Taxonomical notes on Anthaxia olympica species-group (Coleoptera: Buprestidae) Pavel Svoboda

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1 FOLIA HEYROVSKYANA Vol. 8(3-4): ISSN editum: December 31, 2000 Taxonomical notes on Anthaxia olympica species-group (Coleoptera: Buprestidae) Pavel Svoboda Svákov 710 / III, CZ Soběslav, Czech Republic Abstract. Anthaxia {Haplanthaxia) denesi sp. n. from Turkey, and Anthaxia (Haplanthaxia) kantneri sp. n. from Tunisia, are described, illustrated and compared with related species. New statuses arc proposed for Anthaxia {Haplanthaxia) oreas Peyerimhoff, 1919 and Anthaxia {Haplanthaxia) astoreth Obenberger, New synonym is proposed for Anthaxia {Haplanthaxia) praeclara praeclara Mannerheim, 1837 = Anthaxia {Haplanthaxia) praeclara var. chlorogastra Obenberger, 1938 syn. n. Key to complexes and species of Anthaxia {Haplanthaxia) olympica Kiesenwcttcr, 1880 species-group is given. Taxonomy, new species, new statuses, synonymy, key, Buprestidae, Anthaxia, Palaearctic region Introduction The Anthaxia olympica species-group was established by Obenberger (1917) and divided by Novak (1988) into two complexes; this species-group was partly keyed by Novak (1988: A. scutellaris complex,) and by Niehuis (1991: East-Mediterranean species). Since this time several new taxa were described: A. samai Curletti et Magnani, 1992, A. praeclara creticola Brandl et Muhle, 1998 and A. francescoi Sparacio et Svoboda, 1999; this fact, the discovery of further new species and necessity of change of some statuses induce me to key complete species-group inclusive of dividing it into three complexes. The following abbreviations are used in the text: MNHN collection of Museum national d'histoire naturelle, Paris, France; NMPC collection of National Museum, Prague, Czech Republic. Anthaxia {Haplanthaxia) denesi sp. n. (Figs 14,35,52,59) Type material. Holotype (<$): Turkey, Karadut env., 2.vii. 1993, K. Denes jr. leg. Type deposition. Holotype deposited in NMPC. Description. Holotype: small (4.7x1.8 mm), rather vaulted, lustrous species; frons golden green, vertex black, pronotum blue-green with golden green lateral margins and with two small oval, indistinctly delimited blue-black spots, elytra golden-red, elytral base blue-green, triangular postscutellar field reaching nearly

2 138 Taxonomical notes on Anthaxia olympica species-group 1 Coleoptera: Bnprestidae) fourth of elytral length, blue-green with golden margins, ventral side of body golden green, sternites black with intensive golden green lustre, legs and antennae black; irons and ventral side of body with short, elytra with very short white pubescence (Figs 35, 59). Head small, clypeus with shallow triangular incurvation, frons flat, vertex narrow, about 1.4 times as wide as width of eye, eyes rather large, not projecting beyond outline of head, with moderately S-shapcd inner margins; sculpture of head consisting of small rounded cells with well visible central grains; antennae long, 1.6 times longer than length of pronotum, the first antennal segment pear-shaped, the second one relatively long, twice longer than wide at apex, antenno-meres 3 and 4 triangular, of the same length, antennomeres 5-10 trapezoidal, 1.5 limes longer than wide, apical segment slender, narrowed towards apex. Pronotum moderately vaulted, about 1.6 times wider than long, with small, rounded, shallow lalerobasal depressions, anterior pronotal margin slightly lobate medially, posterior one bisinuous, lateral margins angulated near anterior fourth, straightly tapering anteriorly and almost parallel at posterior three fourth, maximum pronotal width before anterior third; sculpture of pronotum homogeneous, consisting of regular network of deep polygonal cells with sharp, distinct central grains, basal microsculplure undeveloped. Scutellum blue, pentagonal, as long as wide, roughly microsculptured. Elytra slightly wedge-shaped, 1.9 times longer than wide at humeri, humeral swellings well developed, basal transverse