Catalog of the Family Cordylidae

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1 Catalog of the Family Cordylidae Harold De Lisle 2016

2 Cover photo : Platysaurus orientalis, by Nilsroe CC BY-SA 4.0 Back cover : Ouroborus cataphractus in defence posture, by Hardaker

3 CONTENTS INTRODUCTION... 1 CATALOG... 6 ACKNOWLEGEMENTS REFERENCES TAXON INDEX... 32

4 The Cordylidae is a family of small to medium-sized lizards that occur in southern and eastern Africa. They are commonly known as girdled lizards, spinytail lizards or girdle-tail lizards. Cordylid lizards are diurnal and mainly insectivorous. They are terrestrial, mostly inhabiting crevices in rocky terrain, although at least one species digs burrows and another lives under exfoliating bark on trees, and one genus lives mainly on grassy savannahs. They have short tongues covered with long papillae, flattened heads and bodies. The body scales possess osteoderms and have large, rectangular, scales, arranged in regular rows around the body and tail. Many species have rings of spines on the tail, that aid in wedging the animal into sheltering crevices, and also in dissuading predators. The body is often flattened, and in most species is box-like on cross-section, with lateral fold. The morphology and organization of osteoderms is highly variable in the Cordylidae. High-resolution X- ray Computed Tomography (CT) is ideal for recovering information about individual osteoderm structure, as well as overall arrangement of these features. The images above show just a small part of the variation in armor across the Cordylidae. Courtesy of the AMNH

5 Most species have four limbs, but those in the genus Chamaesaura are almost entirely limbless, with only tiny spikes in place of the hind limbs. The family includes both oviparous and ovoviviparous species. Currently (2016) ten genera are recognized: Chamaesaura, with five species Cordylus, with twenty-one species Hemicordylus, with two species Karusasaurus, with two species Namazonurus, with five species Ninurta, with one species (montotypic) Ouroborus, with one species (montotypic) Platysaurus, with sixteen species Pseudocordylus, with six species Smaug, with eight species The phylogeny of the Cordylidae is shown below (from Pyron et al., 2013) with position of each genus. This Catalog is an attempt to summarize the genus and species taxonomic names of extant and recently extinct cordylid lizards. Several species concepts have been applied to these lizards, and changes in the species level taxonomy have been proposed and many more are to be expected for that reason alone. The systematic arrangement of the catalog is intuitive once a few principles are stated. Subspecific entries for each species are indented. Dates in brackets indicate that these have been subsequently determined from sources external to the original document.

6 Misspellings are generally excluded except where they have been widely used or misinterpreted. Junior subjective synonyms are listed under each valid name. In order to enhance the usefulness of the present list, the type species of each genus has been added keeping in mind the famous quote of Harold Cogger, "Taxonomy is a matter of opinion" (Golay et al., 1993) based, of course, on the evaluation of the best evidence. The taxonomy used herein is my evaluation and we accept full responsibility for its use. We have deliberately omitted most lapsus calami, nomina nuda, nomina dubia, nomina oblita, nomina substituta, etc., from the synonymy because these names are not available for use. The synonymy herein is limited to new descriptions. Entries for species consist of species or subspecies name, genus, author, year of description, publication page, type locality, location of primary (name-bearing) type specimen(s), if known, and known distribution. Type localities are those in the original description, except those in obvious error or those amended subsequently by authoritative sources. Acronyms of type collections employed in the catalogue follow those used by American Museum of Natural History. Abbreviations of museums holding the types are as follows ANSP - Academy of Natural Sciences, Department of Herpetology, 19th and the Benjamin Franklin Parkway, Philadelphia, Pennsylvania 19103, USA. BMNH The Natural History Museum, Department of Zoology, Cromwell Road, London SW7 5BD, United Kingdom. CAS - California Academy of Sciences, Department of Herpetology, Golden Gate Park, San Francisco, California 94118, USA MBL - Museu Bocage, Universidade de Lisboa, Lisbon, Portugal [destroyed by fire in March 1978] MCZ - Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts 02138, USA. MD - Museu Regional do Dundo, Dundo, Luanda-Norte, Angola MNHNP - Museum National dhistoire Naturelle, Laboratoire des Amphibiens et Reptiles, 25 Rue Cuvier, Paris, France MSNG - Museo Civico di Storia Naturale di Genova Giacomo Doria, Via Brigata Liguria 9, Genova, Italy. NHMW - Naturhistorisches Museum, Zoologische Abtheilung, Postfach 417, Burgring 7, 1014 Wien, Austria NHRM - Naturhistoriska Rijkmuseet, Section for Vertebrate Zoology, Roslagsvägen 120, Box 50007, Stockholm 50, Sweden. NMP KwaZulu-Natal Museum, Pietermaritzburg 3201, KwaZulu-Natal, South Africa NMZB - National History Museum of Zimbabwe, Bulawayo, Zimbabwe.

7 PEM - Port Elizabeth Museum, Port Elizabeth, Republic of South Africa. QM - Queensland Museum, Gregory Terrace, Fortitude Valley, Brisbane, Queensland 4006, Australia. RMNH - Naturalis (Nationaal Natuurhistorisch Museum; formerly Rijksmuseum van Natuurlijke Historie), Postbus 9517, 2300RA Leiden, The Netherlands SAM - South African Museum, P.O. Box 61, Cape Town 8000, Western Cape Province, South Africa. TMP - Transvaal Museum, P.O. Box 413, Pretoria 0001, Gauteng, South Africa. UTEP - University of Texas at El Paso, Department of Biological Sciences and Centennial Museum, Laboratory for Environmental Biology, 500 W University Ave, El Paso, TX 79968, USA ZMA - Universiteit van Amsterdam, Zoologisch Museum, Plantage Middenlaan 53, Amsterdam: C, Netherlands (now incorporated into NCB Naturalis, Leiden). ZMB - Universität Humboldt, Zoologisches Museum, Invalidenstrasse 43, Berlin, Germany. ZMH - Zoologisches Institut und Museum, Universität Hamburg, Martin-Luther-King Platz 3, D-2000 Hamburg 13, Germany.

