi Paleontologiche intorno ai Cirripedi terziari della provincia de Messina. Con

110 G.B. Zevina, O.P. Poltarukha appendice intorno ai Cirripedi viventi nel Mediterraneo e sui fossili terziari dell Italia Meridonale. Parte. II. Lepadidi. Atti della Accademia Pontaniana. Vol.10. P.267 481. Shreider Ì.U. 1994. [Benthic Lepadomorpha of the western Indian Ocean] // Trudy Instituta Oceanologii. Vol.129. P.156 164 [in Russian, with English summary]. Stubbings H.G. 1936. Cirripedia // Sci. Rep. John Murray Exped. 1933 34. Vol.4. No.1. P.1 70. Stubbings H.G. 1967. The cirriped fauna of tropical West Africa // Bull. British Mus. (Nat. Hist.). Zoology. Vol.15. No.6. P.229 319. Tarasov N.I. 1936. [To the fauna of Cirripedia Thoracica from the Arctic Ocean] // Trudy Arcticheskogo Nauchno-Issledovatel skogo Instituta. Vol.33. P.45 49 [in Russian]. Tarasov N.I. 1937. [To the fauna of Cirripedia Thoracica from the Arctic Ocean. III] // Trudy Arcticheskogo Nauchno-Issledovatel skogo Instituta. Vol.50. P.35 59 [in Russian]. Tarasov N.I. 1945. [To the fauna of Cirripedia Thoracica from the Arctic Ocean. IV] // Problemy Arktiki. Vol.4. P.51 61 [in Russian]. Tarasov N.I., Zevina G.B. 1957. [Cirripedia Thoracica from the seas of the USSR] // Fauna SSSR. Rakoobraznye. Vol.6. No.1. P.1 267 [in Russian]. Thomson C.W. 1873. Notes from the Challenger // Nature. Vol.8. P.347 349. Thomson C.W. 1877. The Voyage of the Challenger. The Atlantic. 2. London: MacMillan 396 p. Weisbord N.E. 1979. Lepadomorph and Verrucomorph barnacles (Cirripedia) of Florida and adjacent waters with an addendum on the Rhizocephala // Bull. Amer. Paleont. Vol.76. No.306. P.1 156. Weltner W. 1897. Verzeichnis der bisher beschriebenen recenten Cirripedien Arten // Archiv für Naturgeschichte. Bd.63. H.3. Nr.1. S.227 280. Weltner W. 1922. Cirripedia der Deutschen Tiefsee-Expedition // Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Exped. auf dem Dampfer Valdivia. Bd.23. Nr.2. S.59 112. Withers T.H. 1953. Catalogue of fossil Cirripedia. III. Tertiary. British Mus. (Nat. Hist.) London 396 p. Young P.S. 1992. Lepadomorph cirripeds from the Brazilian coast. II Family Scalpellidae // Bull. Mar. Sci. Vol.50. No.1. P.40 55. Young P.S. 1998a. Cirripedia (Crustacea) from the Campagne Biaçores in the Azores region including a generic revision of Verrucidae // Zoosystema. Vol.20. No.1. P.31 92. Young P.S. 1998b. The Cirripedia (Crustacea) collected by the Fish. Steamer Meteor in the Eastern Atlantic // Arquivos do Mus. Nac. Rio de Janeiro. Vol.58. P.1 54. Young P.S. 2000. Cirripedia Thoracica (Crustacea) collected during the Campagne de La Calypso (1961 1962) from the Atlantic shelf of South America // Zoosystema. Vol.22. No.1. P.85 100. Young P.S. 2001. Deep-sea Cirripedia Thoracica (Crustacea) from the northestern Atlantic collected by French expeditions // Zoosystema. Vol.23. No.4. P705 756. Young P.S. 2002. Revision of the Scalpellidae (Crustacea Cirripedia) in the collection of the Museum national d Histoire naturelle, France, studied by Abel Gruvel // Zoosystema. Vol.24. No.2. P.309 345. Young P.S. 2004. On the validity of little-known species of Scalpellidae (Crustacea Cirripedia) collected by the Campagnes scientifiques du S.A.S. Prince de Monaco from North Atlantic // Arquivos do Museu Nac. Rio de Janeiro. Vol.62. No.1. P.25-30. Young P.S., Leta A.C. 1996. Antarctic Cirripedia (Crustacea) collected by the Brazilian Antarctic program (1983 1986) // Iheringia, Porto Alegre, Zoologia. Vol.80. P.121 126. Zevina G.B. 1970. [Cirripedia of the north-western Pacific] // Trudy Instituta Oceanologii. Vol.86. P.252 276 [in Russian, with English summary]. Zevina G.B. 1971. [Verrucomorpha (Cirripedia) of the south-east Pacific] // Zool. Zhurn. Vol.50. No.3. P.439 441 [in Russian, with English summary]. Zevina G.B. 1972. Benthic Lepadomorpha (Cirripedia Thoracica) from the southeast Pacific // Crustaceana. Vol.22. No.1. P.39 63. Zevina G.B. 1973a. [Scalpellidae (Cirripedia) of the Indian Ocean 1. Species of the subgenera Scalpellum and Arcoscalpellum of the genus Scalpellum] // Zool. Zhurn. Vol.52. No.6. P.842 848 [in Russian, with English summary]. Zevina G.B. 1973b. [Scalpellidae (Cirripedia) of the Indian Ocean 2. Species of the subgenera Annandaleum, Mesoscalpellum and Neoscalpellum of the genus Scalpellum] // Zool. Zhurn. Vol.52. No.7. P.1000 1007 [in Russian, with English summary]. Zevina G.B.1973c. [Scalpellidae (Cirripedia) of the Gulf of Alaska] // Trudy Instituta Oceanologii. Vol.91. P.136 140 [in Russian, with English summary]. Zevina G.B. 1975.