depressions shallow; lateral margins of elytra sinuately narrowed, almost straight and finely serrate at apical third, elytral epipleura not reaching elytral apex, each elytron widely rounded separately; elytral surface lustrous, with regular, rather rough grainy sculpture without basal microsculpture. Ventral side of body with fine ocellate sculpture, sternites almost smooth; anal sternile widely truncate and s-lightly incurved apically, with shallow but distinct trapezoidal preapical depression, lateral margins finely serrate before apex. Legs relatively short, slender, protibiae moderately bent inwards, pro- and mesotibiae with fine inner serration at apical third, inner margin of metatibiac incurved and roughly serrate before apex (Fig. 52); trochanters simple. Acdeagus short, subparallel, parameres somewhat enlarged at their midlcnglh (Fig. 14)/' Female: unknown. Host plant: unknown. N a m e d e r i v a t i o n. This species is named after my friend Karel Denes sen. Differentia] d i a g n o s i s. Anthaxia denesi sp. n. differs from other species of Anthaxia olympica species-group by regular pronotal sculpture consisting of deep polygonal cells with well developed central grains, by form of male metatibiae, form of aedeagus and by the other characters given in the key.

3 Folia Heyrovskyana, vol. 8(3-4): , I ';1 Figs Aedeagi of Anthaxia (Haplanthaxia) olympica species-group (dorsal view). 1 -A. (//.)praeclara praedara Manncrhcim; 2 - A. (H.) p. creticola Brandl et Miihle; 3-/4. (H.) samai Curletti et Magnani;.4 - A. (H.) scutellaris scutellaris Gene; 5 -A. (//.) 5. atlasica Thcry; 6 - A. (H.) s. semireducta Pic; 7 - A. (H.) israelita Abeille de Perrin; 8 - A. (H.) escalerina Obenberger; 9 - A. (H.) escalerinella Novak; 10-/4. (//.) franceskoi Sparacio et Svoboda; 11 -A. (H.)olympica Kiesenwetter; 12-A. (//.)astorelh Obenberger stat. n.

4 140 Taxonomical notes on Anthaxia olympica species-group '.Coleoptera: Baprestidae) Anthaxia (Haplanthxia) kantneri sp. n. (Figs 18,36,56,60) T y p c m a t e r i a I. Holotype ($): Tunis, Ain El I Iamaraya, 7.vi. 1994, F. Kantner leg. T y p e deposition. Holotype deposited in NMPC. Description. Holotype: medium sized (5.75x2.05 mm), rather flattened, slightly lustrous species: head, pronotum, ventral side of body, elytral base and narrow, indistinctly differentiated triangular postscutelar field golden green, vertex and two small indistinct pronotal spots black, elytra golden-brown, legs and antennae black with golden green lustre; frons, anterior pronotal angles and ventral side of body with short, elytra with very short white pubescence (Figs 36, 60). Head small, clypeus distinctly triangularly incurved anteriorly, frons slightly convex with shallow longitudinal depression; vertex narrow, as wide as width of eye, inner margin of eyes distinctly S-shaped, eyes relatively large but not projecting beyond outline of head; sculpture of head consisting of dense network of small polygonal cells without central grains; antennae about 1.4 times longer than length of pronotum, the first antennal segment pear-shaped, the third segment only slightly longer than the second or the fourth ones, antennomeres 5-10 trapezoidal, about as long as wide, the last segment slender, narrowed to apex, twice longer than wide at apex. Pronotum slightly convex, subcordiform, 1.6 times as long as wide, with transverse, narrow, and shallow laterobasal depressions; anterior pronotal margin only slightly lobate medially, posterior one bisinuous, lateral margins almost regularly rounded and incurved at apical third, maximum pronotal width at anterior third; pronotal sculpture consisting of network of small oval and polygonal cells with minute but well distinct central grains laterally, central part of pronotum with transversely prolonged cells without central grains, sides of cells form distinct irregularly bent transverse wrinkles; basal microsculpture absent. Scutellum black with green lustre, triangular, about 1.1 times as wide as long. Elytra