8 Cordylidae Mertens, 1937 Mertens, R Reptilien und Amphibien aus dem südlichen Inner-Afrika. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt, vol. 435, p Type genus : Cordylus Laurenti, 1768 Chamaesaura Schneider, 1801 Schneider, J.G Historiae Amphibiorum naturalis et literariae. Fasciculus secundus continens Crocodilos, Scincos, Chamaesauras, Boas. Pseudoboas, Elapes, Angues. Amphisbaenas et Caecilias., Frommani, Jena. 364 pp. Type species: Lacerta anguina Linnaeus, 1758: 210 Chamaesaura aenea (Fitzinger, 1843) Cricochalcis aenea Fitzinger 1843: 21 Type locality: South Africa Holotype: ZMB 1172 Distribution: South Africa: Northwest, Free State, Eastern Cape, KwaZulu-Natal provinces; Swaziland Chamaesaura anguina Linnaeus, 1758 Chamaesaura anguina anguina (Linnaeus, 1758) Lacerta anguina Linnaeus 1758: 210 Type locality: Cape of Good Hope, South Africa Holotype: BMNH Distribution: South Africa: Eastern Cape, Western Cape, Gauteng, Mpumalanga, KwaZulu-Natal provinces Chamaesaura anguina oligopholis Laurent, 1964 Chamaesaura anguina oligopholis Laurent, 1964: Type locality: Calonda, Lunda, Angola Holotype: MD? Distribution: Angola

9 Chamaesaura macrolepis (Cope, 1862) Mancus macrolepis Cope, 1862: 339 Type locality: Umvoti, KwaZulu-Natal, South Africa Syntypes: ANSP Distribution: South Africa: Northwest, Mpumalanga, KwaZulu-Natal provinces; Swaziland; southestern Zimbabwe Chamaesaura miopropus Boulenger, 1895 Chamaesaura miopropus Boulenger, 1895: 724 Type locality: Fwambo, northeastern Zambia Holotype: BMNH Distribution: northern Zambia; southern Tanzania, eastern DR Congo Chamaesaura tenuior Günther, 1895 Chamaesaura tenuior Günther, 1895: 524 Type locality: Kampala, Uganda. Syntypes: : BMNH Distribution: Kenya: central highlands; northern Tanzania; eastern Rwanda; southern Uganda; northeastern DR Congo Cordylus Laurenti, 1768 Laurenti, J.N Specimen medicum, exhibens synopsin reptilium emendatam cum experimentis circa venena et antidota reptilium austracorum, quod authoritate et consensu. Vienna, Joan. Thomae, 217 pp. Type species: Cordylus verus Laurenti, 1768 Cordylus angolensis (Bocage, 1895) Zonurus angolensis Bocage, 1895: 25 Type locality: Caconda, Huila District, Angola Holotype: MBL (probably lost in fire 1978) Distribution: Angola highlands., northern Namibia, southcentral DR Congo Photo: Xesko Cordylus aridus Mouton & Van Wyk, 1994 Cordylus aridus Mouton & Van Wyk, 1994: 15 Type locality: Farm Botterkraal, Prince Albert District, Western Cape Province, South Africa Holotype: SAM Distribution: South Africa: Western Cape Province

10 Cordylus beraduccii Broadley & Branch, 2002 Cordylus beraduccii Broadley & Branch, 2002: 22 Type locality: Ndukusi village, 10 km SE of Lolkisale, Kiteto District, Arusha Region, Tanzania. Holotype: NMZB Distribution: northern Tanzania and southern Kenya (plateau areas). Photo: Petra Bejuk Cordylus cloetei Mouton & Van Wyk, 1994 Cordylus cloetei Mouton & van Wyk, 1994: 12 Type locality: Farm De Hoek, Nuweveld Mountains, Fraserburg District, South Africa. Holotype: SAM Distribution: South Africa: Northern Cape Province Photo: Petra Bejuk Cordylus cordylus (Linnaeus, 1758) Lacerta cordylus Linnaeus, 1758: 202 Type locality: Africa Syntypes: NHRM Synonyms: Cordylus verus Laurenti, 1768 Type locality: Africa Holotype: not located Cordylus dorsalis Cuvier, 1829 Type locality: Africa Holotype: MNHNP, not located Cordylus griseus Cuvier, 1829 Type locality: Africa Holotype: MNHNP, not located Zonurus vertebralis Gray, 1838 Type locality: Cape of Good Hope, South Africa Syntypes: BMNH V.2.a-e, V.3.f-g Zonurus cordylus flavus Rose, 1926 Type locality: Robben Island, Western Cape Province, South Africa Holotype: not located Zonurus cordylus tasmani Power, 1930 Type locality: Dunbrody, Uitenhage District, South Africa. Photo: A Shawka Syntypes: PEM R13539, , 13522, , 13582, 13583, Distribution: South Africa: Northern, Western. Eastern Cape southeastern Free State Provinces; Lesotho

11 Cordylus imkeae Mouton & Van Wyk, 1994 Cordylus imkeae Mouton & van Wyk, 1994: 14 Type locality: Lower slopes of the Rooiberg on the farm Wolfhok (30 24'27"S, 18 06'11" E), Garies District, Namaqualand, South Africa Holotype: SAM Distribution: South Africa:Western Cape Province: Rooiberg Mts, Namaqualand Photo: Petra Beju Cordylus jonesii (Boulenger, 1891) Zonurus jonesii Boulenger, 1891: 417 Type locality: Murchison Range, northeast Transvaal, Northern Province, South Africa Holotype: BMNH Distribution: eastern Botswana; southern Zimbabwe; southern Mozambique; South Africa: Northern, Mpumalanga, KwaZulu- Natal Provinces; eastern Swaziland. Photo: C.K. Willis Cordylus machadoi Laurent, 1964 Cordylus vittifer machadoi Laurent, 1964: 49 Type locality: Leba, Humpata, near Sá da Bandeira, Húila, Angola. Holotype: MD? Distribution: Angola: Huila Province; Namibia: extreme northern Kaokoveld Cordylus macropholis (Boulenger, 1910) Zonurus macropholis Boulenger, 1910: 494 Type locality: Little Namaqualand, South Africa. Holotype: SAM 434 Distribution: South Africa: Northern Cape Province: coastal areas of Little Namaqualand Cordylus marunguensis Greebaum, Stanley, Kusamba, Moninga, Goldberg & Bursey, 2012 Cordylus marunguensis Greebaum, Stanley, Kusamba, Moninga, Goldberg & Bursey, 2012: 26 Type locality: Pepa ( S, E), Marungu Mountains, Katanga Province, DR Congo Holotype: UTEP Distribution: DR Congo: Katanga Province: Marungu Mountains