[Cirripedia collected on the 11 cruise of the RV Akademik Kurchatov in the Atlantic sector of the Antarctic] // Trudy Instituta Oceanologii. Vol.103. P.183 192 [in Russian, with English summary]. Zevina G.B. 1976. [Deep-sea species of cirripedians from the North Atlantic] // Zool. Zhurn. Vol.55. No.8. P.1149 1156 [in Russain, with English summary]. Zevina G.B. 1978a. [New system of the family Scalpellidae (Cirripedia Thoracica). I. Subfamilies Lithotryinae, Calanticinae, Pollicipinae, Scalpellinae, Brochiinae and Scalpellopsinae] // Zool. Zhurn. Vol.57. No.7. P.998 1007 [in Russain, with English summary]. Zevina G.B. 1978b. [New system of the family Scalpellidae (Cirripedia Thoracica). II. Subfamilies Arcoscalpellinae and Meroscalpellinae] // Zool. Zhurn. Vol.57. No.9. P.1343 1352 [in Russain, with English summary]. Zevina G.B. 1981a. [Cirripedians of the suborder Lepadomorpha of the world ocean. I.] Family Scalpelidae. Leningrad: Nauka. 406 p. [in Russian, with English summary] Zevina G.B. 1981b. [Deep-sea cirripedians from Australia and New Zealand] // Trudy Instituta Oceanologii. Vol.115. P.76 93 [in Russian, with English summary]. Zevina G.B. 1982. [Cirripedians of the suborder Lepadomorpha of the world ocean. II.] Leningrad: Nauka. 222 p. [in Russain, with English summary] Zevina G.B. 1983. [Cirripedians from seamounts from the Nazca Ridge] // Zool. Zhurn. Vol.62. No.11. P.1635 1642 [in Russain, with English summary].

Deep-sea fauna of European seas. Cirripedia 111 Zevina G.B. 1987a. [Abyssal Verrucomorpha (Cirripedia Thoracica) of the Atlantic and Indian Oceans] // Zool. Zhurn. Vol.66. No.9. P.1304 1313 [in Russain, with English summary]. Zevina G.B. 1987b. [Abyssal Verrucomorpha (Cirripedia Thoracica) of the Pacific. 1. Northern Pacific] // Zool. Zhurn. Vol.66. No.12. P.1812 1821 [in Russain, with English summary]. Zevina G.B. 1988. [Abyssal Verrucomorpha (Cirripedia Thoracica) of the Pacific. 2. Southern Pacific] // Zool. Zhurn. Vol.67. No.1. P.31 40 [in Russain, with English summary]. Zevina G.B. 1990. [Cirripedia Thoracica from the guyot summits of Nazca and Sala y Gomez] // Trudy Instituta Oceanologii. Vol.124. P.174 186 [in Russian, with English summary]. Zevina G.B. 1993. [Abyssal species of scalpellids (Cirripedia Thoracica) from the Atlantic sector of the Antarctic] // Trudy Instituta Oceanologii. Vol.127. P.122 128 [in Russian, with English summary]. Zevina G.B., Galkin S.V. 1992. [Altiverruca beringiana sp. n. (Crustacea Cirripedia) first finding of Verrucomorpha in the boreal Pacific] // Zool. Zhurn. Vol.71. No.8. P.140 144 [in Russain, with English summary]. Zevina G.B., Galkin S.V. 1993. [New species of Catherinum (Cirripedia Lepadomorpha)] // Zool. Zhurn. Vol.72. No.3. P.147 149 [in Russain, with English summary]. Zevina G.B., Kolbasov G.A. 2000. [Barnacles of the genus Heteralepas (Thecostraca Cirripedia Thoracica) from the Canary Islands and the Azores. Description of mantle ultrastructure] // Zool. Zhurn. Vol.79. No.11. P.1275 1283 [in Russain, with English summary]. Zevina G.B., Shreider Ì.U. 1992. [New species of Cirripedia from the Indian Ocean] // Zool. Zhurn. Vol.71. No.10. P.39 46 [in Russain, with English summary]. Zevina G.B., Tarasov N.I. 1964. [To the fauna of cirripedians (Cirripedia Thoracica) of the Arctic Ocean] // Trudy Arkticheskogo i Antarcticheskogo Nauchoissledovatel skogo Instituta Vol.259. P.229 240 [in Russain, with English summary]. Zevina G.B., Yakhontova I.V. 1987. [A new genus of Scalpellidae (Crustacea Cirripedia) from the North Atlantic] // Zool. Zhurn. Vol.66. No.8. P.1262 1264 [in Russain, with English summary]. Responsible editors A.V. Gebruk, H. Thiel, M. Thurston