wedge-shaped, flattened dorsally, about twice longer than wide at humeri, humeral swellings rather small, transverse basal depressions well developed, elytral epipleura not reaching apex of elytra. Elytral lateral margins widely S-shaped with fine serration at apical third, each elytron rounded separately; elytral surface with fine regular grainy sculpture. Ventral side of body with indistinct ocellate sculpture; anal sternite roundly truncate apically with shallow trapezoidal preapical depression, lateral margins distinctly serrate before apex. Legs relatively short, slender, tibiae straight, pro- and mesotibia with blunt inner serration, metalibia with distinct tooth at apical fourth and wide inner preapical incurvation (Fig. 56); trochanters simple. Aedeagus rather long, parameres enlarged before their midlength with distinct outer serration before roundly enlarged apical part (Fig. 18). Female: unknown. Host plant: unknown.

5 Folia Heyrovskyana, vol. 8(3-4): , Figs Anthaxia {Haplanthaxia) olympica species-group : Aedeagi in dorsal view; 13 -A. (H.) niehnisi Brandl; 14 - A. (//.) denesi sp. n.; 15 - A. (//.) klessi Niehuis; 16 - A. (//.) ignipennis Abeille de Perrin; 17 - A. {H.)fulgidipennis Lucas; 18 - A. (//.) kantneri sp. n.; 19 - A. (H.) paralella Gory et Laporte; 20 - A. (//.) oreas Peyerimhoff stat. n : anal sternite (not in scale); 21 - A. (H.) praedara Mannerheim; 22 -A. (H.) samai Curletti et Magnani male mcsotrochanter of A. (//.) praedara; 24 - male metatrochanter of A. (//.) praedara; 25 - male mesotrochanter of A. (//.) samai; 26 - male metatrochanter of A. (//.) samai.

6 142 Taxonomical notes on Anthaxia ulympica specics-group i Coleoptera: Buprestidae) Name d e r i v at i o n. This species is named after the collector Dipl. Ing. František Kantner. Differential diagnosis. Anthaxia kantneri sp. n. resembles by its pronolal sculpture Anthaxia parallela Gory el Laporte, 1839 and Anthaxia oreas Peyerimhoff, 1919 stat. n., and it differs from both these species by its coloration, length of antennae, form of male metatibiae and aedeagus, and by other characters given in the key. Anthaxia (Haplanthaxia) oreas Peyerimhoff, 1919 stat. n. Anthaxia (Haplanthaxia) parallela oreas Peyerimhoff', 1919: 195. After having studied an extensive material of A. parallela oreas including one paratype (MNHN, the other type specimens were not found) and A. parallela parallela Gory et Laporte, 1839, I consider A. oreas Peyerimhoff, 1919 to be a good species. It differs from A. parallela by quite different male genitalia (Figs 19, 20) and male metatibiae (Figs 57, 58). Anthaxia (Haplanthaxia) astoreth Obenberger, 1934 stat. n. Anthaxia (Haplanthaxia) olympica astoreth Obenberger, 1934: 175. After having studied a rich material of both subspecies including three Obenberger's syntypes (1 male, 2 females in NMPC) I consider A. astoreth Obenberger, 1934 to be a good species. It differs from A. olympica Kiesenwetter, 1880 by its more lustrous dorsal side of body, elytral sculpture with rows of punctures, form of aedeagus (Figs 11, 12) and especially form of male metatrochanlers (Figs 31, 32), which character I consider as very important for distinguishing the Anthaxiaspecies. 1. e c t o t y p e d e s i g n a t i o n. I utilize the occasion of studying all the 3 syntypes to select a lectotypc, a holotype not been indicated. I hereby designate as lectotype the only male and the remaining two females as paralectotypcs. All type material is deposited in NMPC. Anthaxia (Haplanthaxia) praeclara praeclara Mannerheim, 1837 Anthaxia praeclara Mannerheim, 1837: 87. Anthaxia praeclara var. chlorogastra Obenberger, 1938: 186, syn. n. After having studied a male syntype (number of type species not mentioned in the original description) of A praeclara var. chlorogastra (NMPC) I have found no differences between the variety and nominotypical subspecies A. praeclara praeclara and I consider A. praeclara var. chlorogastra Obenberger to be a junior synonym of A. praeclara praeclara Mannerheim, 1837.