12 Cordylus mclachlani Mouton, 1986 Cordylus mclachlani Mouton, 1986: 319 Type locality: Koue Bokkeveld, near Ceres, South Africa. Holotype: SAM Distribution: South Africa: Western Cape Province: Koue Bokkeveld to Nieuwoudtville Photo: Dorse & van Rooyen Cordylus meculae Branch, Rödel & Marais, 2005 Cordylus meculae Branch, Roedel & Marais, 2005: 132 Type locality: Serra Mecula, Niassa Game Reserve (12 03'11"S, 37 38'49"E), Niassa Province, northern Mozambique Holotype: PEM R16166 Distribution: Mozambique: Serra Mecula Cordylus minor Fitzsimons, 1943 Cordylus cordylus minor FitzSimons, 1943: 458 Type locality: just north of Matjesfontein,Western Cape Province, South Africa. Syntypes: TMP Distribution: South Africa: Western Cape Province: southwestern Karoo Cordylus namakuiyus Stanley, Ceríaco, Bandeira, Valerio, Bates & Branch, 2016 Cordylus namakuiyus Stanley, Ceríaco, Bandeira, Valerio, Bates & Branch, 2016: 12 Type locality: large rock outcrop near Caraculo (15 0'59.40"S, 12 38'31.30"E), on the road from Lubango and Namibe, Namibe province, Angola Holotype: CAS Distribution: Angola: south and central parts of Namibe Province, southwest of the Leba escarpment Cordylus niger Cuvier, 1829 Cordylus niger Cuvier, 1829: 33 Type locality: Africa. Syntypes: MNHNP Synonyms: Calyptoprymnus verecundus De Vis, 1905 Type locality: unknown Holotype: QM? Zonurus cordylus atrus Power, 1930 Type locality: Cape Peninsula, South Africa. Holotype: TMP? Distribution: South Africa: Western Cape Province: Table Mountain, Cape Peninsula and Saldanha Photo: P Head Cordylus nyikae Broadley & Mouton, 2000 Cordylus nyikae Broadley & Mouton, 2000: 169 Type locality: near Chelinda Bridge, Nyika National Park (10 35'S, 33 48'E), Malawi. Holotype: NMZB Distribution: northern Malawi: Nyika Plateau, Misuku Hills and at Chitipa

13 Cordylus oelofseni Mouton & Van Wyk, 1990 Cordylus oelofseni Mouton & van Wyk, 1990: 34 Type locality: Dasklip Pass near Porterville, Western Cape Province, South Africa. Holotype: SAM Distribution: South Africa: Western Cape Province: western borders of the Cape Fold Mountains Photo: TH Cordylus rhodesianus (Hewitt, 1933) Zonurus cordylus rhodesianus Hewitt, 1933: 48 Type locality: Monte Cassino Mission, Macheke, Zimbabwe Syntypes: PEM Distribution: Zimbabwe: Eastern highlands; Mozambique: Chimanimani Mountains Cordylus rivae (Boulenger, 1896) Zonurus rivae Boulenger, 1896: 8 Type locality: Giacorsa, Gallaland, Ethiopia. Holotype: MSNG Distribution: Ethiopia: Sidamo region Cordylus tropidosternum (Cope, 1869) Zonurus tropidosternum Cope, 1869: 169 Type locality: Mozambique Holotype: MCZ R-5742 Synonyms: Zonurus frenatus Pfeffer, 1889 Type locality: Mhonda, Tanzania Holotype: ZMH, probably lost in WWII Zonurus parkeri Cott, 1934 Type locality: Amatongas, Sofala Province, Mozambique Holotype: BMNH Distribution: southern coastal Kenya; eastern Tanzania; to central Mozambique west to southeast DR Congo (Katanga); northern Zambia; Malawi; northeastern Zimbabwe Photo: Petra Beju Cordylus ukingensis (Loveridge, 1932) Zonurus ukingensis Loveridge, 1932: 378 Type locality: Tandala, Ukinga mountains, southwestern Tanzania Holotype: MCZ Distribution: Tanzania: southern highlands, Ukinga and Udzungwa Mountains Cordylus vittifer (Reichenow, 1887) Zonurus vittifer Reichenow, 1887: 25 Type locality: Transvaal, South Africa. Holotype: ZMB Photo: M Robinson Synonym: Zonurus tropidogaster Boulenger, 1910 Type locality: Barberton, Mpumalanga Province, South Africa Holotype: not located

14 Distribution: southeastern Botswana; South Africa: Northwest, Northern, Gauteng, Mpumalanga, Free State,KwaZulu- Natal Provinces; Swaziland; southwestern Mozambique: eastern slopes of the Lebombo Range Hemicordylus Smith, 1838 Smith, A Contributions to South African zoology. Art. VI. Annals and Magazine of Natural History (1) 2: Type species: Hemicordylus capensis Smith, 1838: 32 Hemicordylus capensis (Smith, 1838) Cordylus capensis Smith, 1838: 32 Type locality: Hottentots Holland Mountains, Western Cape Province, South Africa Holotype: BMNH Synonym: Zonurus robertsi Van Dam, 1921 Type locality: Klaver, Van Rhynsdorp District, Western Cape Province, South Africa Holotype: TMP 3747 Distribution: South Africa: Western Cape Province Photo: A Shawka Hemicordylus nebulosus (Mouton & Van Wyk, 1995) Pseudocordylus nebulosus Mouton & van Wyk, 1995: 391 Type locality: northern slope of Landdroskop, Hottentots-Holland Mountains, South Africa Holotype: SAM Distribution: South Africa: Western Cape Province: Hottentots Holland Mountains Karusasaurus Stanley, Bauer, Jackman, Branch & Mouton, 2011 Stanley, E.L., A.M. Bauer, T.R. Jackman, W.R. Branch & P.L.N. Mouton Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizard (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58(1): Type species: Cordylus polyzonus Smith, 1838: 34 Karusasaurus jordani (Parker, 1936) Zonurus jordani Parker, 1936: 133 Type locality: Hoffnung, near Windhoek, Namibia Holotype: BMNH Distribution: central Namibia, north of Aus Karusasaurus polyzonus (Smith, 1838) Cordylus polyzonus Smith, 1838: 31 Type locality: South Africa Syntypes: BMNH Distribution: South Africa: Northern Cape, Western Cape, Eastern Cape, Free State Provinces; southern Namibia Photo: Gawie

15 Namazonurus Stanley, Bauer, Jackman, Branch & Mouton, 2011 Stanley, E.L., A.M. Bauer, T.R. Jackman, W.R. Branch & P.L.N. Mouton Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizard (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58(1): Type species: Zonurus pustulatus Peters, 1862: 19 Namazonurus campbelli (Fitzsimons, 1938) Zonurus campbelli Fitzsimons, 1938: 189 Type locality: Farm Barby, 25 km west of Helmeringhausen, Great Namaqualand, Namibia Holotype: TMP Distribution: Namibia: Bethanie District Photo: J Marais Namazonurus lawrenci (Fitzsimons, 1939) Zonurus lawrenci Fitzsimons, 1939: 208 Type locality: Lekkersing, Richtersveld, Little Namaqualand, South Africa Holotype: SAM Distribution: South Africa: North Cape Prov., Richtersveld; southwestern Namibia Namazonurus namaquensis (Methuen & Hewitt, 1914) Zonurus namaquensis Methuen & Hewitt, 1914: 137 Type locality: Farm Narudas Süd, Great Karasberg District, Great Namaqualand, Namibia Holotype: TMP 3163 Distribution: southern Namibia: Great Karasberg district Namazonurus peersi (Hewitt, 1932) Zonurus peersi Hewitt, 1932: 116 Type locality: Garies, Little Namaqualand, South Africa Syntypes: SAM Distribution: South Africa: North Cape Prov: Little Namaqualand Namazonurus pustulatus (Peters, 1862) Zonurus pustulatus Peters, 1862: 19 Type locality: Otjimbingue, Namibia Syntypes: ZMB , 4361 Distribution: central Namibia Photo: Guy

