Sheep Breeding in Norway

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Sheep Breeding in Norway Sheep Breeders Round Table 2015 Thor Blichfeldt Ron Lewis Director of Breeding Professor, University of Nebraska-Lincoln The Norwegian Association of Sheep and Goat Breeders (NSG)

Outline Sheep production in Norway Breeds Sheep recording NSG the sheep breeding company Traits, EBVs and total merit index Genetic gain R&D Genomics Key points about sheep breeding in Norway

Sheep production in Norway (1) Production per year Sheep farmer economy 24 million kg of meat 4 million kg of wool Heavily subsidised - 2/3 of the income Consumption 5 kg per capita per year Self sufficient Some import Tax protection Per animal Per hectare Good farmer prices 4-5 per kg carcass weight 3 per kg of wool Norwegian costs - high (oil related) Net income from sheep Low, but improving

Sheep production in Norway (2) Structure Housed during winter Farmers: 12 000 Ewes (1+ years): 700 000 Before mating until 1-2 weeks after lambing Small flocks: Average 55 ewes Few 300+ ewes Intensive care during lambing time Assist the ewe Production very seasonal Indoor lambing April-May Slaughter in August-November Assist the lamb Colostrum One lost lamb is one too many Age 160 days Carcass weight 20 kg

Sheep production in Norway (3) - Feeding Norway: 3% arable land Grass silage in the winter Spring pasture on farm 2-6 weeks Aprile May June Summer pasture in the woods or the mountains (mid June - mid Sept.)

Photo: Grethe Ringdal

Photo: Grethe Ringdal

Photo: Grethe Ringdal

Photo: Grethe Ringdal

Sheep production in Norway (3) - Feeding Grass silage in the winter Autumn pasture on farm Spring pasture on farm 2-6 weeks Aprile May June Summer pasture in the woods or the mountains (mid June - mid Sept.) First group of lambs: Directly to the abattoir September Second group: October Third group: November The rest: Jan. - Feb. Concentrate Ewes Lambs

Breeds Norwegian Spælsau 3 lines: 15 % Most important: Flock instinct Short tail ( spæl ) Fleece: Dual coated wool, white - coloured Polled - Horned Norwegian White Sheep (NWS): 75 % Long tail Fleece: Crossbred type, white Polled 10-15 other breeds: 10 %

Norwegian Spælsau of today

Norwegian White Sheep - definitely a composite NWS: A population, not a proper breed

NWS: - Sire line or maternal line? 90,000 ewes in 950 breeding flocks (2014) Number of lambs born: 2.29 Age of slaughter, days; 156 Carcass weight, kg; 21.1 Carcass conformation: R+ (9.3) NWS is a dual purpose breed Used as pure No appreaciable crossbreeding in Norway

Animalia: Marit presents a poster The Norwegian Sheep Recording System Sheep Recording in Norway Central database On farm data Sheep producers: 30% Web / Mobile app Ewes: 43% Abattoir data Slaughtered lambs: 49% File transfer Output Management tool Benchmarking EBVs R&D

Recording (2) Individually recorded Abattoir info Electronic ear tags (EID) Carcass weight Birth info Dam and Sire Total born EUROP conformation and fat score Fleece weight and quality Live born Lambing ease (code) Weights of lambs Birth 6 weeks 20 weeks (weaning) NOT DONE IN NORWAY Ultrasound scanning for meat and fat CT scanning Fecal egg count Disease Mastitis

The breeding company: - NSG and the Ram Circles (1) NSG: - The breeding plan - EBVs - AI 80% of sheep farmers are members of NSG (10,000) NSG budget for breeding: 1.3 mill. NSG Breeding Council Gov. support: 45% 5 breeders Levy on meat: 15% 2 from the abattoirs AI sales: 40% 1 from the Agr. University Central office at Ås Breeding and AI is half of the activities in NSG Responsible for the breeding programme Calculating EBVs 13 runs per year Director: Thor B. AI 3 geneticists R&D 3 breeding consultants

The breeding company: - NSG and the Ram Circles (2) The Ram circle: - Select rams - Circulate rams - Plan elite matings Ram circle: A small financially independent organization that has breeders as members More than 50 years of good work Rams are owned by the ram circle and used among member flocks Ewes are owned by the members themselves Cooperating with NSG Regulations Guidelines Financial support from NSG 150 per test ram that qualifies

NWS breeding AI: The key to success - Selection intensity of rams - Connectedness among flocks The breeding population Elite matings 150 ram circles 950 members 15% of ewes in ram circles are AI d 90,000 ewes Progeny testing of rams - Selection within ram circle Test rams (0.5 y.): 1,800 Elite rams (1.5 y.): 300 AI - Selection across ram circles 20 rams (2.5 years) 5 rams (3.5 years) Sired by an AI ram in ram circle flocks (2014) Lambs born: 9% Lambs slaughtered: 5% Ewes lambing: 21% AI sires in ram selection Test rams sired by an AI ram: 85%