7 Folia Heyrovskyana, vol. 8(3-4): , Figs : male mesotrochanters (not in scale); 27 - Anthaxia (Haplanthaxia) scutellaris scutellaris Gene; 28 - A. (//.) olympica Kicscnwetter; 29-/4. (//.) astoreth Obcnbcrger stat. n : male mctatrochanters (not in scale); 30 - A. (H.) scutellaris scutellaris; 31 - A. (H.) olympica; 32- A. (//.) astoreth : male mcsotibiae; 33 - A. (H.) praeclara praeclara Mannerhcim; 34 - A. (H.) p. creticola Brandl et Miihle : outline of male body; 35 - A. (H.) denesi sp. n.; 36 - A. (//.) kantnteri sp. n.; 37 - A. (H.) paralella Gory ct Laportc; 38 - A. (H.) oreas Pcycrimhoff stat. n.

8 144 Taxonomical notes on Anthaxiu olympica species-group (Coleoptera: Buprestidae) Anthaxia (Haplanthaxia) olympica Kiesenwetter, 1880 species-group C h a r a c t e r i s t i c s of the species-group. Small to medium sized ( mm), slender, moderately wedge-shaped, mostly multicolorous species: male frons, pronotum, elytral basis and postscutellar triangular field golden green to blue, pronotum with two dark spots, elytra coloured by various shades of red or purple, exceptionally species with different coloration (see key); ventral side of body golden green or blue-green, less frequently black with intensive golden green or blue lustre, or bronze. Frons and ventral side of body with short, elytra with very short white pubescence. Sexual dichromatism only slightly developed: females differ usually by black frons and darker ventral side. Frons slightly convex or almost flat, moderately depressed medially, vertex narrow, about times as wide as width of eye; male antennae long, about times longer than length of pronotum, exceptionally shorter; antennomeres 5-10 trapezoidal. Pronotum slightly, less frequently distinctly vaulted, about times as wide as long, with shallow or almost indistinct laterobasal depressions and without distinct medial groove; lateral pronotal margins regularly rounded or moderately angulated, the widest part of pronotum at anterior half. Elytra slender, more (males) or less (females) sinuately narrowed, rarely subparallel, about twice as long as wide at humeri, exceptionally distinctly shorter; elytral epiplcura narrow, not reaching elytral apex, lateral margins of elytra finely serrate in apical third. Sculpture of head and pronotum consisting of a network of oval and polygonal cells with or without central grains, more rarely central part of pronotum with traces of transverse wrinkles; elytral sculpture usually grainy, rarely with rows of punctures; dorsal side of body with or without basal microsculpture; sculpture of ventral side ocellatc, less distinct on abdominal sternites than that on sternum. Legs short and slender, tibiae almost straight, exceptionally male pro- and metatibiae strongly bent inwards; male mesotibiae and less often protibiae with inner serration, metatibiae with deep or shallow inner prcapical incurvation and usually distinctly serrate in apical third. Pro- and metatrochanter with or without spine, mesotrochanter simple, less frequently transversely enlarged and prolonged to sharp lateral spine. Anal sternite widely truncate and slightly incurved apically, exceptionally regularly rounded with small apical incurvation, with fine lateral serration and with more or less developed preapical depression. Sexual dimorphism. Female differs from male by shorter antennae and by simple trochanters and tibiae.