16 Ninurta Stanley, Bauer, Jackman, Branch & Mouton, 2011 Stanley, E.L., A.M. Bauer, T.R. Jackman, W.R. Branch & P.L.N. Mouton Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizard (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58(1): Type species: Zonurus coeruleopunctatus Hewitt & Methuen,1913: 110 Ninurta coeruleopunctatus (Methuen & Hewitt, 1913) Zonurus coeruleopunctatus Methuen & Hewitt, 1913: 110 Type locality: Buffel s Nek, between Knysna and Avontuur, South Africa Holotype: TMP? Distribution: South Africa: Western Cape & Eastern Cape provinces: Mossel Bay to Witelsbos Photo: Shaun Swanepoel Ouroborus Stanley, Bauer, Jackman, Branch & Mouton, 2011 Stanley, E.L., A.M. Bauer, T.R. Jackman, W.R. Branch & P.L.N. Mouton Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizard (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58(1): Type species: Cordylus cataphractus Boie, 1828: 140 Ouroborus cataphractus (Boie, 1828) Cordylus cataphractus Boie, 1828: 140 Type locality: South Africa Holotype: RMNH 3603 Synonym: Cordylus nebulosus Smith, 1838 Type locality: South Africa Syntypes: BMNH Distribution: South Africa: southern Richtersveld, Northern Cape Prov., to Matjiesfontein, Western Cape Prov. Platysaurus Smith, 1844 Smith, A Illustrations of the zoology of South Africa, Reptilia. Smith, Elder, and Co., London Type species: Platysaurus capensis Smith, 1844: pl 40 Platysaurus attenboroughi Whiting, Branch, Pepper & Keogh, 2015 Platysaurus attenboroughi Whiting, Branch, Pepper & Keogh, 2015: 185 Type locality: Fish River Canyon (27 52'21.7 S, 17 31'15.7 E), Karasburg District, southern Namibia Holotype: TMP Distribution: South Africa: Northern Cape Province: lower Orange River from Goodhouse to the Richtersveld; southern Namibia: Hunsberg, Huamsib and Ploegberg Mountains and the Fish River Canyon Photo: M Whiting

17 Platysaurus broadleyi Branch & Whiting, 1997 Platysaurus broadleyi Branch & Whiting, 1997: 126 Type locality: Augrabies Falls National Park, Gordonia District, Northern Cape Province, South Africa Holotype: TMP Distribution: South Africa: Northern Cape Province: from Augribies to Klein Pella Photo: B Dupont Platysaurus capensis Smith, 1844 Platysaurus capensis Smith, 1844: pl 40 Type locality: Kamiesberg region, Namaqualand, Northern Cape Province, South Africa Lectotype: BMNH Distribution: South Africa: Northern Cape Province: greater Kamiesberg region, from Carolusberg to near Springbok south of Garies; southern Namibia: southern Lüderitz, Bethanie, and Karasburg districts Platysaurus guttatus Smith, 1849 Platysaurus guttatus Smith, 1849: 8 Type locality: near the Limpopo River, South Africa Holotype: BMNH Distribution: South Africa: Northwest, Limpopo provinces; eastern Botswana; southwestern Mozambique; southern Zimbabwe Photo: G Matz Platysaurus imperator Broadley, 1962 Platysaurus imperator Broadley, 1962: 816 Type locality: Matowa, 60 km NE of Mtoko, Zimbabwe Holotype: NMZB 4819 Distribution: northeast Zimbabwe; northwestern Mozambique Photo: ElinorD

18 Platysaurus intermedius Matschie, 1891 Platysaurus intermedius intermedius Matschie, 1891 Platysaurus intermedius Matschie, 1891: 606 Type locality: Station Mphome, near Haenertsburg, Limpopo Province, South Africa Lectotype: ZMB 3466 Distribution: South Africa: Limpopo Province: Soutpansberg Mountains Platysaurus intermedius inopinus Jacobsen, 1994 Platysaurus intermedius inopinus Jacobsen, 1994: 139 Type locality: Farm Glen Alpine 304LR, 23 12' S, 28 40'E, Potgietersruts (=Mokopane) district, Limpopo Province, South Africa Holotype: TMP Distribution: South Africa: Limpopo Province: foothills of the Blouberg Platysaurus intermedius natalensis Fitzsimons, 1948 Platysaurus guttatus natalensis Fitzsimons, 1948: 74 Type locality: Louwsburg, northern KwaZulu-Natal Holotype: TMP Distribution: South Africa: KwaZulu-Natal Province; Swaziland Platysaurus intermedius nigrescens Broadley, 1981 Platysaurus intermedius nigrescens Broadley, 1981: 161 Type locality: Shoshong Hills ( S, E), 20km west of Mahalapye, Botswana Holotype: NMZB Distribution: northeastern Botswana Photo: Sputniktilt Platysaurus intermedius nyasae Loveridge, 1953 Platysaurus guttatus nyasae Loveridge, 1953: 237 Type locality: rocks by roadside on Blantyre-State road south of Mpatamaya Bridge, Malawi Holotype: MCZ R Distribution: Malawi; west-central Mozambique Platysaurus intermedius parvus Broadley, 1976 Platysaurus intermedius parvus Broadley, 1976:? Type locality: Blouberg, Soutpansberg District, Limpopo Province, South Africa. Holotype: TMP? Distribution: South Africa: Limpopo Province: Blouberg Massif Platysaurus intermedius rhodesianus Fitzsimons, 1941 Platysaurus guttatus rhodesianus Fitzsimons, 1941: 31 Type locality: Vumba Mountain, Zimbabwe Holotype: TMP? Distribution: southern Zimbabwe; northeastern Botswana; South Africa: northern Limpopo Province Platysaurus intermedius subniger Broadley, 1962 Platysaurus guttatus subniger Broadley, 1962: 814 Type locality: Watsomba, a granite peak 16 miles NNW of Umtali, Zimbabwe Holotype: NMZB 1392 Distribution: northeastern Zimbabwe Platysaurus intermedius wilhelmi Hewitt, 1909 Platysaurus wilhelmi Hewitt, 1909: 32