AI the Norwegian way - 35.000 semen doses per year A shot in the dark No synchronisation, no hormone treatment Oestrus detection 2-3 times per day Walk the ram; Leach and apron Inseminate once 18-24 hours after onset of oestrus Frozen semen 240 mill. sperm cells Vaginal deposition Done by the farmer Non-return: 70% Cost: (Rent of shipper + freight + 20 doses) Per semen dose: 23

Traits in the breeding work - heritabilities and weighting Selection on total merit index NKS Lamb traits Heritability h 2 Weight in the total merit index Growth, carcass weight at 22 w. 0.12 24 % EUROP conformation score, at 20 kg 0.19 18 % EUROP fat score, at 20 kg 0.19 11 % Fleece weight, at 20 kg 0.33 2 % Fleece grade, at 20 kg 0.08 0 % Ewe traits Maternal ability, at 6 weeks 0.06 15 % Maternal ability, at 22 weeks 0.05 24 % Litter size, total born 0.13 6 %

The name of the game (1): Genetic gain litter size Enough is enough! Inger Anne presents a poster

The name of the game (2): Genetic gain lamb growth Goal: Slaughtered straight from the mountains Jette and Thor presents a poster

R&D The breeders: Have to do more recording! New traits? Improved models Early lamb loss Adjusting weights for age Lambing ease (new scoring) Lamb vigour Suckling assistance Udder and teat conformation Mastitis Longevity Adjusting EUROP scoring for age or weight Heterogeneous variances Litter size: Reduce variability Contemporary groups within flock-year Genomic information

Genomic selection The key equation BV X t = r BV X,P X (i X )σ BVX L Genomic tools may allow Increased accuracy ( r BVX,P X ) Via Genome-enhanced EBV Decreased generation interval ( L)

Opportunities Facilitate improvement of otherwise difficult-to-measure traits (r BVX,P X ; L) Traits expressed later in an animal s life Longevity Traits expressed in only one sex Fertility, litter size Traits that are expensive and/or challenging to measure Lamb survival, mastitis, maternal bonding, eating quality

Opportunities Facilitate improvement of otherwise difficult-to-measure traits (r BVX,P X ; L) Form a better pedigree (r BVX,P X )

Challenges Requirement is likely for large reference populations in individual breeds Large to capture genetic diversity within a breed Within breeds because genomic predictions do not extend well across breeds In many industries, there are lots of breeds Validation (training) must be ongoing Accuracies deteriorate as ancestors used to form genomic predictions become more distant Structure of ram circles likely well suited to form industry-based reference populations

Challenges Requirement is likely for large reference populations in individual breeds To keep up-to-date, need to measure about 12.5% new animals per year (Goddard, 2009; Hayes et al., 2009; van der Werf et al., 2011)

Challenges Requirement is likely for large reference populations in individual breeds Costs of genotyping Value per animal unit is relatively low in sheep However, the per animal cost of genotyping is nearly the same across species

Benefits vs. costs The utility of genomic selection in the Norwegian industry will depend on Gain in accuracy realized Function of the number (diversity) of performance recorded sheep, e.g., 1/4 million Norwegian White ewes Economic importance of traits being considered Cost and thereby industry uptake of genotyping

Norwegian lamb finishing system Lambs grazed over-summer on mountain or forest pasture Once gathered, drafted on-farm Over-weight & over-finished lambs marketed immediately Target weight & finished lambs also marketed immediately Under weight and/or under-finished lambs retained Grass and/or concentrate fed until achieve a target end-point or end-of-season

Carcass weight (kg) Carcass weight by age (2014) 25 23 21 19 17 15 80 100 120 140 160 180 200 220 Days of age

Norwegian lamb finishing system What then is the target end-point for adjustment for genetic evaluation? Weight (20-25 kg)? Age (20 weeks)? Finish? A combination of several? Given varying drafting strategies, how should contemporary groups be defined? Will (and should) producers provide more delineating contemporary group designations?

Sheep breeding Making results together! - what is unique for Norway? Centrally financed The sheep breeders 1.3 million Recording One central database Carcass data transfer Large breeding population 90,000 ewes A strong belief in the breeding theory Confidence in the central breeding management The same breeding goal for all breeders within a breed 1,800 rams AI The very best rams Intensively used Collaboration, not competition Within ram circle Across ram circles Substantial genetic and phenotypic gain

Invitation to World Championship in Ewe Productivity Norwegian contestant Hove 2010-00003 Year Lambs 20 week weight 2011 2 60 + 61 2012 4 62 + 67 + 64 + 61 2013 3 63 + 66 + 72 2014 3 61 + 59 + 60 2015 2 62 + 57 Sum 14 875 kg 175 kg per year Progeny for breeding: - 6 daughters - 2 AI rams Concentrate given to the ewe, not the lambs: 56 kg per year