9 Folia Heyrovskyana, vol. 8(3-4): , Figs Male metatibiac of Anthaxia (Haplanthaxia) olympica species-group A. (H.) praeclara praeclara Mannerhcim; 40-/1. (//.) p. creticola Brandl et Miihle; 41-/1. (H.) samai Curletti et Magnani; 42 - A. (H.) scutellaris scutellaris Gene; 43 - A. (H.) s. atlasica Théry; 44-/1. (Ji.) s. semireducta Pic; 45-/1. (H.) israelita Abeille de Perrin; 46- A. (//.) escalerina Obenbcrgcr; 47-/1. (//.) escalerinella Novak; 48 -A. {H.)franceskoi Sparacio et Svoboda; 49 -A. (//.) olympica Kiesenwetter; 50 - A. (H.) astoreth Obenberger stat. n.; 51 - A. (H.) niehuisi Brandl; 52 - A. (//.) dene si sp. n.; 53 - A. (H.) klessi Niehuis; 54-A. (//.) ignipennis Abeille de Perrin; 55 -A. (H.) fulgidipennis Lucas; 56-A. (//.) kantneri sp. n.; 57 - A. (H.) paralella Gory et Laporte; 58-/1. (//.) oreas Peyerimhoff stat. n.

10 146 Taxonomical notes on Anthaxia olympica species-group (Coleoptera: Buprestidae) Key to complexes and species of Anthaxia (Haplanthaxia) olympica species-group 1 (4) Rather short, robust species, pronotum distinctly, elytra moderately vaulted, male mesotrochanter transversely enlarged and prolonged to sharp lateral spine (Figs 23, 25): elytra distinctly wedge-shaped, about times longer than wide at humeri, protochanler with distinct spine on anterior margin {A. praeclara complex). 2 (3) Male antennae as long as pronotum, male metatrochanter with distinct spine on posterior margin (Fig. 24), anal stcrnite regularly rounded and slightly incurved apically with deep triangular prcapical depression (Fig. 21), occllate pronotal sculpture with central grains, elytra times longer than wide at humeri, clytral sculpture with distinct rows oť punctures; pronotum golden green or blue-green with two black spots, elytra purple, elytra! base and postscutellar triangular field golden green to bluc-grccn, lateral margins of elytra very frequently golden green; mm A. praeclara Mannerheim, 1837 a(b) Sculpture of pronotum consisting of finely bordered, somewhat longitudinally elongate cells with minute central grains, elytra shorter, about 1.6 times longer than wide at humeri in both sexes, male mesotibiae enlarged apically and more distinctly incurved before apex (Fig. 33); male mctatibia as in Fig. 39; aedcagus as in Fig. I; mm; Italy, the Balcans, Turkey, Cyprus, Syria, Jordan, Lebanon, Israel A. p. praeclara Mannerheim, 1837 b(a) Sculpture of pronotum consisting of distinctly bordered polygonal cells with small but distinct central grains, elytra rather slender, about 1.6 (females) to 1.8 (males) times longer than wide at humeri, male mesotibiae subparallcl with very shallow preapical incurvation (Fig. 34); male mctatibia as in Fig. 40; aedcagus as in Fig. 2; mm; Crete A.p. creticola Brandl et Miihle (2) Male antennae longer, about 1.6 times longer than length of pronotum, male metatrochanters without distinct apical spine (Fig. 26), anal stcrnite widely truncate and moderately incurved apically, without distinct depression (Fig. 22); occllate pronotal sculpture without central grains, elytra times longer than wide at humeri, with regular grainy sculpture; pronotum blue-green or blue without distinct black spots, elytra purple, clytral base and postscutellar field golden green; male mctatibia as in Fig. 41, aedeagus as in Fig. 3; mm; Morocco, Tunisia A. samcti Curletti et Magnani, (1) More slender species, body rather flattened, male mesotrochantcrs not modified (Figs 27, 28, 29), elytra less wedge-shaped, about twice longer than wide at humeri, rarely shorter, protrochanter exceptionally with spine on anterior margin. 