19 Type locality: Nelspruit, Barberton District, Mpumalanga province, South Africa Holotype: TMP? Distribution: South Africa: Mpumalanga and KwaZululand-Natal provinces; Swaziland Platysaurus lebomboensis Jacobsen, 1994 Platysaurus lebomboensis Jacobsen, 1994: 140 Type locality: Mananga hill (25 58'S, 31 52'E), Barberton district, Mpumalanga province, South Africa Holotype: TMP Distribution: South Africa: KwaZululand Province: Middle Lebombo Range; southern Mozambique; Swaziland Photo: J Marais Platysaurus maculatus Broadley, 1965 Platysaurus maculatus maculatus Broadley, 1965 Platysaurus maculatus maculatus Broadley, 1965: 3 Type locality: Mitucué Mountain, Niassa Province, Mozambique Holotype: NMZB 8054 Distribution: southern Tanzania; northern Mozambique Platysaurus maculatus lineicauda Broadley, 1965 Platysaurus maculatus lineicauda Broadley, 1965: 3 Type locality: 14 mi. west of Morrumbala, Zambezia Province, Mozambique Holotype: NMZB? Distribution: Mozambique: Zambezia Province Platysaurus minor Fitzsimons, 1930 Platysaurus guttatus minor Fitzsimons, 1930: 30 Type locality: at Vygeboompoort, Waterberg District, north Limpopo Province, South Africa Syntypes: TMP 2109, 2367 Distribution: South Africa: Limpopo Province: Waterberg Massif to the Blouberg Massif Photo: R van Huyssteen Platysaurus mitchelli Loveridge, 1953 Platysaurus mitchelli Loveridge, 1953: 234 Type locality: Ruo River Forest, Mt Mlanje, Malawi Holotype: MCZ R Distribution: southern Malawi: Mulanje Mountains Platysaurus monotropis Jacobsen, 1994 Platysaurus monotropis Jacobsen, 1994: 136 Type locality: Farm Blackhill 317 LR, Potgietersrust district, Limpopo Province, South Africa Holotype: TMP Distribution: South Africa: Limpopo Province, foothills of the Blouberg

20 Platysaurus ocellatus Broadley, 1962 Platysaurus ocellatus Broadley, 1962: 810 Type locality: western slopes of the Chimanimani Mts., above Dead Cow Camp, 8 miles east of Melsetter, Zimbabwe Holotype: NMZB 1199 Distribution: east-central Zimbabwe: Chimanimani Mountains; west-central Mozambique Photo: Brian Gratwicke Platysaurus orientalis Fitzsimons, 1941 Platysaurus orientalis orientalis Fitzsimons, 1941 Platysaurus minor orientalis Fitzsimons, 1941: 280 Type locality: Sekororo, on the lower slopes of the Drakensberg, about 40 miles south of Leydsdorp, Limpopo Province, South Africa Syntypes: TMP Distribution: South Africa: Drakensberg Mountains, southern Limpopo Province, South Africa Platysaurus orientalis fitzsimonsi Loveridge, 1944 Platysaurus guttatus fitzsimonsi Loveridge, 1944: 88 Type locality: Lydenburg, Mpumalagna Prov., South Africa Holotype: MCZ 8982 Distribution: South Africa: Mpumalagna Escarpment, Mpumalagna Prov. Platysaurus pungweensis Broadley, 1959 Platysaurus pungweensis pungweensis Broadley, 1959 Platysaurus guttatus pungweensis Broadley, 1959: 314 Type locality: Pungwe River (near the Mozambique border), eastern Zimbabwe Holotype: NMZB 1975 Distribution: eastern Zimbabwe: Eastern Highlands Platysaurus pungweensis blakei Broadley, 1964 Platysaurus intermedius blakei Broadley, 1964: 1 Type locality: 15 miles south-east of Vila de Manica, Mozambique Holotype: NMZB? Distribution: western Mozambique: Chimoio Plateau, Manica Prov., Photo: Milii Platysaurus relictus Broadley, 1976 Platysaurus relictus Broadley, 1976:? Type locality: Waterpoort, Soutpansberg District, Limpopo Prov., South Africa Holotype: TMP Distribution: South Africa: Limpopo Province: northern side of the Soutpansberg Photo: R van Huyssteen Photo: Joachim

21 Platysaurus torquatus Peters, 1879 Platysaurus torquatus Peters, 1879: 10 Type locality: Tete, Mozambique Syntypes: ZMB , , MCZ R-37203, NMHW Distribution: northeastern Zimbabwe, west-central Mozambique, southern Malawi Pseudocordylus Smith, 1838 Smith, A Contributions to South African zoology. Art. VI. Annals and Magazine of Natural History (1) 2: Type species: Cordylus montanus Smith, 1838: 32 Pseudocordylus langi Loveridge, 1944 Pseudocordylus langi Loveridge, 1944: 73 Type locality: Mont-aux-Sources, Drakensberg, Lesotho Holotype: MCZ R Distribution: South Africa: KwaZulu-Natal: Drakensberg mountains; Lesotho Photo: Alexander R Pseudocordylus melanotus (Smith, 1838) Cordylus melanotus Smith, 1838: 32 Type locality: South Africa, hills between the main branches of the Orange River east of Philippolis, Orange Free State Holotype: lost Distribution: South Africa: northern KwaZulu-Natal, Gauteng, eastern Mpumalanga, northeastern Free State Provinces Photo: DK Pseudocordylus microlepidotus (Cuvier, 1829) Pseudocordylus microlepidotus microlepidotus (Cuvier, 1829) Cordylus microlepidotus Cuvier, 1829: 33 Type locality: South Africa Holotype: iconotype, based on Seba pl 62 Synonyms: Zonurus wittii Schlegel, 1834 Type locality: South Africa Holotype: not located Photo: nvlooi

22 Cordylus montanus Smith, 1838 Type locality: South Africa Holotype: BMNH, not located Distribution: South Africa: Cape Fold Mountains, from Cedarberg to Port Elizabeth, Western Cape and Eastern Cape Provinces. Pseudocordylus microlepidotus fasciatus (Smith, 1838) Cordylus fasciatus Smith, 1838: 32 Type locality: South Africa, rocky hills near Grahamstown Holotype: lost Synonym: Cordylus algoensis Smith, 1838 Type locality: Algoa Bay, Cape Province, South Africa Holotype: BMNH Distribution: South Africa: Eastern Cape Province Pseudocordylus microlepidotus namaquensis Hewitt, 1927 Pseudocordylus microlepidotus namaquensis Hewitt, 1927: 392 Type locality: Namaqualand, Northern Cape Prov., South Africa Holotype: SAM 872 Distribution: South Africa: Northern Cape Prov.: Nuweveld Mountains, from Sutherland to Beaufort West Pseudocordylus spinosus Fitzsimons, 1947 Pseudocordylus spinosus Fitzsimons, 1947: 116 Type locality: Cathkin Peak area, Drakensberg, KwaZulu- Natal Province, South Africa Holotype: TMP Distribution: South Africa: lower slopes of Drakensberg, from Giant's Castle in KwaZululand-Natal to Golden Gate in Free State Photo: T Ping Pseudocordylus subviridis (Smith, 1838) Cordylus subviridis Smith, 1838: 33 Type locality: South Africa Holotype: lost Distribution: South Africa: from Amatola Mountains, Eastern Cape Province, to Mont-Aux-Sources (Drakensberg), Lesotho Photo: Armada44 Pseudocordylus transvaalensis Fitzsimons, 1943 Pseudocordylus subviridis transvaalensis Fitzsimons, 1943: 469 Type locality: Woodbush, Pietersburg District, Limpopo Province, South Africa Syntypes: TMP 1695, 1699 Distribution: South Africa: southern Limpopo Province Photo: Arno Meintjs