5 (14) Postscutellar triangular field large, sharply differentiated from basal clytral coloration; proand metatrochanter with or without spine (A. scutellaris complex). 6 (7) Male protrochanter with small, metatrochanter with long hook-shaped spine (Fig. 30), male mesotibiae moderatclly. pro- and metatibiae strongly bent inwards mm A. scutellaris Gene a(d) Black pronotal spots smaller, distinctly narrowed posteriorly. b(c) Pronotal sculpture consisting of polygonal cells with weakly developed central grains along lateral margins, black part of pronotum roughly microsculptured with indistinct, shallow cells without central grains; male frons golden green, pronotum golden green to blue with two blueblack spots, scutellar triangle golden green or bluc-grccn, elytra puiple; male mctatibia as in Fig. 42; aedeagus as in Fig. 4; mm: Portugal, Spain, Fiance, Italy. Croatia. Greece, Morocco, Algeria, Tunisia A. s. scutellaris Gene, 1839 c(b) Pronotal sculpture consisting of polygonal cells with distinct central grains laterally, black part of pronotum lustrous without microsculpture, cells more distinct, with traces of central grains; male frons black with golden green lustre, pronotum and scutellar field golden green, elytra purple; male mctatibia as in Fig. 43; aedcagus as in Fig mm; Morocco A. s. atlasica Théry. 1928

11 Folia Heyrovskyana, vol. 8(3-4): , d (a) Pronotal black spots larger, only slightly narrowed posteriorly; pronotal sculpture consisting of well developed polygonal cells with sharp (laterally) or flat central grains; darker species: male frons golden green anteriorly and black posteriorly, almost whole pronotum black, only lateral margins narrowly golden green and narrow medial stripe blue, scutellar triangle smaller, golden green, elytra purple; male metatibia as in Fig. 44; aedcagus as in Fig. 6; mm; Corse A. s. semireducta Pic, (6) Male trochanters not modified, tibiae almost straight. 8 (11) Pronotum about 1.6 times wider than long, flattened, laterobasal depressions distinct, frons distinctly depressed medially. 9(10) Smaller ( mm) species, ocel late pronotal sculpture with flat, weakly developed central grains, scutellar triangle narrower than elytral base, lateral margins of elytra without posthumeral green spot; male frons, pronotum and scutellar triangle golden green, pronotal spots usually black, elytra purple-red; male metatibia as in Fig. 45; aedcagus as in Fig. 7; Syria, Lebanon, Jordan, Israel A. israelita Abeille de Perrin, (9) Large ( mm) species, ocellate pronotal sculpture with rather sharp central grains, scutellar triangle as wide as elytral base, lateral margins of elytra with narrow blue-green posthumeral spot; male frons green, pronotum green to blue-green with two black-violet spots, scutellar triangle golden green or green with blue-green lateral margins, elytra purpleviolet; male metatibia as in Fig. 46; aedeagus as in Fig. 8; Morocco A. escalerina Obcnbcrgcr, (8) Pronotum narrower, about times as wide as long, rather vaulted, laterobasal depressions rather indistinct or missing, frons only indistictly depressed medially. 12 (13) Pronotum about 1.5 times as wide as long, laterobasal depressions shallow, almost indistinct, frons slightly depressed medially; sculpture of lateral pronotal parts consisting of longitudinally elongate cells with small central grains, basal microsculpture almost absent, pronotum relatively lustrous; male frons black with golden green lustre, pronotum green with two large black spots, scutellar triangle golden green, shorter, not reaching anterior third of elytral length, elytra purple-violet; male metatibia as in Fig. 47; aedcagus as in Fig. 9; mm; Morocco A. escalerinella Novak, (12) Pronotum about 1.4 times wider than long, laterobasal depressions almost missing, frons indistinctly depressed medially; lateral part of pronotum with oval and polygonal cells with large, flat central grains, basal microsculpture well developed, pronotum less lustrous; male frons golden green, pronotum green or blue-green with somewhat smaller blue-black spots, scutellar triangle golden-green, long, reaching anterior third of elytral length, elytra purple with golden tinge laterally; male metatibia as in Fig. 48; aedeagus as in Fig. 10; mm; Tunisia A.francescoi Sparacio et Svoboda, (5) Postscutellar triangular field smaller, narrower, indistinctly differentiated from basal elytral coloration; trochanters not modified: mctatrochantcr only exceptionally with apical spine (A. olympica complex). 15 (20) Ocellate pronotal sculpture without distinct central grains. 16 (19) Medium-sized ( mm) species; pronotal sculpture consisting of small polygonal cells without central grains, only lateral part of pronotum with minute, irregularly situacd central grains. 17 (18) Male trochanters simple (Figs 28, 31), elytral sculpture regularly grainy, less lustrous with fine basal microsculpture; male frons golden green, pronotum golden green to blue with two large indistinctly delimited black spots, scutellar triangle golden green to blue-green, elytra usually red-brown with golden tinge; male metatibia as in Fig. 49; aedeagus as in Fig. 11; mm; Central and SE Europe, W Turkey : A. olympica Kiescnwettcr, 1880

12 148 Taxonomical notes on Anthaxia olympica species-group (Coleoptera: Buprestidae) 18 (17) Male metatrochanter with distinct apical spine (Fig. 32), elytral grainy sculpture with rougher microsculpture, more lustrous, with traces of rows of punctures; male frons golden green, pronotum golden green or rarely blue-green with two smaller black or blue-black spots, scutellar triangle mostly blue-green, elytra purple, less frequently golden-red; male metatibia as in Fig. 50; aedeagus as in Fig. 12; mm; SE Turkey, Syria, Jordan, Israel A. astoreth Obenberger, 1934 stat. n. 19 (16) Small species; pronotal sculpture consisting of relatively large shallow polygonal cells without central grains, only several cells in latcrobasal depressions contain central grains: male frons dark blue-green or blue, pronotum black with blue-green or blue lateral margins, elytra dark red-brown or almost brown with indistinct blue-green scutellar field; male metatibia as in Fig. 51; aedeagus as in Fig. 13; mm; S Turkey" A. niehuisi Brandt, (15) Ocellate pronotal sculpture with distinct central grains. 21 (24) Central grains present over the entire pronotum. 22 (23) Pronotal sculpture consisting of regular network of deep oval and polygonal cells with sharp central grains; male frons blue-green, pronotum blue-green with golden green lateral margins, with two smaller but distinct blue-black spots, elytra goden-red, scutellar field large, blue-green with golden margins; dorsal side of body without basal microsculpture, lustrous; male metatibia as in Fig. 52; aedeagus as in Fig. 14; 4.7 mm; E Turkey A. denesi sp. n. 23 (22) Sculpture of pronotum consisting of shallow oval and polygonal cells with large, flat central grains; male frons and pronotum blue, pronotal black spots less distinct, elytra darker, red-brown with small, indistinct blue-green triangular scutellar field; dorsal side finely microsculptured, matt; male metatibia as in Fig. 53; aedeagus as in Fig. 15; mm; Israel A. klessi Niehuis, (21) Central grains present only along lateral margins of pronotum. 