23 Smaug Stanley, Bauer, Jackman, Branch & Mouton, 2011 Stanley, E.L., A.M. Bauer, T.R. Jackman, W.R. Branch & P.L.N. Mouton Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizard (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58(1): Type species: Cordylus giganteus Smith, 1844: pl 35 Smaug barbertonensis (Van Dam, 1921) Zonurus barbertonensis Van Dam, 1921: 240 Type locality: Barberton, Mpumalanga Province, South Africa Holotype: TMP 4273 Distribution: South Africa: southeastern Mpumalanga, northern KwaZululand-Natal; eastern Swaziland Photo: DR McKenzie Smaug breyeri (Van Dam, 1921) Zonurus breyeri V Van Dam, 1921: 239 Type locality: Farm Geelhoutkop, Waterberg District, Limpopo Province, South Africa Holotype: ZMA Distribution: South Africa: Limpopo Province: Waterberg Massif area Photo: L Ghoul Smaug depressus (Fitzsimons, 1930) Zonurus barbertonensis depressus Fitzsimons, 1930: 24 Type locality: Newgate, near Louis Trichardt, Limpopo Province, South Africa Holotype: TMP 4621 Distribution: South Africa: Limpopo Province: middle and eastern Soutpansberg, to northern Krüger National Park and south to Duiwelskloof

24 Smaug giganteus (Smith, 1844) Cordylus giganteus Smith, 1844: 35 Type locality: Interior districts of South Africa on pinnacles of Quathlamba Mountains Syntypes: BMNH , Synonym: Zonurus derbianus GRAY, 1845 Type locality: South Africa Syntypes: BMNH Distribution: South Africa: northeastern Free State, extreme KwaZululand-Natal, southeastern Mpumalanga Smaug mossambicus (Fitzsimons, 1958) Cordylus warreni mossambicus Fitzsimons, 1958:? Type locality: Gorongoza Mountain, near Vila Paiva, Mozambique Holotype: NMP 1237 Distribution: Mozambique: Gorongoza Mountains; Zimbabwe: Chimanimani Mountains Photo: KS Bader Smaug regius (Broadley, 1962) Cordylus warreni regius Broadley, 1962: 808 Type locality: Dora, 8 miles S.S.W. of Umtali, Zimbabwe Holotype: NMZB 1210 Distribution: Zimbabwe: southeastern Mutare District: Bvumba Mountains Photo: J Pfleiderer Smaug vandami (Fitzsimons, 1930) Zonurus vandami Fitzsimons, 1930: 25 Type locality: Gravelotte, near Leydsdorp, Limpopo Province, South Africa Holotype: TMP 7407 Synonym: Zonurus vandami perkoensis Fitzsimons, 1930 Type locality: Farm Perkoe, Olifant River District, Mpumalanga Province Holotype: TMP Distribution: South Africa: Mpumalanga Province: eastern escarpment of Mpumalanga Photo: scavenger

25 Smaug warreni (Boulenger, 1908) Zonurus warreni Boulenger, 1908: 232 Type locality: Ubombo, Zululand Holotype: BMNH Distribution: South Africa: Lebombo Range, southeastern Mpumalanga, northern KwaZululand-Natal Provinces; eastern Swaziland Photo: James Culverwell ACKNOWLEGEMENTS This work could not have been completed without the help of numerous individuals. We thus acknowledge the help and advice of Patrick D Campbell (BMNH), Esther Dondorp (RMNH), Lemmy Mashinini (TMP), Werner Conradie (PEM), Jan Grathwohl The author expresses his to the photographers who have contributed to this work, in particular, James Culverwell, J. Pfleiderer, K. S. Bader, Petra Bejuk, Abu Shawka, C.K. Willis and many others. If any of the readers have any of the missing data (designated by?), I would appreciate if they would contact me

26 REFERENCES Boie, F Über eine noch nicht beschriebene Art von Cordylus Gronov. Cordylus cataphractus Boie. Nova Acta Academiae Caesareae Leopoldino-Carolinae, (Halle), 14 (1): Bocage, J.V. du B Herpétologie d'angola et du Congo. Lisbon: Imprimerie Nationale, i-xx, 203 pp. Boulenger, G. A Description of a new lizard of the genus Zonurus from the Transvaal. Annals and magazine of natural history (6) 7: 417 Boulenger, G.A Second report on additions to the lizard collection in the Natural History Museum. Proceedings of The Zoological Society of London 1894: [1895] Boulenger, G.A A list of the reptiles and batrachians collected by the late Prince Eugenio Ruspoli in Somaliland and Gallaland in Annali del Museo civico di storia naturale di Genova, Giacomo Doria, (2) 17: 5-14 Boulenger, G On a collection of fresh-water fishes, batrachians, and reptiles from Natal and Zululand, with description of new species. Annals of the Natal Government Museum (Pietermaritzburg) 1(3): Boulenger, G.A A revised list of the South African reptiles and batrachians, with synoptic tables, special reference to the specimens in the South African Museum, and descriptions of new species. Annals of the South African Museum 5: Branch, W.R., M.O. Rödel and J. Marais A new species of rupicolous Cordylus Laurenti 1768 (Sauria: Cordylidae) from Northern Mozambique. African Journal of Herpetology 54 (2): Branch, W.R. & M.J. Whiting A new Platysaurus (Squamata: Cordylidae) from the Northern Cape Province, South Africa. African Journal of Herpetology 46 (2): Broadley, D.G Sauria, Cordylidae: Platysaurus guttatus pungweensis n. subsp. Occasional Papers of the National Museum of Southern Rhodesia 23 (B): Broadley, D.G On some reptile collections from the North-Western and North-Eastern Districts of Southern Rhodesia , with descriptions of four new lizards. Occasional Papers of the National Museum of Southern Rhodesia 26 (B): Broadley, D.G A new Platysaurus from central Mozambique. Arnoldia (Rhodesia) 1 (5): 1-2 Broadley, D.G Two new forms of Platysaurus from the northern Transvaal (Sauria: Cordylidae). Arnoldia (Rhodesia) 8 (8): 1-3 Broadley, D.G A revision of the Genus Platysaurus A. Smith. Occasional Papers of the National Museum of Southern Rhodesia B, 6 (4): Broadley, D.G A new subspecies of Platysaurus intermedius from Botswana (Sauria: Cordylidae). Botswana Notes and Records, 13: 161 Broadley, D.G & P.le FrasN. Mouton A new species of rupicolous Cordylus Laurenti from Malawi (Sauria: Cordylidae). African Journal of Herpetology 49 (2): Broadley, D. G. & W. R. Branch A review of the small east African Cordylus (Sauria: Cordylidae), with the description of a new species. African Journal of Herpetology 51 (1): 9-34

27 Cope, E.D Notes upon some reptiles of the Old World. Proceedings of the Academy of Natural Sciences of Philadelphia 14: Cope, E.D Seventh contribution to the herpetology of tropical America. Proceedings of the American Philosophical Society, 11: [1869] Cott, H The Zoological Society's expedition to the Zambesi, No. 5. On a collection of lizards, mainly from East Africa, with description of new species of Zonurus, Monopeltis, and Chirindia. Proceedings of The Zoological Society of London (no. 5): Cuvier, G. J. L. N. F. D Le Regne Animal Distribué, d'apres son Organisation, pur servir de base à l'histoire naturelle des Animaux et d'introduction à l'anatomie Comparé. Nouvelle Edition. Vol. 2. Les Reptiles. Déterville, Paris, i-xvi, de Vis, C.W A new genus of lizard. Annals of the Queensland Museum (Brisbane) 6: FitzSimons, V.F Descriptions of new South African Reptilia and Batrachia, with distribution records of allied species in the Transvaal Museum collection. Annals of the Transvaal Museum 14: Fitzsimons,V.F.M Transvaal Museum Expedition to South-West Africa and Little Namaqualand, May to August Reptiles and Amphibians. Annals of the Transvaal Museum (Pretoria) 19 (2): Fitzsimons,V.F.M Descriptions of some new species and subspecies of lizards from South Africa. Annals of the Transvaal Museum. (Pretoria) 20 (1): 5-16 Fitzsimons,V.F.M Descriptions of some new lizards from South Africa and a frog from southern Rhodesia. Annals of the Transvaal Museum (Pretoria) 20 (3): FitzSimons, V.F The lizards of South Africa. Transvaal Museum Memoir No.1 (Pretoria), 528 pp. Fitzsimons,V.F.M Descriptions of new species and subspecies of reptiles and amphibians from Natal, together with notes on some other little known species. Annals of the Natal Museum (Pietermaritzburg) 11 (1): Fitzsimons,V.F.M Notes on some reptiles and amphibians from the Darkensberg, together with a description of a new Platysaurus, from northern Natal. Annals of the Transvaal Museum (Pretoria) 21 (1): FitzSimons, V.F A new Cordylus from Gorongoza, Mocambique. Annals of the Natal Museum (Pietermaritzburg) 14 (2): Fitzinger, L Systema Reptilium, fasciculus primus, Amblyglossae. Braumüller et Seidel, Wien: 106 pp Gray, J.E Catalogue of the slender-tongued saurians, with descriptions of many new genera and species. Part 3. Annals and magazine of natural history (1) 1: Gray, J.E Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of the British Museum/Edward Newman, London: xxvii pp. Greenbaum, E., E.L. Stanley, C. Kusamba, W.M. Moninga, S.R. Goldberg & C.R. Bursey A new species of Cordylus (Squamata: Cordylidae) from the Marungu Plateau of south-eastern Democratic Republic of the Congo. African Journal of Herpetology 61 (1): Günther, A Notice of Reptiles and Batrachians collected in the eastern half of tropical Africa. Annals and magazine of natural history (6) 15:

28 Hewitt, J Description of a new species of Platysaurus and notes on the specific characters of certain species of Zonuridae, together with synoptical keys to all the known South African species and a résumé of our knowledge on their distribution: and a key to the known Genera of South African Lizardst Annals of the Transvaal Museum (Pretoria) 2 (1): Hewitt, J Further descriptions of reptiles and batrachians from South Africa. Records of the Albany Museum (Grahamstown) 3 (5): Hewitt, J Some new species and subspecies of South African batrachians and lizards. Annals of the Natal Museum (Pietermaritzburg) 7 (1): Hewitt, J Descriptions of some new reptiles and a frog from Rhodesia. Occasionnel Papers of the Rhodesian Museum 2: Jacobsen, N. H.G The Platysaurus intermedius complex (Sauria: Cordylidae) in the Transvaal, South Africa, with descriptions of three new taxa. South African Journal of Zoology 29 (2): Laurent, R.F Reptiles et batraciens de l'angola (troisième note). Companhia de Di amantes de Angola (Diamang), Serviços Culturais, Museu do Dundo (Angola), No. 67, 165 pp. Laurenti, J.N Specimen medicum, exhibens synopsin reptilium emendatam cum experimentis circa venena et antidota reptilium austracorum, quod authoritate et consensu. Vienna, Joan. Thomae, 217 pp Linnaeus, C Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii, Holmiæ. 10th Edition: 824 pp Loveridge, A New reptiles and amphibians from Tanganyika Territory and Kenya Colony. Bulletin of the Museum of Comparative Zoology at Harvard College 72: Loveridge, A Revision of the African lizards of the family Cordylidae. Bulletin of the Museum of Comparative Zoology at Harvard College 95 (1): Loveridge, A Zoological Results of a Fifth Expedition to East Africa, III. Reptiles from Nyasaland and Tete. Bulletin of the Museum of Comparative Zoology at Harvard, 110 (3): Mertens, R Reptilien und Amphibien aus dem südlichen Inner-Afrika. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt, vol. 435, p Hewitt, J. & Methuen, P.A Descriptions of some new Batrachia and Lacertilia from South Africa. Transactions of the Royal Society of South Africa (Cape Town) 3: Matschie, G.F.P Ueber eine kleine Sammlung von Reptilien und Amphibien aus Süd-Afrika. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere. (Jena) 5: Methuen, P.A. & Hewitt, J The Percy Sladen Memorial Expedition to Great Namaqualand Records and descriptions of the reptiles and batrachians of the collection. Annals of the Transvaal Museum (Pretoria) 4 (3): Mouton, P. le F.N Description of a new species of Cordylus Laurenti (Reptilia: Cordylidae) from the southwestern Cape, South Africa. South African Journal of Zoology 21 (4) 1986:

29 Mouton, Ple FN. & J.H. van Wyk Taxonomic status of the melanistic forms of the Cordylus cordylus complex (Reptilia: Cordylidae) in the south-western Cape, South Africa. South African Journal of Zoology 25 (1): Mouton, P. Le F.N. & J.H. Van Wyk Taxonomic status of geographical isolates in the Cordylus minor complex (Reptilia: Cordylidae): A description of three new species. The Journal of the Herpetological Association of Africa 43: Mouton, P. Le F.N. & J.H. Van Wyk A new crag lizard from the Cape Folded Mountains in South Africa. Amphibia-Reptilia 16 (4): Parker, H.W Dr. Karl Jordan's expedition to South-West Africa and Angola: herpetological collections. Novitates Zoologicae (London) 40: Peters, W.C.H Übersicht einiger von dem, durch seine afrikanische Sprachforschungen, rühmlichst bekannten, Hrn. Missionär C.H. Hahn bei Neu-Barmen, im Hererolande, and der Westküste von Afrika, im 21 südl. Br. gesammelten Amphibien, nebst Beschreibungen der neuen Arten. Monatsberichte der Königlichen Preussische Akademie des Wissenschaften zu Berlin 1862: Peters,W.C.H Eine neue Art der Säugetiergattung Hyrax (H. nigricans) aus Chinchoxo und über eine neue Eidechse, Platysaurus torquatus, aus Mossambique. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1879 (2): Pfeffer, G Übersicht der von Herrn Dr. Franz Stuhlmann in Ägypten, auf Sanzibar und dem gegenüberliegenden Festlande gesammelten Reptilien, Amphibien, Fische, Molluskten und Krebse. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 6 (2): 1-36 Power, J.H On the South African species of the genus Zonurus. Annals of the Transvaal Museum (Pretoria) 14 (1): Pyron, R.A., F.T. Burbrink and J.J. Wiens A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes. BMC Evolutionary Biology,13:93 Reichenow, A Neue Wirbelthiere des Zoologischen Museums in Berlin. Zoologischer Anzeiger 10: Rose, W Some notes on the lizards of the Cape Peninsula. Annals of the South African Museum 20: Schlegel, H Zonurus wittii. Tijdschrift Voor Natuurlijke Geschiedenis en Physiologie 1: 218 Schneider, J.G Historiae Amphibiorum naturalis et literariae. Fasciculus secundus continens Crocodilos, Scincos, Chamaesauras, Boas. Pseudoboas, Elapes, Angues. Amphisbaenas et Caecilias., Frommani, Jena. 364 pp Smith, A Contributions to South African zoology. Art. VI. Annals and Magazine of Natural History (1) 2: Smith, A Illustrations of the zoology of South Africa, Reptilia. Smith, Elder, and Co., London Smith, A Illustrations of the zoology of South Africa, Reptilia. Smith, Elder, and Co., London

30 Stanley, E.L., A.M. Bauer, T.R. Jackman, W.R. Branch & P.L.N. Mouton Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizard (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58(1): Stanley, E.L., L.M.P. Ceríaco, S. Bandeira, H. Valerio, M.F. Bates & W.R. Branch A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola, with the description of a new species from the Pro-Namib desert. Zootaxa 4061 (3): Van Dam, G.P.F., Descriptions of new species of Zonurus, and notes on the species of Zonurus occurring in the Transvaal. Annals of the Transvaal Museum 7, pls. i iv. Whiting, M.J., W.R. Branch, M. Pepper & J.S Keogh A new species of spectacularly coloured flat lizard Platysaurus (Squamata: Cordylidae: Platysaurinae) from southern Africa. Zootaxa 3986 (2): TAXON INDEX A aenea (see Chamaesaura) algoensis (see Pseudocordylus microlepidotus) angolensis (see Cordylus) anguina (see Chamaesaura) aridus (see Cordylus) atrus (see Cordylus niger) attenboroughi (see Platysaurus) B barbertonensis (see Smaug) beraduccii (see Cordylus) blakei (see Platysaurus pungweensis) breyeri (see Smaug) broadleyi (see Platysaurus) C campbelli (see Namazonurus) capensis (see Hemicordylus) capensis (see Platysaurus) cataphractus (see Ouroborus) cloetei (see Cordylus) coeruleopunctatus (see Ninurta) cordylus (see Cordylus) D

31 derbianus (see Smaug giganteus) depressus (see Smaug) dorsalis (see Cordylus cordylus) E F fasciatus (see Pseudocordylus microlepidotus) fitzsimonsi (see Platysaurus) flavus (see Cordylus cordylus) frenatus (see Cordylus tropidosternum) G giganteus (see Smaug) griseus (see Cordylus cordylus) guttatus (see Platysaurus) H I imkeae (see Cordylus) imperator (see Platysaurus) inopinus (see Platysaurus intermedius) intermedius (see Platysaurus) J jonesii (see Cordylus) jordani (see Karusasaurus) K L langi (see Pseudocordylus) lawrenci (see Namazonurus) lebomboensis (see Platysaurus) lineicauda (see Platysaurus maculatus) M

32 machadoi (see Cordylus) macrolepis (see Chamaesaura) macropholis (see Cordylus) maculatus (see Platysaurus) marunguensis (see Cordylus) mclachlani (see Cordylus) meculae (see Cordylus) melanotus (see Pseudocordylus) microlepidotus (see Pseudocordylus) minor (see Cordylus) minor (see Platysaurus) miopropus (see Chamaesaura) mitchelli (see Platysaurus) monotropis (see Platysaurus) montanus (see Pseudocordylus microlepidotus) mossambicus (see Smaug) N namakuiyus (see Cordylus) namaquensis (see Namazonurus) namaquensis (see Pseudocordylus microlepidotus) natalensis (see Platysaurus intermedius) nebulosus (see Hemicordylus) nebulosus (see Ouroborus cataphractus) nigrescens (see Platysaurus intermedius) niger (see Cordylus) nyasae (see Platysaurus intermedius) nyikae (see Cordylus) O ocellatus (see Platysaurus) oelofseni (see Cordylus) oligopholis (see Chamaesaura) orientalis (see Platysaurus) P parkeri (see Cordylus tropidosternum) parvus (see Platysaurus intermedius) peersi (see Namazonurus) perkoensis (see Smaug vandami) polyzonus (see Karusasaurus) pungweensis (see Platysaurus)

33 pustulatus (see Namazonurus) Q R regius (see Smaug) relictus (see Platysaurus) rhodesianus (see Cordylus) rhodesianus (see Platysaurus intermedius) rivae (see Cordylus) robertsi ( see Hemicordylus capensis) spinosus (see Pseudocordylus) subniger (see Platysaurus intermedius) subviridis (see Pseudocordylus) S T tasmani (see Cordylus cordylus) tenuior (see Chamaesaura) torquatus (see Platysaurus) transvaalensis (see Pseudocordylus) tropidosternum (see Cordylus) tropidogaster (see Cordylus vittifer) U ukingensis (see Cordylus) vandami (see Smaug) verecundus (see Cordylus niger) vertebralis see Cordylus cordylus) verus (see Cordylus cordylus) vittifer (see Cordylus) V W warreni (see Smaug) wilhelmi (see Platysaurus intermedius) wittii (see Pseudocordylus microlepidotus)

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