25 (26) Central part of pronotum with distinctly separated, very small rounded cells anteriorly and with dense network of small polygonal, slightly transversely elongate cells on prescutellar part; large, brightly coloured and relatively lustrous species: pronotum blue-green with two distinctly delimited black or black-violet spots, elytra purple, postscutellar triangular field blue-green with golden margins; male metatibia as in Fig. 54; aedeagus as in Fig. 16; mm; Portugal, Spain, S France A. ignipennis Abeille dc Pcrrin, (25) Central part of pronotum with transversely confluent cells, sides of cells form more or less distinct transverse, irregularly bent wrinkles. 27 (30) Scutellar triangle golden green or blue-green, ventral side green to blue-green, male antennae shorter, about times as long as pronotum. 28 (29) Only prescutellar part of pronotum with short transverse wrinkles, multicoloured species: male frons, pronotum and scutellar triangle golden green to blue-green, large pronotal spots black, elytra purple; male metatibia as in Fig. 55; aedeagus as in Fig. 17; mm; Morocco, Algeria, Tunisia A.fulgidipennis Lucas, (28) Central part of pronotum with fine, well visible transverse wrinkles, less conspicuously coloured species: male frons, pronotum and scutellar triangle golden green, pronotum with two indistinct black spots, elytra golden-brown; male metatibia as in Fig. 56; aedeagus as in Fig. 18; 5.75 mm; Tunisia A. kantneri sp. n. 30 (27) Scutellar triangle blue-black with golden green margins, ventral side of body bronze, male antennae long, about times as long as pronotum. 31 (32) Male elytra subparallel, eyes not projecting beyond outline of head (Fig. 37); frons black with violet lustre (male) or black (female), pronotum black, lateral pronotal margins with violet lustre, elytra purple-violet; male metatibia as in Fig. 57; aedeagus as in Fig. 19; mm; Morocco, Portugal, Spain, S France A.parallela Gory ct Laporte, (31) Male elytra moderately wedge-shaped, eyes slightly projecting beyond outline of head (Fig. 38); frons black with golden green (male) or violet (female) lustre, pronotum black with golden or golden green lateral margins, elytra purple-violet; male metatibia as in Fig. 58; aedeagus as in Fig. 20; mm; Morocco, Algeria, Tunisia A. oreas Peyerimhoff, 1919 stat. n.

13 Folia Heyrovskyana, vol. 8(3-4): , Figs Anlhaxia {Haplanthaxia) denesi sp. n., holotype. 60-A. (H.) kantneri sp. n., holotypc. References BRANDLP. & MOHLEH. 1998: Anthaxia praeclara crcticola n. ssp. von Kreta. (Coleoptera, Buprcstidae, Anthaxiini). Ada Coleopterologica 14: MANNERHEIM C. G. 1837: Enumeration dcs Buprestides, et desription de quelcqucs novucllcs cspeces de cette tribu de la fainilie des Stcrnoxcs, dc la collection de M. le Comte Mannerheim. Bulletin de la Societě Imperiále des Naturalistes de Moscou 8: 87. NlEHUiS M. 1991: Anthaxia (Haplanthaxia) klessi sp. n. ein neuer Prachtkafcr aus Israel (Coleoptera: Buprestidae). Nachrichlenblatt der Bayerischen Entomologen 40: NOVAK G. 1988: Anthaxia (Haplanthaxia) escalerinella sp. n., einc bischcr uncrkannte Art der Anthaxia scutellaris-gruppc aus Siidmarokko (Coleoptera, Buprestidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 40: OBENBERGER J. 1917: Holoarktische Anthaxicn. Beitrag zur Monographic dcr Gattung. Archiv fiir Naturgeschichte, Ser. A 82(8): OBENBERGER J. 1934: De novis generis Anthaxia (Col. Bupr.) formis. Acta Societatis Entomologicae Cechosloveniae 31: OBENBERGER J. 1938: Etudes sur les especes du genre Anthaxia Eschsch. (Col. Bupr.). Acta Entomologica Musei Nationalis Pragae IB, 2: