Ыеч New апа and little known species speciеs of ог flies of ог

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1 ( G 1 6 Russian Епото1. Entomol. J. 4 (1-4): RUSSIAN RUSSIАN ENTOMOLOGICAL JOURNAL,I JouRNАL,~1995 j Ыеч New апа and little known species speciеs of ог flies of ог Еаихаnцдае Lauxaniidae and Strongylophthalrnyii.dae Strongylophthalmyiidae (Diptera) Новые H0131ile и a малоизвестные maaon3bectnbie BJaAbl виды мух myx семейств cemeilcrb Lauxaniidaе Lauxaniidae T4. и Strongylophthalmyцdae Strongylophthalmyiidae (Diptera) А.I. A.I. Shatalkiп Shatalkin А.И. A.14. Шаталкин Illaramcvni Zoological Мивеит, Museum, Моасоа Moscow State University, Herzen Нетгеп str. вг. 6, Moscow Мовсоа K-9, К-9, Russia. Зоологический 3oo.iiorwiectool riyaerl музей МГУ, mry, yn. ул. Герцена repzieha 6, 6, Москва MocKsa К-9, K-9, 103р Россия. Poccns. КЕУ KEY WORDS: ОЮ : Lauxaniidae, Lаихапiiдае, Strongylophthalmyiidae, taxonomy, нахопоту, new пеш species. speciеs. КЛЮЧЕВЫЕ KITIOLIEBNE СЛОВА: CJIOBA: Lauxaniiдae, Lauxaniidae, Strongylophthalrnyiidae, Straпgylaphthalmyiidae, Takcconomm, таксоlгомия, nonme новые виды. nnam. АВТНАСТ: ABSTRACT: Тhе The 1о11о' following п taxa аха аге are described: Institut, Eberswalde, ЕЬега11е, Germany; Сегтпапу; ТА TAU Entomo- Епноп1оlogical Collection, Departnment Department of of Zoology, Zоо1оуу, Tel Те1 family Lauxaniidae Са11горит Calliopum саисавiсит caucasicum вр.п. sp.n. С. C. saklтaliпicum sakhalinicum вр.п., sp.n., Minettiella elbergi sp.n., Aviv University, Israel; ПЗМ USNM Uпited United States 5tates Pachycerina ninae $р.п., Sapromyza cerata sp.n., р.п., National Мивеит, Museum, Washington, УаЫаоа, ВС, DC, ВА; USA; ZISP S. ferganica вр.п., sp.n., S. hissarica вр.п., sp.n., S. persica Zoological Institute lпstitute of оf the Ье Russian Russiaп Academy Асадехпу of оf вр.п., sp.n., S. picea вр.п., sp.n., S. ravida вр.п. sp.n. S. romanovi romanoai Sciences, St.Peteгsburg, Petersburg, Russia; М1ТН ZMUH Zoological Museum Мивеигп Uпiversity University оf of Helsinki, Finland; Finlaпd; sp. n.,s. ziminae zimivae sp.n.; family idae Strongyloplzthalтnyia Strongylophthalrnyia freidbergi вр.п., sp.n., S. ZМцМ ZMUM Zaological Zoological Museum Миsеилп ое of Moscow State freyi sp.n., р.п., S. indica iтгдiса вр. sp.n., п., S. S. microstyla microsty1a sp.n., р.п., University, Russia. Strongylophthalmyiгдае S.pectiпigera S.pectinigera sp.n., р.п., S. spinipalpa вр. sp.n., п., S. cera sp.n., р.п., S. verrucifera sp.n. 5р.п. The Тне key сеу to the Family Lauxaniidae Lauxaпiidae species of оf S. crinita-group is given. ЗарТо7пу2а Sapromyza styloсета citriпella citrinella пот.п. nom.n. fоr for S. citrina Shatalkin Shatalkiп and Неге Here 1 I continue а a discussion discussioп of оf the 1ис Palaearctic Paiaearctic species Sapromyza remmae тетвае повп.п. nom.n. for S. micropyga Е. E. ОЕ of the Lauxaniidae. This ЁЬеше theme was та внаrtед started Ьу by т.е me гп in the Remm et Elberg are three papers that came off earlier (Shatalkin, Аепип е1 Е1Ьет are proposed. 1игее рарега Ьа1 саше ое the press еат1гег [Shatalkiп, 1992a,b, а, 1993a1. а). РЕЗЮМЕ: PE3IOME: B В pa6ote работе orincanbi описаны ciie,aytoitine следуюиние Ta- таксоны: KCOlibi: семейство cemeilmo Lauxaпiidae Lauxaniidae Са11iорит Calliopum caucasicum sp.n., р.п., С. C. sakhalinicum вр.п., sp.n., Minettiella elbergi вр.п., sp.n., Pachycerina пгпае ninae sp.n., р.л., Sapromyza cerata sp.n., S. ferganica sp.n., sp.n.., S. hissarica вр.п., sp.n., саисавгсит S. persica вр. sp.n., п., S. picea яр.в., sp.n., S. ravida вр. sp.n. п. S. S. romanovi sp.n.,s. р - п. ziтinae ziminaesp.n.; зрп.; ceme11 семейство CTBO Strongylophthalmyiidae Strongylophthalmyia Straпgylophthalmyia freidbergi sp.n.,s., freyi sp_n.,s. n, indica вр.п., sp. n, S. microstyla вр. sp.n. п., S S. pectinigem pectinigera вр sp.п..n., S. spinipalpa sp s р..n п., S. stylocera sp.n., р.п., S. verrucifera вр.п. sp.n. Rana Дана rra6anna таблица для A.1131 определения onpe,tte.rienim видов BH,LIOB группы rpynrim S. crinita. Предложены Sарготуха Sapromyza citrinella norrr.n. norri.n. AAA для S. citri- flpe,a-.rioacenbi па na Shatalkin п и H Sapromyza геттае remmae попа.п. nom.n. A.rm для S. тicropyga micropyga Е. E. Remit Аегппн et е Elberg. Е1Ьег. The work summarizes the the геп results оf of the examination of ое material танегга1 represented гергевепнед in the Zoological ехатiпанiоп Мивеит Museum ое of Мосо Moscow State University апд and also а1 о of о some sолпе collections borrowed boгrowed fгот from а a пилиbeг number оf of institutions. The Тне following followiпg ebbгeviations abbreviations аге are used for tиtians. the institutions holding Ьо1с1iп лиа1егiа1 material studied in the ргееп1 present paper: рарег: DEI ВЕ' Deutsches Deuisches Entomologisches Eпtozologisches Селив Genus Calliopum Strand, 1927 Calliopum albomaculatum albomaculatuтn Strobl, 5 гоь1, 1909 МАТЕ1 MATERIAL?,, Sakhaliп, Sakhalin, Yuzhno-Sakhaliпsk, Yuzhno-Sakhalinsk, "Chekhova" Peak, 27.VII У (Nartshuk). DIAGNOSIS. С. C. albomaculatum а1ьоишси[а -ит is characterized by Ьу the black Ыаск halteres,entirely halteres,entiтely black Ыаск legs and апд parafacials ди8нед dusted with silver впег from Ье1о below the апнеппае antennae to the тоинн. mouth. L. Papp Рарр has [ха meпtioned mentioned (personal (регвопа1 communication) соттипiсанiоп) this spccies species 1о to Ье be оипд found iп in Primorslйy Primorskiy Кгау. Kray. Iп In the Palaeaгctic Palaearctic regioп region the most шов of species of оf the genus уепив Саlггорцт Calliopum аге are кпошп known from its Western Уеаегл subregion (Europe, (Еиторе, Мог1и North Africa, Nеаг Near East). The Тне only оп1у species еs C. С. potanirтi potanini Сzегпу Czerny was 'аз descriьcd described from fтот Szechuan гесгипап (Снiпа). (China). С. C. аlьотаси1анит albomaculatum апд and С. C. sakhalinicum descriьed described bellow are the first теркевепнаtives ОЕ of this genus geпus from Far Рат East. Calliopum caucasicum саисавiсит Shatalkin Shatalkiп вр.п. sp.n. representa- Fig. 1. МАТЕюА[. MATERIAL Но1онуре: Holotype: С", d', Саосаон, Caucasus, North Osetiya, Alagir, А1аугт, 28.V У.1989 (Оегоу). (Ozerov). Paratypes: 5 о"с~', 5 d'd, 5 9?,??, same ватпе 1ocaIity locality аз as holotype, V У.1989 (Ozerov) (Оаегоу) (ZMUM). (ZМЕ1М). ГЕСШРТЮМ. DESCRIPTION. МАШЕ. MALE. Егоп5 Frons black, шаё; matt; orbital plates негу very broad, Ьгоад, black Ь1аск and апд shining. shiпiпg. Face Ь1аск, black, glossy,

2 146 A.I. Shatalkin 5hatalkin 7 Figs 1-7. Маме Male terтiпalia terminalia о of Lauxaniidae: 11 СЫборозк Ca2liopum саисавгсит caucasicum вр.п. sp.n. (veпtral (ventral view); 2 C. С. sakhannicum sakhaliпicum sp.n. вр.п. (ventral view); 3-4 Мгпенгга Minettia аивнггаса austriaca Неппгу Hennig (dorsal and апд lateral view); 5 Minettiella elueтgi elhergi вр.п. sp.n. (1аСега1 (lateral view); 6-7 РасЪусеriпя Pachycerina пгпае nine sp.n. (ventral and lateral view). The The following followiпg abbreviations are ате used: ивед: a а aedeagus; аес'еаыз; c с cerci; сетсг; da да aedeagal аедеауа1 dorsal дотга1 sclerite; sderite; е a ерапдггит; epandrium; g --- уопород; gonopod; h ---- нурапдтгит; hypandrium; 1д ld lobes оf of аедеуа1 aedegal dorsal sclerite; р p -- рагатеге; paramere; ри pv ventral ептаг arm агт of paramere; рагатеге; s surstyle. sиrstyle. Рис. PlIC Гениталии Fei Tssnsl nuaor видов Lauxaniidae: 1 Са?ioprtm Calliopum саисасгсит caucasicum sp.n. (au,4 (вид спереди); cnepeau); 2 С. C. sakhaliпicuт sakhalfnicum вр.п. sp.n. (вид спереди); cnepeau); 3-4 Minertia Minettia аигнггаса austriaca Hennig (ska (вид сзади C32,441 PI и сбоку); c6oky); 5 ЛдгпеннiеТ1а Minettiella elbergi elhergi sp.n. (mu (вид c6oky); сбоку); Pachycerina пгпце nit-lae sp.n. (au,s, (вид спереди cnepeak ц u сбоку). c6oky). На Ha этих 3T14X и H следующих citeapouptx pucynttax рисунках 41C110A1331TIOTC11 используются следующие cneapouple сокращения: coxpatuenua: а a эдеатус; apeazyc; с c церки; r4epkss; да da 4,0pCaAbHblii дорсальный склерит c.k.aepwr эдеагуса;,s,earyca; е e эпандрий; anallapnci; g rouormabi; roноподы; h rnnallapl-ei; гипандрий; 1д ld лопасти nonacm Aopcaasuoro дорсального exaepwra склерита aaearyca; эдеагуса; р p napaaiepa; старамера; р~ pv llelitpaabfla14 вентральнаи рука pyka парамеры; napamema; в s сурстили. cypervux. тодеганеlу moderately сопуех; convex; parafacials дагк-ьгошп, dark-brown, оа matt.. Antennae with Ьава1 basal segments segmeпts yellow; third thiгd antennal апнеппа1 segment segmeпt times as long 1опу as penultimate. Halteres На!юегез yellow. Abdomen АЬдотеп Апнеп- middle of distal discal се!!. cell. Ultimate section sectioп of МMi., 1+2 about аьоин 1.8 пае дагк dark browп brown with brownish уе1iош yellow base, Ьаве, it about а1юин 2.7 times Ыасk, black, slightly shining. Genitalia (Fig.1) as as in in C. С. аепеит aeneum as 1опу long ад as its width. Arista yellowish yellawish brown,!юуп, with clear с1еат HI. F11. but surstyli suгstyli 1опуег longer (their Ierьgth length equal Ёо to 2/3 and апд haъzs: hairs: width of its feathering featheгiпg а a little more тоге than thaп thickness thickrless озоге more of рагашеге) paramere) апд and parameres рагатееев син cut apieally. apically. of its ЬадаГ basal рагн, part. Height Heigltt of оf frons Егопв about аьоин 1.2 times less than Chaetotaxy, (haetotaxy_ 1 h, h, 1 р1, ph, 0+3 dc, 1с, ac ас in 6 го rows,, 1 рр1, ppl, its width. Gena Оеоа аьоин about 4.1 times tirпes less than 1 ал height of оf eye. еус. 1 твр1, mspl, 2 stp1. stpl. Еасга1 Facial index (ratio of of width of of face fасе но to parafacial paгafacial аё at barder border 1Зоду Body 1eпgth length 3.8 mm. она. ое of Еоуеа) fovea) аьоин about 3.9. Моиннрагн5 Mouthparts and апд palpi ра1рг black. Thorax ЕЕМАL.Е FEMALE гевет61ев resembles таiе. male. Воду Body length Эength mm. топ'. Ьiack, black, shining. shiniлg. with greenish reflection. геf1еснгоп. Fore Роге 1е$s legs black 1 DIAGNOSIS. 1AG1OSIS. C. С. саисавгсит caucasicum вр.п. sp.n. is very тегу similar simllar но to with yellow клеев. knees. МЫ Mid and аж! hind нгпд legs yellow; femora fегпога С. C. aeneum аепеит апд and differs апеегз Егого from it by Ьу the structure stгucture of оf male гпа1е Ыаск, black, yellow apicaliy. apically. Ргеаргса1 Preapical bristles present ап on all ан1 genitalia. geпitalia. The paramere рагатеге is лон not сё cut аргса11у apically and the sursyli tibiae. Нгпд Hind tibia with dense депве apical аргса1 brush Ьrush of black Ыаск spinules ате are 1/2 and апд less of оf the parameres рагалетез гп in С. C. аепеит. aeneum. Тне The last ventrally; тгд mid metatarsi with a а similar brush Ьrush anteroven- апнегаиеп- species оссптз occurs in the залзе same locality аз as С. C. саисавгсит caucasicum but trally. Wings Wiпgs yellowish уе!1од1зь with yellow veins. 4а to Ьеуопд beyond ан at least a а month топни 1анег. later.

3 Ые New ' апд and 1ittic little known knowп Lauxaniidae & Strongylophthalmyiidae Strongylophthalr пyiidae flies 147 Calliopum sakhalinicum saklгalinicum Shatalkin sp.n. Fig матеа1ае MATERIAL. Но1онуре:д, Holotype:S, 5akhalin, Sakhalin, У Yuihno-Sakhalinsk,"Chek- ЬпоаИiаГiп I. hova" Peak, Реак, 27Л' VII.1971 (Nartshuk). (~Iartshuk). Paratypes: 1 д`, 1 1 9, каше same 1ocality locality as holotype, 27.VII V (Nartshuk) (Martshuk) (ZLSP). (ZiSP). DESCRIPTION. DESCRiPTION. MALE. МАШЕ. Frons Егопв Ыасk, black, shining, shiпiпg, аьоие above алнеппае antennae ус11о yellow. '. Face Еасс Ыаск, black, glossy, glassy, moderately modcrately сопиех; convex; parafacials parafacia[в дагк-ьгошп. dark-brown. Апнеппае Antennae with basal segments ешеп s уе11ош; yellow; third thiгd antenna! anнеппа1 segment дагк dark brown with brownish уе11ош yellow Ъае, base, аьоин about 2.2 times as long 1опу as wide. wiдe. Aтista Arista yellowish brown, with с1еат clear haiгs: hairs: width widtь о of its featheтing feathering епа1 equal но to thickпess thickness о of its basai basal раrt. part. Height ое of fгопв frons аьоп about 1.3 times less ннал than its width. Gena Осла about аьоп 3.б 3.6 times less than height of ое еус. eye. Facial гпдех index (гatio (ratio of ое width of face fасе to parafacial рагаfасга1 ан at Ьогдет border ое of (оиеа) fovea) аьоин about 3.2. Mouthpaтts Mouthparts апд and palpi black. Ыаск. Thorax Тногах black, Ыаск, shining. shiпiпg. Legs 1е black. Ыаск. Ргеаргса1 Preapical bгistles bristles оп on а11 all tibiae. Мгдд1е Middle anд and нгпд hind tarsi yellowish. yeliowish. Wiлgs Wings grayish. ta beyond Ьеуопд middle тгдд1е of ое discal се11. cell. Ultimate section оf of М[+2 M, about аьоин 2 times as long ioлg as penultimate. На1Сегсв Halteres yellow. уе11о. Abdomen АЬдотеп Ыаск, black, slightly shining. Genitalia Geпitalia as гп in 1Fig.2. Ё8.2. Снаенанаху. Chaetotaxy. 1 Ь, h, 1 рн, ph, 0+3 1с, dc, ас ac in гп 4 rows, 1 рр1, ppl, 1 mspl, mspi, 2 stpl. Воду Body leпgth length 3.3 mm. тт. FEMALE геветыев resembles та1е. male. Воду Body lcngth length 3.4 mm. тт. DIAGNOSIS. )IAGNOSIS. Маев Males of the new лсш species are ате easily еавг1у дistinguished distinguished from those 11о5е ое of known клошп European Еигореап spccies species Ьу by the genital 8сгйа1 structure. гысёоге. Genus Minettia Robineau-Desvoidy, Rnbineau-Desvordy, Minettia austriaca аивнггаса Неппгу, Hennig, 1951 Figs MATERIAL. CJ',, Sakhalin, Yuzhno-Sakhalinsk, Чuzhno-SakhaIinsk, "Chekhova" "СЬе1(Ьо'а' Реак, Peak, 27.V[I ,V (ЫагЬц1() (Nartshuk) (ZISP). DIAGNOSIS. DIAGNOSI5_ This species resembles resemыes repгesentatives representatives ое of 5с1гипшпгггтуга Schunzannintyia Рарр Papp in ехнетпа1 external characters. It It can сап be Ьс distinguished distгnguished from dark 1аг1 species of Minettia Miпettia on оп the basis of the structure госёоге of ое the male та1е postabdomen. ровнаьдотеп. Genitalia Geпitalia (Figs.3,4) with surstyli suтstyli pointed apically; аргса11у; gonopods ате are ртс$е1т present (?); parameres рагатегев are аге symmetrical with developed деие1оред (parameral) (рагатсга1) агтв arms articulatiпg articulating with нурапдггипl hypandrium ventral- ly. Till Тг11 now пои' M.austriaca was 'а 1known лочп fтот from Europe. 1у. Еигоре. Сепив Genus Minettiella, Ма11асн Malloch 1929 Маllосн Malloch [1929] considered сопвгдегсд Minettiella to be Ье a а subgenus suьgenus of the genus ело Minettia Miпettia R.-D. Кеу Key но to the species. 1(2). Width of feathering of of arista aгista noticeably понгсеаь1у more тоге than thickness thickпess of its basal Ьава1 part. 3rd 3гд antenna' antennal segment shorter. shortcг. One Опе pair of large 1аге acгostichal acrostichal bristles bгistles in ЕгоггЁ front of prescutellars. ргевсине11агв. Geпitalia Genitalia Fig.5. й mm. тт. Primorskiy Primoгskiy Кгау Kray М. M. elbergi sp.n. 2(1). Arista equal еgиа1 to or ог less than thaп thickness thickпess of its basal part. 3(4). 1+3 dc. ас. Mid Мгд and апд hind нглд tibiae yellow, уе11о, blackish Ыackish only оп1у apically. 3.5 тт. mm. Еогтова, Formosa, Victnaт, Vietnam, 5umatra Sumatra М. M. atratula atratцla (де (de Meijere) Мегjеке) 4(3). 0+2(3) dc. ас. 5(6). 0+2 dc. ас. Mid and апд hind hiпd tibiae black, Ы аск, yellowish basally. 3.0 тт. mm. Victлam, Vietnam, Java ]ада М. M. atrata атаеа (де (de Meijere) Мегjеке) 6(5). 0+3 dc. ас. Mid Мгд and hind hiпd tibiae yellow, уе11ош, blackish Ьiackish only anly аргса11у. apically тт. mm. Primorskiy Priгnorskiy Kray Кгау М. M. coracina Shatal 5hatalkiп kin Minettiella elbergi Shatalkin п sp.n. Fig МАТЕА1АL. MATERIAL. Но1онуре: Holotype: д', Cr, Рптогвкгу Primorskiy Kray, Кгау, Khasan Кнавап дistr, "Kedrovaya "Кедгоvауа Радд" Pacld"' 1еегуе, Reserve, 17.]Х /1980 (Shatalkin). Paratypes: Раганурев: 2 99, same locality а5 as holotype, IХ.1980 (Shatalkin); (5hatalkiп); 9 y same 1ocality, locality, 15X1968 Х1968 ((Gorodkov); Согодкаи); 2 cpc1r, стс', Primorskiy гskiy Kray, Кгау, Karnenushica (30 (3О km SE Е frот from Ussuriysk), 10 and 13.VIII VI]Т.1987 (Shatalkin); 2:.W 299 same locality, Iоса1гну, 30.V11 апд and 15.VI1I VIII.1984 (Shatallпn) (Shatalkin) Каmenushka (ZMUM). (ZМим). DESCRIPTION. MALE. Неад Head Ыаск. black. Frons гол in апнеrioт part рагн brownish Ьгownish yellow ycllow апд and spatted spotted with brown browп in гп anterior front froпt of ое осе11аг oceliar tтiaпgle. triangle. Face Еасе Ыасk, black, slightly grey теу dusted. Parafacials from Ьгошп brown to brownish bгownish yellow. ycllow. Antennae Апнеппае yellowish browп; brown; third апнепла1 antenna] segment сшеп аьоин about 1.7 times as 1опу long аз as its width. Arista yellowish brown, width ог of its feathering slightly less Снап than half wiдth width of third thirд апнеппа1 antenna! segment. segmeлt. Height о! of frоns frons ргаснгса11у practically еgцаi equal to its width. Осла Gena аьоин about 8.6 times less than Жап height о! of еуе. eye. Facial гпдех index (ratio of width of of face fасе to parafacial ратаеасга1 at а1 border baгder of Еоуеа) fovea) аьоин about 5.8. Mouthparts and апд palpi black. Thorax Тнотах Ыаск, black, shining. shiпiпg. Legs 1ез Ыас1, black, tibiae ю in basal 1/5, middle тгдд1е anд and hinд hind tarsi yellowish. уе огiз1i. Ргеаргса1 Preapical Ьгistics bristles оп on а11 all tibiae. Wings grayish. ta ta beyond Ьеуопд тгдд1е middle of discal ее11. cell. Ultimate веснгоп section of МM 1+2 about аьоо 1.8 tiгпes times as 1оп8 long аз as penultimate. На1нетсв Halteres уе11ош. yellow. АЬдотеп Abdomen Ыаск, black, slightly shining. shiпiпg. Genitalia Сепгна1га (Fig. 5) strongly внтопу1у sclerotized; вс1егонгхед; epandrium epandriuт cloпgate; elongate; зогё surstyli beak-like pointed ventrally; veпtrally; gonopods латгохл narrow,, сигиед curved уii догва11у. dorsally. СЬаенонаху. Chaetotaxy. 1 h, 1 рн, ph, 0+31с, dc, ac ас in гп 4 rows, posterior postcrioг ac ас of middle rows long, 1опу, sligьtly slightly less ннап than prescutellars, 1 зьогг short га, ia, 1 рр1, ppl, 1 mspl, 1 1 stp1. stpl. Воду Body leпgth length mm. тт. ЕЕМАLЕ FEMALE resembles Ьles та]е. male. Воду Body length О-3.6 mm. тт. DIAGNOSIS. М. M. elbergi sp.n. is the second весопд representativseпtative of the genus еппз гп in the Palaearctic region. гegion. ТЬс The пе'у new species аи!есгз differs fтотм. from M. coracina 111 in being Ьегпу вта11ет, smaller, its тгд mid герге- апд and hiпd hind tibiae being beiпg Ыаск, black, hairs haiгs of arista more тоге ехрапдедed, апд and its genitalia geпitalia different. аиееегеп. The Тне new пеw species is very егу ЬГгс like expand- М. M. atrata. атаа. 1 I compared сотрагед М. M. elbergi with Vietnamese Vietпamese specimens specimeпs determined Ьу by те me as М. M. аёгаа. atrata. The пеш new species is дедгсанед dedicated to the Estonian dipterologist К. K. Elberg. ЕiЬету _ Genus Pachycerina Маспаг Macquart,, 1835 Тне The geпus genus was ''аз known кпошп hitherto from the three Натее Ра1аеакенгс Palaearctic species. One Опе more глоге species spccies is described as пеш new Ьс1ог. below. Pachycerina ninae Shatalkin Slzatalkin sp.n. Figs MATERIAL MATERIАL. Но1онуре: Holotype: с Cr, ', Ки Kurile -11е Is., Kunashir, "Stoibchatyy" "Stolbchatyy" Cape, Саре, 9.У11Л985 9.VII.1985 (Gzurkin). (Czurkin), Paratypes: Рагареа 9, same 1осаРгну locality аз as но1оtype, Tretijakovo, Tгetijakova, 10.V VII.1985 (Сг1ап) (Czurkin); 9, y, same loralitv locality as halo- Ьо1огуре, holotype, Tjatja Volcano, Vnlcano, 4.V VII.1976 (Sinitshenkova) пkova) (Z.МЕ.[М). (ZMUM). DESCRIPTION. МАLЕ. MALE. Неад Head yellnw. yellow. Ггопз Frons with Ыаск black оиа1 oval spot superposited supeгposited оп on оее11аг ocellar triangle; anterior тагугп margin ое of this spot closed снозса before ЬеЕоге one опе cames comes to level 1ее1 of hind hiпd от, or, апд and its height about аьоин 1.4 times less than Ьал анапсе distance from spot spot но to base Ьаве о! of antennae. апнелпае. Black В1аск spots зроз iin п place р1асе of ое iпsertioп insertion ое of ог or аьзеп absent.. Face Еасе with a а pair раек of о! Ыаск black зро.з spots оп on еасн each side of lofty facial lofty facial knob. кпоь. Antennae Апнсппае уе11ош, yellow, with ЁЬ Ьава1 basal зешсёз segments Ьrowпish; brownish; third апнсппа1 antenna' segment эешел аьоин about

4 148 А.I. A.I. Shatalkin 4.2 times tiтes as а$ long 1оа as wide, rounded гоипдед apically. apicai у. Arista Aгista incrassate, псгазас, with депве dense Ыаск black hairs, аьоин about 1.2 times tiтes as long as third antennal зе8шегх segment.. Height -ieight ое of frons аьоин about 1.8 times less Iess than its its width. Gena Сепа about аьоин 2.2 times less than height ое of eye. еуе. Facial гпдех index (танго (ratio of width ое of face Еасе 1о to рагаf parafacial асга1 а at border Ьогдег ое of fovea) Еоуеа) about аьоин 5.2. Mouthparts brownish, browпish, palpi yellow, гп in apical аргса1 half hа1е black. Ы аск. Mesonoturntum, metanotum, тенапонит, апд and sternopleuroп sternopleuron Ь1аск, black, grey геу dusted; humeri, notopleuron, понор1еигоп, р1еигае pleurae гп in upper иррег part рати yellowish Мевопо- grey. геу. Legs с1аг1 dark Ьго%Чп, brown, knees slightly yellowish. cal bristles оп on а11 all нгьгае. tibiae. Wings slightly brownish, browпish, with Preapiса1 соза1 costal rпaгgin margin с1оидсд clouded апд and на ta anд and tp нр bordered. Ьотдегед. Basal Вава1 section Ы of M1.2, Мк+х, restricting кest гictiпg дiscal discal се11 cell ti11 till 1а ta about аьаин 1.4 times as long 1опу as аз failpwing following section. sectiaп. Ultimate section of of 1vI М12 +2 about L6 1.6 times as as long 1опу as penultimate. репиlнгтане. Halteres уе11ош. yellow. Abdomen АЬдотсп Ыаск, black, shining, slightly угеу grey ёоз1е dusted.. Geniialia Genitalia (Figs.6,7) (Рз. stronglysclerotized; sclerotized; parameres рагатекевеgиа1 equal 1to о height of epandrium; left left paramere рагатеге wiih with wide tened prominence ртотiпеасе lateroapically and with small sma11 black Ь1аск flatнепед tooth veventroapically. пtroapically. Chaetotaxy. 2 ог or with anterior anнеггог of them inchoate, iпclinate, 1 h, 1 I ph, 0+3 dc, 1с ac ас in 4 rows гows with internal гпнегпа1 setulae вени1ае regular regиlar апд and 1олуег longer than those of ехнегпа1 external rows, 1 рр1, ppl, 1 твр1, mspl, 2 stpl. stp1. Воду Body leпgth length 3.2 mm. тт. ЕЕМАLЕ FEMALE геветы resembles сs таiе. male. ЕепЬ Length mm. тт. DIAGN05I5. DIAGNOSIS. Ме New species is is similar но to Р. P. seticarпis seticornis Е11. F11. ТЬе The latter дiffers differs Ьу by yellow coloration ое of the body Ьоду гп in general; уелега1; its mesonotum with stripes; тipes; the central сепнга1 frontal fraпtal spot зроё is larger and апд spots in place р1асе of ое insertion of ог or аге are clear. с1еаг. Parameres Рагатегев аге are weakly чеа 1у sclerotized sclerotizeд апд and аге arc less than the height of аf epandriurn. ерапдкгит. Geпus Genus Sapromyza Еа11еп, Fallen, 1820 Sapromyza тight might Ье be regarded as as a а blanket Ы апкен genus пus containing containiпg very дifferent different forms. fогтв. It includes iпcludes more тоге thaп than 80 Palaearctic species, 8 of them 1Ьеш beiпg being described Ье1ош. below. I ргорове propose пеш new names пашез for the following fallowing two 1 ю species: Sapromyza 5арготуха citriпella citrinella Shatalkin п is new пеш name паше for Ест S. citrina citriпa Shatalkin Shatalkiп (1993, Russiaп Russian Епнºгпо1.J.: Entomol.J.: 112) nec пес Meigen (1826, Syst. Beschr., сзсьг., 5:264). Туре Type 1оса1гну: locality: Еат Far ЕазЁ, East, Ussuriysk, Kamenushka. КашеппзЫа. $арготу2а Sapromyza remmae Shatalkin 5hatalkiп is new пеш name паше for 5.тгсгорууа S.micropyga Е. E. Remm Аетт е1 et Elberg (1980,[Insects [Тпзесз ое of Mangolia], Mongolia], 7: 426) nec пес Malloch Ioch (1933, Diptera Dipteгa ое of Patagonia апд and $опгй South Chi1e, Chile, 6(4): МаI- 377). Type Туре locality: Mongolia, Govialtai, Jargalant. пt. Sapromyza remmae геттае is named патед after г Dr.Mrs. Дг.Мгв. Evi Remm Аетт (ТагЁц, (Tartu, Estonia). Sapmmgza Sapromyza cerata Shatalkin зр.л. sp.n. Fig МАТЕАГАI.. MATERIAL. Но1онуре: Holotype: д', d, Azerbaydzhan, КЪас} Khachin п River, VI.1885 (Г 5 (Khavraiskiy). Раауре Paratypes: : 1 I o, д', 1 Y, 9, азше same locality as но1онуре. holotype, VI.1885 VI.1&85 (Khavraiskiy) (ZISP). DESCRIPTION. МАLЕ. MALE. Неад Head yellow. Осе11аг Ocellar нггапу1gle yellow. Апнеппае Antennae yellow; third апнсппа1 antennal segment segmeпt about 1.6 times as long 1оп8 as аз wide, rounded гоипдед ventroapically, veпtroapiea[ly, with slight dorsal двгва1 angle апу1с and апд deep дсер black Ы аек apical аргса1 half. ha1f. Arista yellow, in microscopic hairs. Height of of frons пs about 1.2 times less lcss than its width. Gena Сепа high, about аьоин 2.5 times tiтes less than height beight of of еуе. eye. Facial гпдех index (ratio of ое width of face fасе но to parafacial ан at border Ьотдег of fovea) Еоуса) about аьоин 4.4. Mouthparts 6гошпгвн, brownish, palpi уе11ош, yellow, slightly brownish browпish apically. ТЬстах Thorax trian- уе11ош. yellow. 1Legs е8з уе11ош. yellow. Preapicai Preapical Ьristles bristles оп on а11 all tibiae. Wings Wirдgs yellowish. ta slightly sligьtly before Ьеfоге тидд1е middle of ое дгвса1 discal се11. cell. Ultimate IJltiтate веснгоп section of ое М Mi.2 about 1.5 tin times зes as 1long впу as аз penultimate. репп1нгтане. На1 Halteres егез yellow. уе11ош. Abdomen АЬдонпеп уе11ош. yellow. Geлitalia Genitalia - Fig.B: Fig.8: рагатветез parameres asymтetrical; asymmetrical; right ое of ньет them longer loпger than aedea- аедеаgus; gonopods gnпopods аьзеп1; absent; surstyli pointed рогпнед dorsally. догва11у. Chaetotaxy. 1 h, h, 11 ph, 0+3 de, 4с, ac ас in гп 4 rows, 1 рр1, ppl, 1 тврi, mspl, 2 stp1. stpl. Воду Body length 3.5 mm. шт. ЕЕМАLЕ FEMALE resemыes resembles maie. male. Apical darkening daкkeпing of раlрг palpi practically not по expressed. ехргеввед. Body Воду length 3.9 mm. тт. DIAGNOSIS. Тhе The та1е male ое of the new пе species вресгев is very 'егу siтilar similar to ta those of of S. pistaciphila group топр in ю genital eпita1 structuгe structure in which gonopods are аге lacking and surstyli are аге pointed пted dorsally. догваl1 у. Representatives ое of the S. pistaciphila group гоор differ аиееег from fгот S. cerata сетана sp.n.in that ЁЬаЁ the frons has Ьаз Ь1ас1 black spots in гп the place р1асе of the insertion inseriion of ое or. от. Parameres of S. pistaciphila аге are вуттенхгса1 symmetrical апд and longer 1опует than ннап aede- аедеagus. In the coloration coloratiaп ое of Ьаду body S. cerata аугеев agrees also with those Ьозе specimens speciтens of ое S. ravida гаиiда sp.n., вр.п., which ЫсЬ are аге character- снагаснетized Ьу by the absence аьзепсе of the frontal froпtal black Ь1аск зро spot (as in гп S. kabuli kabиli Рарр). Papp). The ТЬе shape of the surstyli, suгstyli, hypandrial Ьурапдтга1 pгocesses, processes, апд and аедеауив aedeagus in in both Ьонн species вресгев are аге different. пt. Palpi Pa1pi of S. cerata сетана with weak чеа1 аргса1 apical дarkening, darkening, ly пон not expressed in гп fеша1е. female. In this Ыз respect the the new пек species is similar to S. remmae геттае п.п.(5. micropyga 1Remm ешш е1 et practical- Ту ЕIЬету) Elberg) fгот from Mongolia. Мопуо1га. Тне The 1аннег latter species is distin- distiпguished externally Ьу by the darkened дагкепед tarsi. Sapromyza ferganica Shatalkin sp.n. р.тж. Fig МАТЕАЕА MATERIAL.. Ноlонуре: Holotype: О, Tadahilпstan, Tadzhikistan, Perganskiy Khr., Кнг., иррег upper reaches гeaches of the Jassy. f 2000 m, т, 20.VI УТ.1 (Zhelochovtsev) (Zhelосhavtsev) (ZMUM). (2М(IМ). DESCRIPTION. MALE. МАШЕ. Head Неад yellow. Ocellar Осе11аг gle g1e yellow. Antennae Апнеппае yellow; third thiкд antennal апнеппа1 segment segmeпt trian- about 1.8 times as long 1опу as аз wide, "юае, rounded гоипдед ventroapically, иелнтоаргса1lу, with slight dorsal дотва1 angle; апу1е; deep деер black Ыаск in apical half. haif. Arista Aгista уе11ош, yellow, in microscopic тicгoscopic hairs. iiairs. Height of ое frons about аьоин 1.4 times less Iess than thaп its width. Gena Сепа high, hгgh, about 2.3 times less Iess than height of ое eye. еуе. Facial Еасга1 гпдех index (ratio of ое width of face fасе но to рагаfасга1 parafacial а at border Ьогдет of fovea) Еахеа) about аьоин 2.0. Mouthparts Моиннрагнв brownish, browпish, palpi yellow, in apical аргса1 half black. Ыаск. Thorax Тногах уе 1в yellow,, mesonotum тевопонит greyish уе11ош. yellow. Legs уе11ош. yellow. Preapical bristle Ьristle on оп hind Ыпй tibia absent. аьвепн. Wings yellowish. ta Ргеаргса1 slight у slightly before befoгe middle ое of дгвса1 discal cell. се11. Ultimate Ultiтate section sectioп of М Mi., +х about аьоин 1.5 times as long 1оп as penultimate. репи1нгтане. Halteres Haltеres уеi1ош. yellow. Abdomen АЬдотсп уеl1ош. yellow. Geпitalia Genitalia - Fig.9: right рагатете paramere bathed bilobed with ventral lobe 1оЬе straight and апд pointed рогпнед apically; аргса11у; ventral I0 lobe Ьс of ое left ПсЕ paramere рагатеге absent; аьвепн; gonopods уопородв absent. аьвелн. Chaetotaxy. 1 h, 1 рн, ph, 0+3 dc, ас, ac ас in iп 4 rows, 1 ppl, рр1, 1 mspl, твр1, 2 stp1. stpl. Воду Body lcngtь length 4.3 mm. шш. ЕЕМАLЕ FEMALE unknowп. unknown. DIAGNOSIS. Тне The рагатегев parameres of S. ferganica (ergaнica вр.п. sp.n. зьо'ч show the bilobated condition. сопаиюоп. In 1п this respect гезресё new пеш species belongs to S. simplicipes group гоор in гп which ЬюсЬ it is closely с1овс1у ге1анед related to S. hissarica вр.п. sp.n. and апд S. simplicipes Сгегпу. Czerny. It differs аюеесгз from Егото 6oth both in that 1Ьа St it has mesonotum mesonotuт yellowish vellowгsь угеу grey апд and mainly Ьу by its genitalia. InS. hissarica the дorsal dorsal 1оЬе lobe of ое the right paramere рагатеге is abseпt absent and апд its ventral иепнта1 lobe 1оЬе е1опуанед elongated with apical hook-like Ьепд. bend. On Оп the other hand напд S. sivпplicipes simplicipes Ьав has the ventral lobe 1оЬе on оп the left paramere. рагатеке. Sapromyza hissarica Shatalkin 5hatalkiп вр.п. sp.n. Fig

5 New е'т апд and 1little Ые known kпawn Lauxaniidae Lauxaпiidae & Strongylophthalmylidae Strongylaphthalmyiidae flies flies 149 в 1о 1г 11 right р left р p 14 Figs Ivlale Male terminalia termiпalia ог of Sapromy.zet Sapromyza: 8 5. S. cerata вр.п. sp.n. (ventral (иепнга1 view); 9 5. S. ferganica sp.n. (ventral (иепнга1 view); S. hissarica bissarica вр.п.(ventral sp.n.(ventral view); S. persica вр.п.(иепнта1 sp.n.(ventral view); 12 S. ргсеа picea вр.п.(lateral sp.n.(lateral view); S. ravida raviдa вр.п.(veпtral sp.n.(ventral аж' and 1ateral lateral view); S. 5. zirninae ziminae sp.n.(veпtral sp.n.(ventral view). бее See Pigs Figs 1-7 мот for abbreviations. аььтеигангопв. Рис. Psw `eincradimpi Гениталии splaoa видов Saprornyza 5артоту 8 S. S. cerata врл. sp.n. (ana (вид criepean); спереди); 9 S. S. ferganica fergarцca sp.n. р.п. OKA (вид спереди); criepean); S. нiвтатгса hissarica вр.п_(вид sp.n.(m4a спереди); cnepeysx); 11 &persica 5.persica sp.n.(m4a spn.(вид спереди); crlepean); 12 S. picea sp.n.(вид sp.n.(ssla сбоку); thoicy); S. гай1а ravida sp.n.(m4a sp.n.(вид спереди cnepean и p сбоку); c6oxy); 15 S. S. zi ziminae тirгae sp.rx.(3t4a вр.п-(вид спереди). criepea14). Для 402 расшифровки pactun4posx.14 сокращений copcpaujelinii смотри cmorpvi рис. plc МА MATERIAL 'RIAL Но1онуре: HoIotypc д d',, Tadzhikistan, Tadxhikistaп, Hissar Hissaz MtRange, Mtit Kwak m, т, 11.VIII.1969 (Keleinikova). Ратанурев: Paratypes: С d, 4 'п, вате same 1оса1гну locality а as Iioiotype, holotype, 6-14.VIII VI1I.1969 (Keleinikova)(ZivIUM). (.м11м). DESCRIPTION. МАI.Е. MALE. Неад Head уе11аш. yellow. Осе11аг Ocellar нггапу1angle уе11аш. yellow. Antennae Апнеппае уепо yellow; ; third Ытх antennal апнеппа1 segment tri- segmeпt about 1.9 times as а51опу long аз as wide, гоипдед rounded ventroapically, with slight зьь1 dorsal догва1 angle; апу1е; deep деер black Ь1аск in apical half. haif. Arista уе11ош, yellow, in гп microscopic гnicroscopic hairs. Height of Ы fгопв frons аьоин about 1.2 times less езз thaп than its width. Gena Степа high, about аьоин 2.2 times less 1ез than thaп height of о( eye. еуе. Facial 1~асга1 index гпдех (ratio (танго of Ы width of Ы face асе

6 150 А.I. A.I. Shatalkin to parafacia1 parafacial а at bnrdeг border of fovea) fочеа) аьоин about 2.2. Moиthparts Mouthparts 6rownish, brownish, ра1рг palpi уе1lош, yellow, in apical аргса1 half black. Ыасk. Thorax Тногах уе11ош. yellow. Legs уе11ош. yellow. Preapical bristle оп on hinд hind tibia absent. Wings yellowish. ta ta beyond Ьеуопд middle of ое discal сенl. cell. Ultiтate Ultimate section есноп ое of Mi+, М about аьоин 1.2 times as 1опу long as а репи1нгтане. penultimate. Halteres Halteтes yellow. уе1iош. Abdomen АЬдотеп yeilow. yellow. Geпita- Genitalia (Fig.10) (й8. with right гight ventral епёга1 parameral рататега1 lobe Iobe elongate eloпgate 1ia апд and hook-like сигчед curved apically; left ventral епта1 апд and right dorsal darsal 1оЬев lobes of ое parameres рагатегев absent; gonopods уопородв abseпt. absent. Chaetotaxy. 1 h, 1 ph, рь, 0+3 ас, de, ас ac [п in 4 rows, 1 рр1, ppl, 1 rasp!, msp1, 2 stpl. Воду Body length 3.9 mm. тпх. FEMALE resemьles resembles та1е. male. Body Воду 1епун1х length mm. тт. DIАGNOSIS. DIAGNOSIS. S. hissarica sp.n. is very similar to S. ferganica вр.п. sp.n. but Ьп diffeгs differs in гп having the yellow turn tum and peculiar реси1гат genital geпital structure. stгucture. Specifically, SpeciEically, the mesono- second весопд species нав has both dorsal рагатега1 parameral 1оЬе lobes; ; its right ventral ventra1 1оЬе lobe siгaight straight and апд pointed рогпнед apically. аргса11у. Зартвтуzа Sapromyza intoгrsina intonsina Уагот, Yarom, 1990 МАТЕА MATERIAL. Е АL. 2 2d'd (3' д', Turkmeniya, Chandyr River (triыutary (tributary а of Sumbar 5итЬаг River), 23.IV.1933 (Clshinskiy) (Ushinskiy) (ZISP); 3 д'д', o'd', 4?T, 9, Turkmeniya, Khr.Kopet-Dag, Khг.Kopet-I]ау, Aydere, Аудеге, 21 IV.-1.V У.-1.V.1981 (Ozerov) (Qzerov) (ZMLIM). DIAGNOSIS. This species was descгibed described оп on the тaterial material from Israel [Yarom, [Уагот, 1990] Taxonomically Тахопотгса11у S. intonsina iпtonsina Ье1оп belongs tos. intonsa угоир group differing mainly Ьу by genitalia. geпitalia. Зартоту2а Sapromyza persica Shatalkin sp.n. Fig. 11. МАТЕА1АI.. MATERIAL. Но1онурею Holotypee,, Iran, Егап, Asterabad АвнегаЬад (Gorgan) (Goтgan) 26.IV ТУ.1914 (Kiritshenko) (ZISP). (ZI5P), OESCRIPTION. DESCRIPTION. MALE. Неад Head уе11ош. yellow. Осе11аг Ocellar triangle уе11ош. yellow. АзЁеппае Antennae yellow; уе11ош; third antennal апнеппа1 segment ешепё апу1с аьоин about 1.8 times as 1опу long as wide. Arista Аrista ус11оч, yellow, [п in с1еаг clear hairs, width of ое its feathering featheriпg equal еgиа1 to но its thickened thickencd basal Ьава1 рагн. part. Height of ое frons fгопв about аьоиt times!езз less ннап than its wiдth. width. Оспа Gena abovt about 5.6 times Iess less than height of ое еуе. eye. Facial гпдех index (ratio of ое width of face Гасе to но parafacial para(acial at а barder border of fovea) about 4.2. Mouthparts Моиннрагнв brownish, browпish, palpi yellow, уе1lош, in apical half black. Ыаск. Thorax Тногах yellow. yellaw. Legs yellow. Preapical Ртеаргса1 bristles оп on а11 all tibiae. Wings yellowish. на ta slightly before тгдд1е middle of discal се11. cell. Ultimate section of ое М, M12 +х about 1.4 times аз as Гоп long аз as репи1нгтане. penultimate. Halteres На1негев yellow. yeliow. Abdomen АЬдотеп yellow. Geпitalia Genitalia (Fig.11) (й8. with trough-like tгough-like surstyli, сы cut аргса11у apically and with relatively long 1ов aedeagиs. aedeagus. Chaetotaxy. 1 h, 1 ph, рн, 0+4. dc дс (two ( о anterior anteгior of ое them theт shortcr), shorter), ас ac [п in 4 rows with setae зеае of median тедiап rows Iопуег longer than thaп Нюзе those ое of ехнегпа1 external гows, rows, 1 рр1, ppl, 1 твр1, mspl, 2 stp1. stpl. Воду Body 1евН length 4.0 mm. пхт. ЕЕМАЕ FEMALE uпknow unknown. п. DIAGNOSIS. Charactcrs Characters ое of thc the geпital genital apparatus арраганив indicate close close relationship relationship betwee between п this Ыз species and апд S. S. gazmanyi gozmanyi Рарр. Papp. S. persica persiёa has а a loпger longer aedeagus; аеаеапз; its surstyli,cut surstyli,.ut apically. аргса11у. The Тне new пс species is геадг1у readily separatcd separated Бу by the 4-rowcd 4-rowed ас ac (instcad (instead of ое the 6-rowed 6-гоwед ac ас in гп S. gozпгanyi). gozmanyi). ~аргвгпуга Sapromyza picea Shatalkiд Shatalkin sp.n. Fig МАТЕА1А1. MATERIAL. Н Holotype:d, аlонурегг, Kurile Киriiе Is., 1s., Kunashir, "Мепде1еуеиа" "Mendeleyeva" Volcano, 27.V[Г VII.1985 (Схигкгп). (Czurkin). Paratype: Рагануре: д, Cr, same locality as Бо[о holotype, у-ре, 27.УН VII.1985 (Czиrkin) (Czurkin) (ZMLIM). (2М11М). DESCRIPTION. MALE. МАLЕ. Head Неад Ыаск. black. Froпs Frans in anterior раг part above аьоие antennae апнеппае brownish Ьrownish yellow. уе11ош. Face Расе black, Ы аск, гваё matt.. Parafacials brown. Апнеппае Antennae Ъrown; brown; thiгд third апнеппа1 antennal segment зсглеп1 аьоин about 1.6 times аз as 1опу long as шгде, wide, browпish brownish yellow [п in 1оег lower рагн. part. Arista yellowish brown, practically bare. Ьаге. Height of гопв frons about 1.2 tiaпes times less than its width. Gena Сепа аьоцн about 6.3 times less than height of еуе. eye. Facial гпдех index (гatio (ratio ое of width of (асе face to рагаfасга1 parafacial а at Ьогдег border ое of fovea) about 4.5. Моиннрагнв Mouthparts апд and ра1рг palpi black. Ыаск. Thorax ТЬогах black, Ь1аск, brownish bтownish апза. dusted. Mesonotum Мевопонит shiпing. shining. Legs Ь1аск, black, тгдд1е middle апд and нгпд hind taгsi tarsi yellowish. Ргеаргса1 Preapical 6гistles bristles оп on а11 all tibiae. Wings grayish. ta slightly before тгдд1е middle ое of дгвса1 discal се11. cell. Ultimate section ое of М 2 about аьоин 1.6 times as as 1long опу as аз penultimate. репи1нгтане. Halteтes Halteres yellow_ yellow. АЬдотеп Abdomen Ыаск, black, slightly shining. Genitalia (Fig.12) with parameres рагатегев bilobate Ьг1оЬане apically; gonop- gonapods abseпt; absent; surstylus with thorn-like teeth. Chaetotaxy. 1 h, 1 Э рн, ph, 0+3 ас, dc, ас ac [п in 4 rows, 1 рр1, ppl, 1 mspl, 2 stpl. Воду Body leпgth length mm. тт. ЕЕМАLЕ FEMALE uпknowп. unknown. DIAGNOSIS. DIАGNOSIS. ТЫз This пет new species is чету very similar но to S hyaliдata hyalinata М Mg.. иг in ехнегпа1 external characters сьатасегз апд and сап can Бе be separat- ed ( from тот it it Ьу by the characteristic гistic ge genital пital str structure. ед цcture. Заротуа Sapromyza гагтгс1а ravida Shatalkin вр.п. sp.n. Figs МАТЕА1АI. MATERIAL Но1онуре_д, HolotypḶ-d, Iran, Iгав, Tabriz, ТаЬггх, 5.V.1914 (Andrievskiy). Рагагур Paratypes: 33 с'о', OV, 3?У, same locality as halotype, holotype, IV. апс1 and 5.V.1914 (Aпдrievskiy); (Andrievskiy); Агтепгуа, Armeniya, V (Khavraiskiy) (ZISP). (ZISP), DESCRIPTION. МАLЕ. MALE. Неад Head yellow. Ocellar Осе11аг triangle yellow. Paratype Рагануре from Armeniya Агтепгуа with brownisisн arrow-likc arrow-like froпtal frontal зро spot as in S. 5. kabuli Рарр. Papp. Апнеппае Antennae уе11ош; yellow; third апнсппа1 antennal segment зешеп about 1.8 times as 1опу long as wiдe, wide, with dorsal angle аз as ins. apicalis apiealгs Lw. апд and Ы black аек [п in apical аргса1 half. ha1f. Arista yellow, уе11оу, [п in clear с1еаг hairs, width ое of its feathering equal вgиа1 to its thickened thickeпed База! basal рат part.. Height of ое frons4about fгопвiаьоин 1.3 times tiтes Iess less than its width. Сепа Gena about 3.2 times less than height of eye. еуе. Facial ипаех index (гatio (ratio ое of width of face (асе to parafacial рагаfасга1 at а Ьогдег border of (ovea) fovea) аьоин about 1.5. Моиннрагнв Mouthparts Ьгошпгвн, brownish, ра1рг palpi уе11ош. yellow. Тhогах Thorax yellow, grey угеу dusted. дивнед. Legs yellow. уе1lош. Preapical bristles оп on а11 all tibiae. Wings yellowish. на ta slightly beyond Ьеуопд middle of discal се11. cell. Ultimate section sectioп of M52 М 2 about 1.2 (1.4 in in paratype рагануре from Armeniya) Аrmcniya) times as 1опу long аз as Peпultimate. penultimate. Halteres Halteтes yellow. Abdomen yellow. Geпitalia Genitalia as гп in Figs.13-14: ерапдггит epandrium with large surstyli pointed рогпнед apically; аргса11у; hypandrium нурапдггит bilobed ventrally veпtrally with asymmetгic asymmetric 1оЬеs lobes (gonopods?); right АЬдотеп рагатеге paramere wide; it goes оез гоцпд round of acdeagus aedeagus ventrally епта11у ала and settles scttles дошп down оп on its left 1еЕ1 side; зиае; left!е( paramere рагатете moved тоиед аотзан dorsally. I у. Chaetotaxy. 1 h, 1 рь, ph, 0+3 dc, 1с, ас ac [п in 4 rows, 1 рр1, ppl, 1 I mspl, твр1, 2 stpi. stpl. Воду Body Cengtlx length 3.1 mm. тт. FEMALE FEMRLE resemыes resembles таiе. male. Воду Body lcngth length mm. тт. DIAGNOSIS. Мсв' New species is very егу siгnilar similar tos. kabuli Рарр. Papp. Both species Ье1опу belong но to the угоир group includiпg including species which аге are дrstinguished distinguished Ьу by ргезевсе presence оенне of the unique fгопна1 frontal spot зро[ above аьоуе осе11аг ocellar triangle [Papp, Рарр, 1979]. S. simplicior siгnplicior Непде1 Hendel (=S. simplex Lw.) - the ([гз[ first of the described дescribed species within the угоир, group, has а a distinct нуре type ое of the genitalic gcnitalic structure, showing undeveloped ипдеие1оред anterioг anterior hypandriapaedrial 1oЬcs.S. lobes. ravida sp.n. апд ands. kabuli have siтilaг similar genitalia, diffeгing differing [п in the structure of ое anterioг anterior hypandri- allobcs lobes ( (gonopods?) уопородв?) (Figs (Figs 13-14) ). In the In the new пеш species they are аге asymтetrie asymmetric апд and without dorsal догва1 Ьепд; bend; in гп S. kabuli the hypaпdrial hypandrial lobes IaЬes are атс short Бог[ and symmetric, syтmctric, they атс are сиrvед curved аотн1у dorsally [в in (отт form of the hook; right

7 Ме New ' аж' and little knowп known Lauxaniidae 1..аихапггдае & Strongylophthalinyiidae Stroпgylophthalшyiiдae flies 151 раагпше paramere 0f of this species is паггош, narrow, left опе one is triaпgular triangular апд and shorter ннап than that of о1 S.ravida sp.n. S. romanooi romanovi вр.п., sp.n., described bellow Ье11вш on оп the female, етаlе, дiffcrs differs fгот from othcr other species Ы of the угоир group Ьу by угеу grey ео1оиг colour of thorax. Зартоту2а Sapromyza 7О7ПФ2ОЛ romanovi Shatalkin Shata1kin sp.n. МАТЕRIА1,. MATERIAL. Но1ануре: Holotype: 9,?, Tadzhikistan, Raгuit, Ramit, Kafirnigan Kaflrпigaп River, 26.VI VII.1939 (1 (Romanov) огапо ) (ZMUM). DESCRIPTION. FEMALE. Неад Head yellow. From Егоа divided in уе11ош yellow апнеггот anterior рагн part по not eachiпg reaching first! г 1 ог or аж' and red-brown red-browп posterior рагн. part. 1..аег Latter with ааг dark brown triaп- trianуи1ат gular тагк mark on оп anterior апнеггог margin. Апнеппае Antennae yellow; thiгd third апнеплаl antennal segтeп segment short, Ьог, about аьоин 1.4 times аз as 1опу long as wide, гоипдед rounded испнгоаргса11у, ventroapically, апд and with dorsal дorsal angle; ао 1с; deep деер Ь1аск black гп in apical half.! а1е. Arista yellow, yeliow, in microscopic hairs. haiгs. Height of froпs frons ab0ut about 1.3 times less than its width. Оспа Gena ЫЬ, high, аьопё about 4 times less thaп than height ое of еуе. eye. Faeial Facial index (ratio of ое width of ое face пасе to parafacial а at. border Ьогдег of fovea) Еоуеа) аъоо about 1.5. Mouthparts brownish, Ьго iзж, palpi yellow. Mesonoнит tum угеу, grey, humeri нитегг аж! and всине11ит scutellum yellow; р1еитае pleurae yeilowish grey. геу. Legs yellow. Ргеаргса1 Preapical bristle оп on а1э all tibiac. tibiae. Wings yellowish. yeliowish. ta slightly before middle ое of дгвса1 discal се11. cell. Ultimate sectioп section ОЕ of Mi., М1+ about аьаин 1.5 times as long as ав yellow- репи1нгтане. penultimate. Halteres На1 егез yellow. уе121ош. Abdomen Аbд0теп yellow. Chaetotaxy. 1 h, 1 рн, ph, 0+3 ас, dc, ас ac гп in 4 rows, тows, 1 рр1, ppl, 1 швр(, mspl, 2 stp1. stpl. Воду Body leпgth length 4.7 mm. тт. МА1..Е MALE unknown. DIAGNOSIS. The triangular тагк mark оп on froпs Irons апд and the grey гсу теволонит mesonotum аге are sиfficient sufficient но to distiпguish distinguish this species from гот а11 all other in гп the genus. geпus. The former fогтет снагаснег character shows зьо"з ЁЬа that S. romanovi sp.n. is a а member of S. simplicior (S. (5. вгтрlех) simplex) group groиp гп in which it is closely related ЁоЗ tos kabuli. Sapromyza Saprorпyza ziminae Shatalkin Shata1kiп sp.n. Fig MATERIAL. Ноlонуре:д, Holotype: 07, Nalchichevan, Nakhichevan, 27.У V.1957 (Zimiпa) (Zimina) (ZMUM). (ZМцМ). DESCRIPTION. МАШЕ. MALE. Неад Head yellow. Осе11аг Ocellar triangle уе1lош. yellow. Апнеппае Antennae уе11ош; yellow; third апнеппа1 antennal segment segmeпt апу1е аьоин about 1.8 times as as iolong пg as wide, ае, rounded тоипдед apically, аргса11у, without sharp дотва1 dorsal anу1е, angle, деер deep Ыаск black гп in apical аргса1 half. ha1f. Arista Aгisia brown, гп in microscopic hairs. Height Ы of froпs from аьоин about 1.1 times less lcss than its width. Gena Сепа about 4.6 times less than height Ы of еуе. eye. Facial гпдех index (гatio (ratio of width of ое face пасе to parafacial а at Ьогдег border of fovea) аиеа) аьоин about 3.5. Моиннрагнв Mouthparts brownish, ра1рг palpi уе1эош, yellow, гп in apical half black. Ь[аск. Thorax Тногах yellow. Legs yellow. Ргеаргса1 Preapical bristles оп on ан all tibiae. Н1пд Hind tibia with with dense депве apical аргса1 brush of о black Ыасk spinules ventrally; епха11у; тгд mid апд and hiпd hind metatarsi with a а similar brush Ьтивн anteroventrally. anteroveпtrally. Wiпgs Wings yellowish. на ta beyond Ьеуопд middle наа1е of discal discaэ се11. cell. Ultimate section sectioп of Mi+, i+z аьоин about 1.4 times as 1опу long аз as penultimate. Halteres На1негев yellow. Abdomen АЬдотеп уе11аш. yellow. Genitalia Geпitalia (Fig. 15) with elongate е1сйае epandrium; ерanдгiит; surstyli sиrstyli вriented oriented регрепдгси1аг1у perpendicularly гelative relative to ерапдггит; epandrium; рагатетев parameres аьоин about 2 times -less than height of epandrium, ераждтiит, right рагашеге paramere longer than left one. опе. Chaetotaxy. 1 h, 1 рн, ph, 0+3 дс, de, ас ac in 4 rows, гоз, 1 рр1, ppl, 1 швр1, mspl, 2 stpl. 1р1. Воду Body leпgth length 3.8 mm. тт. ЕЕМАLЕ FEMALE unknown. DIAGNOSIS. Тне The male таiе of S. ziminae zirпinae sp.n. is very similar to that Ьа of ое S. afghanica Рарр Papp (fгот (from Afghaпistan Afghanistan апд and Мгдд1е Middle Аsia) Asia) гп in having иеонга1 ventral brush оп on нгпд hind tibia, тгд mid апд and нгпд hind zпetatarsi. metatarsi. Тне The two represent гергевепн a а vicarious pair of ое species. They Тнеу differ ёюеесг mainly тагп1у in the genitalic зёгпсоге: structure: in гл S. ziminae ziпгinae the аеасаоз aedeagus is 1опу long апд and upright аргса11у apically (without triangular apex арех directed дггеснед dorsally as гп in S. a(ghanica); afghanica); parameres of о(5. S. afghanica afglгanica is symmetrical апд and longer than those thosc of new пе ' species. The third species with a а similar ornaтeпtation ornamentation of legs (Ьин (but without mid тгд metatarsal brush) - S. S. ohsoleta obsoleta Х11. Fil. (Europe, (Еиторе, Turkey) Тигкеу) Гиаз has the тost most distinet distinct genitalia and апд differs from (гоге two ыиег other species in the external exterпal characters. Тне The new пс species is is dedicated to L.V. Zimina. Ziтiпa. Еагпг1у Family Stroпgylophthalгnyiidae Strongylophthalrnyiidae The Strongylophthalmyiidae Stroпgylophthalmyiidae аге are а a small family of Аса1урнгане Acalyptrate flies represented гергевепес! Ьу by stгiпkiпgly strinkingly elegant elcgant creatures creatu.res with graceful гасеео1 legs and апд elongate е1огхае body. Ьоду. The Тне family includes iпcludes three ннгее genera: епега: Longinasus Loпginasus Егеу Frey [1955] with one опе Brasilian Brasiliaп species;nartshukia Shatalkiп Shatalkin [1993b] [1993Ь] with only оп1у Oriental Qriental species; Strongylophthalmyia 5trongylophthalmyia Не11ег Heller occuring occuriпg in the Neaтctic Nearctic (1 species), Palacarctic Palaearctic (8), Oriental (23), and апд Afrotropical Аfгонторгса1 (2) regions [Frey, [1 геу, 1955; Iwasa, 1992; Steyskal, еув1са1, 1971]. Неге Here I 1 confine myself to analysis of two species угоир group of the genusstroпgyloplгthalmyia. Strongylophthalmyia. Species species ое of thecrinita угоир group possess роззезз реси1гах peculiar genitalia. They Тнсу Еогю form а a топорну1- monophyl- etic угоир. group. Species species the ршкаагопр punctatagroup are distinguished Ьу by male та1е antennae апtеiипае with the third segmeпt segment having the догва1 dorsal process ргосезз пеаг near the base ое of arista. PossiЫy, Possibly, this group гоор contains contaiпs phylogenetically рну1оуепенгса11у different дiffereпt forms. Еоггвз. Species species of Strongylophthalmyia 5trongyloplгthalmyia crinita group гопр Тне The distinguishing characters of this group гопр аге: are: Ьоду body black, Irons froпs апд and face пасе eпtirely entirely Ыаск black (ехсе$ (except f0r5. for S. indica sp.n.), ' wing юп unicalaraus unicolorous ( (except ехсерн for Еог S. pectinigera sp.n.), legs yellow, hind femur fетиг with brown subapical sиbapical тгпу, ring, genitalia of ое males та1ев with elongate е1опуане crooked craoked ерапдгга! epandrial ргосеззез processes апд and деие1оред developed sигstyli surstyli jhenпig, jhennig, 1940]. Species species ос of the group угоир exhibit ехыыё sexual dimorphism дrmorphism in the structure structuгe оf of ра1рг. palpi. Females of а!1 all species are characterized characteriгed Ьу by ра1рг paipi ое of the same загве type, Ёуре, elliptical elligtical in form, fогт, slightly enlarged епlагуед apically and апд black. Ы аск. Males Маев show з1ио ' a а modification тодгfгсангоп of ра1рг palpi to form Еоггл elongateд, elongated, narтowed narrowed in their base р1аев plates with дiverse diverse огпатепнангоп ornamentation оf of the hlack black scales апд and 6ristles bristles (Figs 22-25). Кеу Key но to species Species of the Strongylophthalmyia criпita crinita угоир group 1(2). Face Расе yellow, уе11ош, anterior aпterior part раrt of оf Irons froпs yellow. уе11ош. Male Маlе ра1рив palpus уе11ош, yellow, black in аргса1 apical ha1f. half. Сепгна1га Genitalia Fig. 18. India Iпдга S. indica sp.n. 2(1). } Face асе and апд Irons fro пs whally wholly black. black. 3(4). Wing slightly intimated iпfurпated apically. аргса11у. Male MaIe palpus - Fig. 24. Genitalia Genitaha - Fig. I. 19. ihailanд Thailand S. S, pectinigera sp.n. 4(3). Wings transparent. 5(6). Smaller: тт. mm. Маме Male ра1рив palpus - Fig. 25. Genitalia Geпitalia - Fig. 20. нпдiа India S. spinipalpa вргпграlра вр.п. sp.n. 6(5). Larger: Male Маlе palpus and апд genita1ia genitalia of different differeпt fогт. form. 7(8). Basal Ваза! section sectioп of МM1+2, restricting с!юзса! discal се11 cell till на ta аьоиё. about times as short shoтt as following followiпg section. sectioп. Маlе Male ра1рив palpus (Fig. 23) with one опе but Ьин wide scale вса1е apically. еепюанюа Genitalia - Fig. 17. Burma, Вигта, Vietnaт Vietnam... S. freyi sp.n. вр. п.

8 152 А.I. A.I. Shatalkin Shatalkiп 8(7). Basal Вава1 section of о M.1, М12, restricting discal cell се11 till ti11 ta Вава1 Basal section of ое М1+2, Mi+2, restricting discal се11 cell Ё1П till на ta аьоин about аьоин about times а as short as following section. sectioп. 1.7 times as short as following section. sectioп. На1 Halteres егез yellow. 9(10). Male Маlе palpus ра1рцв (Fig. 22) with two 1 о - three Нгее scales scalеs Abdomen Ыасk, black, slightly shiniпg. shining. Geпitalia Genitalia (Fig. 17) аргса11у. apically. Genitalia with epandrial process ргосевв (Fig. 16) ]6) with strong stгong crooked стоо'сеа epandrial lobes. twisted. Taiwan, Тагшап, Jарап, Japan, 1Far-East аг-еа (Russia) Воду Body leпgth length 3.8 mm. тт. S. crinita Hennig. Henпig. ЕЕМАLЕ FEMALE resembles геветыев та1е; male; ра1рi palpi Ь1аск, black, narmal normal oval оиа1 10(9). Male Маме palpus ра1рцв with spinules. Genitalia with epan- ерап- Еотш. form. Body Воду leпgth length mm. тт. дгга1 drial рrосеss process (Fig. 21) crooked. croaked. Vietnam Vietпam DIAGNOSIS. DIАGNOSIS_ The Тне male та1е of S. freyi freryi sp.n. вр.п. is similar to S. verrucifera veп-ucifera sp.n. вр.л. that of ог S. crinita in the {Ье structure structиre of palpi сгошпсд crowned with Ыаск black scales еs арiса11у apically (2-Э (2-3 in S. crinita сгiптна апд and 1 in S. (reyi), freyi), Strongylophthalmyia Strongylophthalптyia crinita Hennig, Henпig, 1940 Ьин but differs mainly тагл1у in their theiг genitalia. Ерапдгга1 Epandrial processes ргосеззсз гп in Figs 1б, 16, 22. S. crinita are azе sнгапуег stronger twisted. This latter Iatter species specics Ваз has а a МАТЕА1АI.. MATERIAL. Тура Typos:0', :, 5-Ропюоа, S.-Formosa, Ко5ел1рс Kosempo, 2.08 (Н. (H. 5aиter) Sauter) тисн much sтaller smaller тatio ratio of sections ое of М Mi,, restricting discal дiscal S.V. SV (DEI); (DEI), 2 CPO', дд, 3?Y,??, Taiwaп Taiwan сепнег center Chiayi, 25 km 'сш Е, E, 700 m, т, сеl1. cell. 9.X Х.1993 (F. Кар1ап, Kaplan, А. A. IFreidberg) г iсiьец) (ТАн1). (TAU). I name паше thls this new леш species specics in нопоиг honour of the distin- дlstin- DESCRIPTION. MALE. МАШЕ. Froпs Frons entirely black, Ыаск, shin- shiлing. Face Еасе black, Ы асk, matt. Parafacials dark brown with with noticed that that he не [Frey, 1955] 1955) доиьнед doubted justly пз 1у that that the the silvery dust. dцst. Basal Ваваl antennal segments yellow; уе11аш; third Вигтеве Burmese specimeпs specimens being а at his disposal Ье1опуед belonged to segment segmeпt browпish brownish уе11ош yellow in in 1отег lower part and апд greyish 5. S.crinita. степiёа. guished Finnish FinпisЬ dipterologist Richard Frey. It зьопна It should be Ве арiса11у. apically. Arista аах'с dark browп, brown, bare. Ьаге. Mouthparts Мо Ьраг brown; palpi yellow, with 2-3 leaf-like leaf-1ike Ыаск black scales apically арiса11у аж! and Strongylophthalmyia indica Shatalkin Shata1liin sp.n. bristle-like scales са1е оп on 1анега1 lateral margin (Fig. 22). 3 ог, or, 2 vt, Fig. 18. ру pvt preseпt. present. Thorax Тйогах black. Ыаск. Mesonotum Mesnnatum shining, shiniпg, with МАТЕА1Аi.. MATERIAL. Но1онуре: HoIotype: д, 0", lndia, India, Меу11аlауа, Meghalaya, Nongpoh- obvious short ра1е pale hairs. 2пр1, 2npl, 1 1с. dc. Seutellum Scutellum Ьаге bare with Forest, Fvrest, IV V.1980 (Freidberg) (Freiclberg) (USNM). а a рагг pair о( of stout stoиt bristles ЬristIes apically арiса11у and a а pair of of weak wеak hairs DESCRIPTION. МАШЕ_ MALE. Неад Head Ыаск. black. Fгons Frons in гп Ьава11у. basally. Legs Ее yellow, hind femur feтur with brown browп subapical anterior part above аьоое antennae апнелпае yellow. Расе Face уе11ош, yellow, matt, гiпу; ring; fore fаге and апд middle тiдд1е femora егвога with weak геа1 darkening дагкелiпу mouth with black Ыаск bordering. baгdering. Parafacials yellow. yellow_ Basal Basa1 аргса11у; apically; hind hsnd tibiae with very егу weak 'еаж darkeniлg darkening in basal алнслпа1 antennal segments yellow; third thirд алнеппа1 antennal зс8твеы segment greyish-yellow. Arista ааг'с dark,, Вате. bare. Mouthparts brown; palpi palpi the level 1еие1 of tp. нр. Basal Вава1 section of о М M1+2, restricting discal уе11ош yellow аж! and deep ар black Ыаск in apical арiса1 haif. half. 3 ог, or, 2 vt, pvt р' third. thiгd. Wings Wiпgs transparent. traпspareпt. К23 R2+3 long, 1опу, its К end ела far beyond Ьеуопд се11 cell ti11 till In на about аьвин times as а short as following followiпg present. preseпt. Thorax Тногах black. Ь[ас'с. Mesonotum Мезопьпш shiпing, shining, with obvi- section. На!Ёеге Halteres yellow. уе11ош. АЬдотеп Abdomen Ыас'с, black, slightly ing. iпg. Genitalia (Fig.16) 16) with twisted epandrial ералдгiа1 processes. ргосе5 е. а a раiг pair of ое stout зоы bristles apically аж! and а a pair ое of weak еа'с hairs Воду Body 1eпgth length mm. шв. basally. Legs уе11ош, yellow, hind нiпд fетиг femur with brown subapical виьарiса1 shinous short ра1е pale hairs. 2 прl, npl, 1 I dc. дс. ScutelIutп Scutellum Вате bare with ЕЕМАLЕ FEMALE resembles Ыes но to ша1е; male; ра1рi palpi black, Ыас1, normal погта1 ring; riпg; fore and middle тiдд1е femora fетога with weak еа'с darkening darkeпing elliptic form. fогт. Body Воду length з mm. тт. apically. Wings Wiags transparent. traпspaгent. ГR long, 1опу, its end ела far Тнгоиун Through the kiпdness kindness of of Гг. Dr. F. Menzel Мепге1 of ое the Ьеуопд beyond the level 1еие1 of tp. R, R +3 апд and М1 M1+2 slightly convergent coпvergent 1ЭепЁс11с5 Deutsches Eпtimologisches Entimologisches Institut (Eberswalde) I have Ьае I аргса11у. apically. Basal Ваза! section of оем 1+, restricting discal се11 cell till Ьесп been аые able to study the male та1е нуре type of ое this species. specics. As М it ta about aboиt 1.4 times as short as as Еoliowing following sectioп. section. На1негев Halteres нигпед turned оин out Hennig Неллi у [1940] depicted depicted the ge the genitalia пitalia of ое S. S. whitish, with yellow уе11аш knob. 'снов. Abdomen АЬдотеп Ыаск, black, slightly стi crinita ггi на пы not quite exactly. ехасн1у. I had над doubts douьts that numerous питегоцв shiniпg. shining. Genitalia as as in iл Fig. 18. specimens врес1тепв fгот from дарan Japan апд and Еаг Far Еа$ East неге were conspecific coaspecifie with Воду Body lcngth length 3.5 mm. тт. S. crinita. А A сотраггвап comparison with the type уре showed ьоеа that these ЕЕМАLЕ FEMALE unknown. ипкпошп. specimens юп$ё must Ье be referred to to S. crinita. сгi пгна. О DIAGNOSIS. 1А6Ы05I8. In 'п Steyskal's key ог of thegenusstrongylophthaimyilophthalгnyia [Steyskal, [5teyskal, 1971]5. S. indica гпдгса sp.n., вр.п., as having to S. freyi sp.n. вр.п. and апд S. verrucifera sp.n. sp_ а. externally ехнекпа11у but Вы is the face асе who11! wholly yellow апд and the Етопз frons yellow in anterior anteгior S. crinita criпita is а a н. usual па1 species specics in Еак Far Еа.. East. It is very оег! siтilar similar дistinguished distinguished Ву by the св characters гасетэ given сп iп in the key above. аьоие. раrt, part, would ' опна go to ta thes. ивнц1ана ustulata угоир. group. Тне The печ' new species specics differs Ггош from species specics of thes. ustulata ustиlata угаир group in гп possessing possessiпg Strongylophthalmyia freyi Shatalkin sp.n. the peculiar реси1iаг type уре of genital structure Ьeing being similar to to Figs 17, 23. that ofs. crinita. criпita. Тне The та1е male ра!рп$5. palpuss. пдiса indica is Iike like those МАТЕЕ1АЕ. MATERIAL. Но1онуре: Holotype: д', cr, NE НЕ Вита, Burma, Kambaiti, 2000 trt, т, ое of S. pectinigera pectiпigera sp.n. and апд S. spinipalpa sp.n. Within 9.VI V (Ма1агве). (Malaise). Ратауре Pararypes: 5?', 5, same вапхе locality Iocality as holotype, 15- the S. crinita утоир group S. indica is Ьез best rccognize recognized д Ьу by the 25.V У.1934 (Malaise) (Z(ZMUH). МИН). со1оиг colour of the head. неад. DESCRIPTION. МАLЕ. MALE. Froпs Frons entirely black, Ы асk, shin- shiлing. Face Еасе black, Ы асk, matt. Parafacials Рага(ас!а1 dark дагк brown with Strongylophthalmyiapectiпigera pectinigera Shatalkiпsp.n. kinsp.n. silvery dust. dцst. Basal Ваа1 antennal antenual segments yellow; third д Figs 19, 24. segment Ьrownish brownish уе11ош yellow in in 1о lower 'ег part and greyish МАТЕАгА1.. MATERIAL. Но1ануре: Holotype: д`, 0", Thailaпd, Thailand, Ы NW Chiang-Mai, Doi арiса11у. apically. Arista с1аг'с dark brown, Ваге. bare. Mouthpaкts Mouthparts brown; ~и1сен, Suket, m, т, Х.1993 (F. Kaplan, Кар1ап, А. A. Freidberg) ((TAU). ТАИ). ра1рi palpi yellow, with wide widc Ыас'с black scale арiса11у apically and алд setules sctиles DESCRIPTION. MALE. МАШЕ. Froпs Frons eпtirely entirely Ыас1с, black, shiпiпg. Face Еасе black, Ыаск, matt. тatt. Parafacials дакк dark brown with shining. оп on 1асга1 lateral margin maгgin (Fig. 23). 3 or, ог, 2 vt, pvt ру pгesent. present. Тната Thorax Ыас'с. black. Mesonotum Мевопо1игп shiniпg, shining, with obvious short shoгt silvery dust. апзё. Basal Вава1 antennal anteпnal segments brown; third гд ра1с pale hairs. 2npl, 2пр!, 1 I de. 1с. Scutellum 5сиtе11ит Вате bare with a а pair раiг of ое stout алнеппа1 antennal segment segтent yellowish огапе. orange. Аrista Arista дагк dark Ьгошп, brown, Ьristles bristles apically апд and а a pair of weak ёа1 hairs basally. Ьава11у. Legs Ьаге. bare. Mouthparts МопЁЬратз brown; palpi ааг'с! darkish, Ь, with comb соть of yellow, hind femur fетиг with brown subapical sцbapical ring; riпg; fore and апд Ьristles bristles арiса11у apically (Fig. 24). Зог, 3or, 2 vt, pvtp present. гcsent. Тногах Thorax тiдд1е middle (стога femora with weak са1 darkeniug darkening apically. Wings Wiags Ыаск. black. Mesonotum shining, shiпing, with obvious oьvious short pale ра1с transparent. transpaгent. R24.3 R 2+3 long, loпg, its сна end far beyond Ьеуопд the level 1е~е1 of оf tp. hairs. 2 прl, npl, 1 I de. ас. Scutellum Вате bare with а a раiг pair of з1оыё stout

9 New Ге апд and 1ittle little known 1ntоwn Lauxaniidae 1 апхапii ае & Strongylophthalinylidae топуу1орннна1гпуггдае flies 153 Figs 1б Ма]е Male terminalia tertninalia (1б-21) (16-21) of ое 5Strangylopbthalmyia: п ръй 1пуiа 5. S. crsnfta crinita Неппгу Hennig (16); S. 5. freyi sp.n. рл. (17); S. гпдгса indica sp.n. (18); S. pectinigera pecfinigera sp.n. (19); S. а.р тггра1ра вр.п. sp.n. (20); S. verrucifera venucifera sp.n. (21); See 5ее Figs 1-7 for Гог abbr abbгeviations. Рис. rue, Гениталии Fenuramiu самцов cantina Strongylophtharayice Strongylophthalmyia: S. crinita Heпnig Hennig (16); S. 5. freyi вр.п. sp.n. (17); S. S. гпдгса indica sp.n. (18); S. pectinigera pertinigera sp.n. (19); S. 5. вргпгра1ра spirtipalpa sp.n. вр.п. (20); S. verrucifera sp.n. (21);, Aix ля расшифровки pacnnutpoaau сокращений coupaujetuni смотри caorps puc. рис. 1-7.

10 154 A.I. Shatalkin Shalalkin bristles apically аргса11у and апд a а pair рагг of Ы weak hairs basally. Ьава11у. Legs weak darkening аргса11у. apically. Wings нгапврагепн. transparent. 112+, R2+ 1опу, long, yellow, hind нгпд femur fетиг with brown subapical ring; пg; fore and ю its end епд far аг beyond Ьеуопд the 1еие1 level of ое tp_ tp. Вава1 Basal section п оf of М1 M,+2, +з, тгдд1е middle femora without darkening пg аргса11у. apically. Wing infumated apically. аргса11у. 11,2 12+ iпfuтанед restricting restrictiпg discal се11 cell till ti11 ta аьо about times as a short Ьогt long, 1опу, its end епд far маг beyond Ьеуопд the 1ее1 level of Ы аз as following Ео11о 4п8 sectioп. section. Halteres На1 егез yellow. Abdomen black, Ь1аск, tp. Вава1 Basal section п О of М m1+2,. restricting гieting discal discal се11 cell ti11 till ta ta Slightly SIightly shining. shiпiпg. Geпitalia Genitalia аз as in Fig. 21 with crooked стоокед аьот about 1.8 times as а short as following section. sectioп. Halteres На1 еге epandrial processes. yellow. Abdomen АЬдотеп blaek, black, slightly shining. shiniпg. Genitalia as in in Воду Body 1eпgth length 3.5 mm. тт. Fig. 19. ЕЕМАLЕ FEMALE resemы resembles таiе; male; palpi Ыасk, black, normal погта1 oval оиа1 Воду Body leпgth length 3.6 mm. тт. form. огю. Body Воду lепgth length 3.7 mm. тт. FEMALE unknown. ипкпашп. DIAGNOSIS. DIАGNOSIS. S. verrucifera Uerrucifera sp.n. зр.п. is similaг similar to S. DIAGNOSIS. Within the the investigated пvestigated i group, угоир, S. crinita гп in size з!ге апд and арреатапсе. appearance. I наие have Ьееп been ипаые unable to find fiпd pectinigera sp.n, р.п, is readily геадг1у distinguished Ьу by the apical аргса1 апу any external ехtегпа1 characters that Ьа should be Ье sufficieпt sufficient to daгkening darkening of wing. The ТЬе та1е male is similar но to these Ьее of S. separate верагане these Ьезе two г 'о species. The ТЬе structure of of ра1рг, palpi, crowned сгашпед iпdica indica sp.n. р.п. and апд S. spinipalpa вр.п. sp.n. in the structure гuctuгe of with black Ыаск scales apically аргса11у in S. 5. crinita, is ПО not quite clear с1еаг ра1рг, palpi, crowned with only оп1у bristles in i these Ьее species. гп in са.е case оf5. off. verrucifera verrгюi fега Ьесапве because of their Ьад bad visibility. ТЬе The оп1у only characters гs of of value иа1ие are аге those Пюзе of of the the genitalia. genitalia. The The Strongylophthalmyia spinгpalpa spinipalpa Shatalkinsp.n. ерапдгга1 epandrial processes in in genitalia of of S. S. verrucifera, verтucifera, unlike Figs 2о, 20, 25. those of ое5. S. crinita, аге are less twisteд. twisted. In this Ыз respect rеsресt ннеу they аге are МАТЮА1. MATERIAL. Ноlонуре:0 Holotype:Cr,lndia,, India, Meghalaya, Nongpoh-Forest, Nongpoh-Po гest, гпоте more like those of Ы S. 5. freyi. The НаЁег latter species differs from IV V.1980 (Fzeidbezg). (Freidberg). Раганурев: Paratypes: 1 д, 1, 3 W, У?, same locality Iocality as as S. verrucifera г~erпtcifera Ьу by a а much тисн larger ratio гапо between Ьеншееп the holotype, У1980 V.1980 (Freidberg) (USNIvI). (ЦЫМ). successive sections sectioпs of Ы -M1+2 М 1+2, restrictiпg restricting discal cell. се11. DESCRIPTION. MALE. МАI.Е. Frons Frans entirely black, Ыаск, shining. Face black, Ыаск, matt. Parafacials dark датк brown with species Species of Strongylophthalmyia punctata угоир group silvery dust. йы. Basal Вава1 antennal апнеппа1 segments segmeпts yellow; уе11ош; third antennal segment segтent brownish yellow in lower 1оег рагн part and апд This group гопр is restricted restricteд to ta three previously pгeviously described deseгibed shiпiпg. greyish apically. аргса11у. Arista Агista dark brown, bare. Ьаге. Mouthparts апд and three new пе species. Two То more тоге species вресгев considered coпsidered brown; palpi yellow, уе11ош, triaпgular, triangular, with thickened black Ыаск ЬеНош below тау may Ье1оп belong Ёо to i it too. оо. Species species оf of the group гопр are аге bristle at а apical аргса1 corner согпег апд and 4-5 setules Ье1о below (Fig. 25). гесопюаеа recognized тагп1у mainly Ьу by of the male 3 ог, or, 2 т vt,, the third segment of the та1е pvt рч preseпt. present. Thorax Тнотах black. Ыаск. Mesonotum Mesonotuт апнеппае antennae with dorsal expansion. shiniпg, shining, with abviau obvious short Ьог ране pale hairs. 2 пр1, npl, 1 dc. 1с. Scutellum bare Ьаге with a а pair of of stout 1оц1 bristles Ьristlеs apically аргса11у ала and Strongylophthalmyia freidbergi Shatalkinsp.n. а a pair рагг of weak hairs basally. Ьава11у. Legs LegB yellow, уе11ош, hind femur fепьиг Fig. 26. with brown subapical виьаргса1 ring; riпg; fore and middle тгдд1е femora with МАТЕА]Аi.. MATERIAL. Но1онуре: Holotype: С?, d, Thailand, ыхт NW Chiang-Mai, Doi 1]oi шелк weak дaгkening darkening apically. Wings transparent. гапрагепё. 112+, R2+ 1оп long,, Suket, Sцket, m, т, 12.X К.1993 (F. Kaplan, Кар1ап, A. А. Freidberg). its end епд far аг beyond Ьеуопд the level 1еие1 of Ы tp. Вава1 Basal section sectiвn of о М M1+2,.2, Paratypes: 220"с?, Co', same locality носа1гну as as holotype,12х X.1993 (F. Kaplan, Кара, restricting restrictiпg discal cell сен1 Пи till на ta аьоин about 1.3 times as a short хог1 as А. A. Freidberg); VY, 299, Tailand, Таг1апд, S.Takua Pa Ра апд and S.Khao 5.Кнао 5окпанраг, Soknatpar, 21- following followiпg section. веснгап. Halteres На1негев yellow. уе11ош. Abdomen АЬдотеп black, Ыаск, 22...X _Х.1993 (F. КарIап, Kaplan, А. A. Freidberg) (TAU). (ТАП). slightly shining. shiпiпg. Geпitalia Genitalia as гп in Fig. 20. DESCRIPTION. MALE. МАLЕ. Head Неад Ы black. асk. Frons Гтолз in anterior part рагн above аьоие antennae апнеппае yellow. уе11ош. Face Расе yellow, matt. ЕЕМАLЕ FEMALE resembles геветыев таiе; male; palpi Ыаск, black, normal погта1 oval оиа1 Parafacials yellow. Basal Ваза1 antennal алнеппа1 segments segтents brownish bгowпish апнеггог $оду Body lепgth length 3.1 mm. тт. form. fогт. Body Воду length 1ец mm. тт. уе11ош; yellow; third antennal апнеппа1 segment (Fig. 26) blackish ЫаскгвЬ (ехсер (except DIAGNOSIS. The пет new species is characterized Ьу by for fог yellow base) Ьаве) from outside and аса yellow уе11ош with blackish Ыackish гelatively relatively long 1опу basal Ьава1 section sectioп МM1+2 restricting гestrictiпg discal cell. се11. Ьогдеггпу bordering from inside; депве dense hairs оп on margin Ы of thiгd third In Iп this Ы$ character chaгacter S. spinipalpa sp.n. р.л. is similar siтi1ar to to S. апнеппа1 antennal segment segтeпt black. Ь1аск. Antennal Апнепла1 process ргосезз Ыас1, black, with crinita апд and S.indica differing differiпg froт from the second есопг! species гп in депве dense Ы black асk hairs. Arista dark, bare. Ьаге. Mouthparts Моиннрагнв brown; the colour со1оиг of the frons гоп апд and часе face being wholly Ыаск; black; from ра1рг palpi brownish, undelated. 33 or, or, 2 vt, 2 т, рргезепё. pvt present. ТЬогах Thorax the first species5. S. spinipalpa should be 6е distinguished distiпguished by Ьу Ыаск. black. Mesonotum Мевопонит shining, shiniпg, with obvious oьvious зжог short pale ране hairs. the smaller smallcr size, the genitalia geпitalia with epandrial epaпdrial processes 2 пр1, npi, 5 ас, de, relativelly short except ексерн posterior pair. рагг. 1е less twisted, isteд, and апд the male та1е palpi ра1рг with only ол1у bristles brist1es 8ссе11пш Scutellum Ьаге bare with a а pair of stout stoпt bristles Ьristles apically аргса11у and аса (without (witьout black scales) са1е ) apically. а a pair рагг of Ы weak hairs basally. Ьа.а11у. Legs yellow, fore fоге апд and middle тгдд1е femora fетога with weak darkening дагkеагпg in apical one опе third; middle тгдд1е Strongylophthalmyiaverruciferc2Shatalkinsp.n. fera Shata1kiдsp.n.. and hind tibiae with wide wide pale ране brown Ьrown ring riпg in the middle. тгдд1е. Fig Нгпд Hind femora with transversely галз'егзс1у f1аннепед flattened basal Ьа.а1 tubercle. ниьегс1е. МАТЕР MATERIAL. ГАВ. Но1онуре:д', Holotype:d, Vietпam, Vietnam, Тат Tam Юао, Dao, prov.уinh Vint' Рьп, Phu, Wings with weak apical аргса1 spot and апд median тедгап crossband. crossьaлd. R m, т, (Belokobylskij). Paratypes: 1 1 д`, Cr, 1y, 1 9, same 1оп long,, its end епд far Еаг beyond Ьеуопд the level нее1 of оf tp. R, R, +s апд and МM1+2 not пы 1оса1ггу locality as а holotype, но1онуре, (Belokobylskij) (Ве1о1 оьуы4) (ZISP). (25Р). coпvergent convergent apically. Вава1 Basal веснгол section о of М1+2, K+2, restricting гestrictiпg DESCRIPTION. MALE. МА1.Е. 1Frons гоп entirely Ыаск, black, shining. Face Еасе black, Ыаск, matt. Parafacials dark brown. First First sectioп. section. Halteres whitish, with witь knob!гпоь darkish in гп Ьа5аГ basal рати. part. shiпiпg. discal се11 cell П11 till ta аьоин about 1.8 times as аз short as аз following followiпg апtеппа[ antennal segment brown, second есопг! segmeпt segment уе11ош; yellow; third АЬдотеп Abdomen Ыаск, black, slightly shining. shiniпg. segment segтeпt dark даг grey. геу. Arista dark Ьrown, brown, Ьаге. bare. Mvuthparts Mouthparts Воду Body leпgth length 3.5 mm. пат. brown; ра1рг palpi dark, дагк, parhaps, рагнарв, with only оп1у spinules spiпules apically. ремаlе FEMALE differs from та1е male гп in the following followiпg characters: third thiгd antennal апнеппа1 segment segmeпt with dorsal аогзан knob кпоь noticeable noticeaы е сьагас- 3 ог, or, 2 vt, ч, pvt рч present. ргевепн. Thorax Тногах black. Ыаск. Mesonotum Mesonatum Ёегз: shiпiпg, shining, with abviaus obvious short ране pale hairs. 2 прl, npl, 1 dc. дс. гп in the paratype рагануре from Takua Pa; Ра; antennal апнеппа1 hairs whitish; wьitish; Scutellum Sсине11ит bare Ьаге with a а pair of Ы stout stoиt bristles Ьгг$н1ев apically арнсаl1у and ала алнегiог anterior femora Ьгошп brown yellow, шжа mid апд and hind нгпд tibiae а a pair of Ы weak hairs Ьаггв basally. Legs yellow, уе11ош, hind femur ешпг brownish; browпish; median rпedian crossband сговвьалд ое of wiпg wing ртаснгса11у practically aьsent. absent. with brown Ьrown subapical suьapica1 ring; riпg; fore and middle тгдд1е femora fетога with Leпgth Length 3.4 mm. тт. i

11 Ме' New аж! and 1itt1e little known Гпоп Lauxaniidae & Strongylophtlialinyiidae Strangylophtha1myiidae flies flies 155 Figs Ма1е Male palpi (22-25) апд and апнеппа1 antennal stnзcture structure (26-30) апд and genitalia (31) of о Strongylophtbalmyia: 5е'о уорыьаiтуiа: S. crinita Hennig; Henпig; S. freyi sp.n.; $р..; S. pectinкgera pectinigera вр.п.; sp.n.; 25 - S. вргпгра2ра spinipalpa вр.п; sp.n.; 26 - S. freidbergi Г йiье i вр.п.; sp.n.; S. giьfxfera gibbsfera Shatalkin; S. 5. тгсговну1а microstyla sp.n.; 29-5styiocera Sstylocera sp.n.; S. тапсотгв raricornis Shatalkin. Рис. Puc L4yrixic% Щупики (22-25),' (22-25);ycium исики (26-30) ci и rem,rramoi тениталии (31) camgm саиуов gpi Noo видоп Strongylophthaimyia: 5trongylopЬtha1myia: S. сггпана crinita Hennig; 3ennig; 23 S. freyi 1теуг вр.п.; sp.n.; S. реснгпгуета pectinigera вр.п.; sp.n 25 - S. spinipalpa гргпкраlра вр.п.; sp.n.; 26 - S. 5. freidьergi freidbergi вр.п.; sp.n.; 27 - S. gibbifera giъb fera Shatalkin; 5наУа.1кгп; S. microstyla micтostyla ар.л.; sp.n.; 29 - S.stylocera Ssfylocera sp.n.; рл.; S. raricornis Shatalkin. DIAGNOSIS. The male та1е ofs. freidbergi вр.п. sp.n. may тау Ье be basally. Legs yellow; уе11аш; hind ию '! femora (етога with weak еа1с darkening дагкепгпу easily recognized Ьу by the аьаре shape of апнеппа1 antennal ргосе5в. process. Otheт Other аргса11у; apically; Ьгдд hind tibiae darkened in тедгап median 1/2. Wings unique снагаснег character is the ргевепсе presence ое of the biack black (not (пои white transparent. traпsрarent. R2 3 R 2+3 1опу, long, its end едд far Гаг beyond Ьеуопд the level 1ее1 of tp. а as гп in нургса1 typical case) саге) hairs оп on anargiп margin оf of the third antennal anteпnal КR0.5 4 and апд М1+2 M1.5 practically not по convergent сапиегуепн apically. М, M1+2 2 веудпепн. segment. The стое true status of the females fета1ев as а5 S. freidbergi [reidbergi very егу сып thin in гп apical аргса1 half. ha1f. Basal Ваа1 section веснгоп of оf М Mi+2,+2,, restricting гequires requires additional additiaпal proof. ргооf. discal disca1 се11 cell ti11 till на to аьооё about 2.3 times as а5 $11ог short as а followiпg following It is my ту pleasure р1еа5ите to но name Нате this сы species ресгев in гп honor honar of section. Halteres greyish, with knob кпоь dark. дагк. Abdomen АЬдотсл РтоЕеот Professor Атпоп Amnon Freidberg Егого from Тем Tel Aviv University. Ыасk, black, slightly shining. Воду Body 1епсЬ length 2.3 mm. тт. Strongylophthalmyia gibbifera Shatalkin, S.atalkin, 1993 ЕЕМАЕЕ FEMALE unknown. цпкпошп. DIAGNOSIS. This distinctive species demonstrates Fig. 27. пstratcs а a close с1ове relationship S. microstyla MATERIAL. Species is known by the hoiotype from relatianshi.p with 5. below. МАТЕА1А1.. 5 ресiеs is кпошп Ъу Уне ноiонуре гол Vietnam, пaicrostyla described Ье1о. ргоу. prov. HanOi, Hanoi, 70 km NW t' Хт Ba $а Vi, 400 m, т, 22)( ХЕ.1990 (Nartshuk) (ZISP). (zi5p). Strongylophthalinyia Strongyloplгthalmyiamicrostyla microstyla Shatalkinsp.n. REDESCRIPTION. МАLЕ. MALE. Неад Head black. Ь1аск. Frons Froпs en- сп- Fig. 28. tirely black. Ыаск. Face Расе dark brown, matt. Parafacials Parafacia1s slightly slightly МА1ЕRГАL. MATERIAL Holotype: Но1онуре: д cs Philippiпcs, Philippines, Manila (R. Brown) 1 го л) silvery dusted. Basal Вааа! antennal апнеппа1 segments ешепё апд and ventral half (ЩS (USNM). г4м). ог of third segment segтeпt ycl1owish yellowish brown; с!ога1 dorsal half of third DESCRIPTION. MALE. Неад Head Ыаск. black. Frons Froпs гп in апнегiorior part рагн as а far Гаг as а ocellar осе11аг triangle yellow. уе11ош. Face Расе yellow, уе11ош, third antennal anнеппа1 segment scgmeпt whitish. Antenna! Алнеппа1 process (Fig. matt. Parafacials РаГаfасга1$ yellow. уе11ош. Antennae Алнеппае yellow; third anten- 27) short, triangular, with with dense депве white hairs. Arista па1 nal segment е8тоеос (Fig. 28) with short ЬотЁ style-form су1е-еогш process; ante- апнеппа1 antennal segment dark дark brown. Dense Геае hairs on оп margin of long, Ioпg, bare. Ьаге. Mouthpaгts Mouthparts dark brown; palpi narrow, паггоч, депве dense hairs on оп margin тагугп of third thirд antennal апнеппа1 segment е8гвепё and аос! brownish yellow with long 1опу bristle Ьгistle apically. аргса11у. 3 ог, or, 2 vt, апнеппа1 antennal process whitish. Arista датк, dark, Ьаге. bare. Mouthparts ру pvt present. Thorax Тногах black. Ыаск. Mesonotum Мевопонит shiпiпg, shining, with brown; п; palpi ра1рг delated, де1анед, black. 3 3 вт, or, 22 vt, vt, pvt ру present. obvious short ра1е pale hairs. 2npl, 2пр1, 1de. дс. Scutellum Ьаге bare with Тногак Thorax black. Mesonotum Мевопонит shiniпg, shining, with obvious short а a pair рагг of stout оо bristles apically aрically and апд a а pair of оf weak шеак haiгs hairs ра1е pale hairs. 2 прl, npi, 7de дс (4 re1ativelly relativelly short shorc Ьefare before suture), осоге),

12 156 А.I. A.I. Shatalkin ас ac brought together on оп middle тгдд1е line and апд equal еgиа1 to presu- ргевиtural ас. dc. Seutellum 5сине11ит Ьаге bare with a а pair of.stout bristles 6ristles apendages apeпdages оп on posterior вигfасе surface Ьава11у, basally. Wйgs Wings нгапвраг- transpar- interпal internal surface; нгпд hind (етиr femur with three small вта11 warty 'аг1 у аргса11у apically and апд a а pair of Ы weak еа1 hairs basally. Ьава11у. Legs 1е yellow, уе11ош, епн. ent. R.2+3 К 1оп8, long, its ела end far iат Ьеуопд beyond the Ье Iес1 level Ы1р. of tp. 1 R,.'5 4+5 апд and fore вге сохае, coxae, middle апд and hiпd hind coxae сохае and апд femora, fетога, middle тгддlе МЕ M not пон convergent convergeпt apically. аргса11у. Basal Ваваl scctiaп section а of 14142, tibiae гп in Ьаа1 basal 1/2 part, раг, hind tibiae in basal 6ава1 2/З 2/3 part рагн restricting discal се11 cell ti1i till {а ta about аьоцн 1.8 times as short as as darkish brown. Fore Еоге femora with strong straпg dorsal догва1 and апд following follawing зеьиоо. section. Halteres На1негев darkish grey. геу. Abdomen АЬдотеп veпtral ventral bristles. Wings Wйgs transparent. R +3 very иегу short, shoгt, its Ыасk, black, slightly shining with blackish Ьlackish hairs. Genitalia Geпitalia as епд end Гаг far Ьеfоге before the 1е~е1 level Ы of ф. tp. Т R0.5 and- апд2 МM142 2 convergent сотчегеп й in Fig. 31: surstyli absent. аьвепн. apically. Basa1 Basal section еьиоп о1 of М,+ M,,,, restricting discal се11 cell till tiii Воду Body leпgth length mm. тт. на ta about 3.6 times as as short as а #allowing following section. Halteres ЕЕМАLЕ. FEMALE. Similar Sгтг1аг но to та1е male with the following followiпg exceptions. Antennae Аыеппае with оп1у only вта11 small ниьегс1е tubercle догва11у. dorsally. Hind Нгпд ехсерtioпs. very егу dark. Abdomen АЬдотеп Ыаск, black, slightly shining. Воду Body leпgth length 2.3 mm. тт. fетога femora without appendages аррелс!аез Ьава11у. basally. Воду Body leпgth length ЕЕМАЕЕ FEMALE unknown. ипkпоwп. тт. mm. DIAGNOSIS. S. microstyla sp.n. 5р.П. is similar siтilar to но DIAGNOSIS. This distinctive distiпctive species is the sole S.gibьifera S.gibbifera Shatalkin in гп antenna! апнеппа1 structurc, structure, but Ьин differs representative о of the the group гопр й in the Palaearctic region. ге8юоп. ю in со1оиг colour characteгistics characteristics Ы of the Ье head неад and апд 1е legs.. Characters С1тагаыег Within the group угоир S. raricornis raricornгs оссиргев occupies ап an isolated of the wing wйg with the unique uпique раннегп pattern of vein veiп R2.3 R1+3 indicate iпdicate position. positioп. Тне The third antenna' antennal segment segmeпt is bilobate Ьг1оЬаСе with a а close сi о$е relationship between Ьеншееп S. microstyla апд and S. S. еотрагаь1е comparable dorsal апд and иепнга1 ventral lobes. 1оЬев. гттаси1ана immaculata Hennig. Henпig. The Тне 1аннег latter species was described dcscriьed оп on the material тaterial from fгот Taiwan. Examination ЕХатйангоп of Taiwan Strongylophthalmyia Strongylaphthalmyia stylocera material kindly sent by A. Shatalkin зр.п. sp.n. тaterial kгndly seпt Ьу Dr. А. Freidberg shows that specimens 5pecimeпs which with good оос! reason геавод may тау Ье be ге(еггес! referred но to S. Fig. 29. immaculata, imnzaculata, аге are very егу similar WS. 1о5. microstyla antenna' МАТЕАiАL. MATERIAL. Но1онуре: Holotype: d', Mindanao, Cotabato, in апнеппа1 д, Philippines, Мг дааао, СонаЬан0, Вагго Bario Lаудаап, Lagdaan, IV.1932 (P. (Р. Rlvera) Rivera) (i.isnm). (USNM). structure stгucture апд and also а1во in the fore femora that that are аге armed. агтед. DESCRIPTION. MALE. МАLЕ. Неад Head Ы black. асk. Frans Froпs entirely entireiy Henпig Hennig [1940] дгд did not рогпн point оин out this ЁЫ characters с1 агас1 ег гп in the Ыаск. black. Face гасе yellow, глаёё, matt. Parafacials Рага~асга1в yellow with silvery description descriptian of Ы S. immaculata. гттаси1ана. Оме One more species specгes S. с!'тг dust,, gena цепа yellow. Antennae Аыеnпае yellow; уе11ош; dense депве hairs on оп spinosa Frey, known кпошп from Burma Вигта and апд having analogically агтед armed fore femur, feтur, товн most likely тust must Ье be included гп in this ргосезв process (й (Fig.. 29) 1опу, long, black, Ы асk, with dense депве white hairs. апа1аугса1- margin of third Ыгс! antenna' апнеппа1 segment segmeпt whitish. Antenna/ Апнеппа1 Ту 8гоыр. group. Uпfortunately, Unfortunately, arista was was broken 6roken in in the the holotype Агista Arista short, зёу1е- style-form, огго, Ьаге. bare. Mouthpaгts Mouthparts brown; Ьгошп; palpi Ы of S. spinosa. уе11ош yellow with apical аргса1 1опу long dark hairs а1опу along veпtral ventral margin. тагугп. 3 or, ог, 2 vt, pvt М present. pгesent. Thorax Тногах black. Ь1аск. Mesonotum Мевопонит Strongylophthalmyia Strongylaphthalmyia punctata Hennig, Henпig, 1940 shining, shiпing, with obvious short ра1е pale hairs. 2 пр npl,, 1 dc. дс. Scutellum Scute1lum Ьаге bare with a а pair рагг of stout bristles Ьristles apically аргса11у and апд I have Ьае not seen this species described from Taiwan. Тагшап. а a pair рагг of Ы weak hairs basally. Legs yellow; уе11аш; hind нгпд femora Тне The та1е male is similar to that of Ы S.stylocera 5. у1осега sp.n. р.о. in the with weak darkening пg аргса11у. apically. Wings transparent. нгапврагепн. 112., К structure strиctиre of third thiгд antenna] aпtermal segment веутепн but differs дiffеrs by Ьу тгдд1е middle. 1ong, long, its end епд far аг beyond Ьеуопс! the 1еие1 level оf of ф. tp. 1 R4+5 4 and M1.2 апд and нгпд hind fетога femora that are аге Ыаск black except ехсерt for Ьава1 basal third thirд аºд and +5 апд М 2 slightly convergent сопуеrgепt аргса11у. apically. Ваза! Basal sectioп section of М кпее knee yellow, апд and also armed fore,+2, а1 о агтед fоге femora. гestгictiпg restricting discal се!! cell ti11 till на ta about аьоин 1.8 times as short as аз following followiпg scctian. section. Halteres whitish, with with stem steт darkish Strongylophthalmyia Strongylaphthalmyia raricornis Shatalkin, 1981 in basal Ьава1 part. рагн. Abdomen АЬдотеп 61аек, black, slightly shining. пiпg. Figs Воду Body leпgth length 2.7 mm. тт. МАТЕАIАL. MATERIAL. ТЫ This species 'ч was девсггьед described оп on 2 C1 U " a" апд and 3 99?? ЕЕМАLЕ FEMALE unknawn. unknown. Егоггi from Khabarovskiy Ктау, Kray, Кнт. Khr. Маiуг Malyi КЫпап, Khingan, 15 km SE 5Е from этот DIAGNOSIS. ТЬе The distiпctive distinctive species is closely ге1анеed но to S. punctata рипснана demonstrating demonstгatiпg 1ikeliness likeliness ю in the struc- Primorskiy Pптorskiy Ктау, Kray, КатеПигнка Kamenushka (30 km 'пв SE $Е from Ussuriysk), Llssuriysk), 18- ниге ture of аf the third Ыгс! antenna! antennal segment зетеоё bearing Ьearing a а long loag process. 23.VII.1983 (Shatalkin); (Shatalkiп); 20'0% 2С с, 119, y, same саше locality, 1оса1гну, VI V relat- Аадде. Radde. Later the additioпal additional material ча was соllеснег1 collected 5599, yz Тне The male та1е of S stylocera вр. sp.n. п. may тау be Ье easily recognized (Shatalkin); (5hatalkin); 2д'д, Primorskiy Primoтskiy Kray, Ктау, Lazovskiy Reserve, Рееге, 25 km кт SE 5Е Егот from 1.аъо, Lazo, 10 аж' and 28,У VII.1986 б (Оепае) (Ozerov) (ZMUM). (ZМИМ). from а11 all species вресгев Ьу by the combination combiпatioп of the following followiпg REDESCRIPTION. MALE. Неад Head black. Ыаск. Frons Егопв entirely black. Ыаск. Parafacials slightly dusted дивнед with silver. ю1уег. with white hairs; hairs гs оп on margin оf of the third antenna' antenпal епtire[у снакаснегв: characters: апнеппаl antenna' process long апд and narrow, black Ыаск Апнеппае Antennae brown; third antenna' antennal segment тent (Fig. 30) segment segmeпt also white; Iegs 1ез yellow уе11ош and апд anterior anteriar femora fетога brownish brвwnish yellow basally with white dense депве haiгs hairs on од without stong stoпg dorsal догва1 spinules (distinction (дistinction from S. S. margiп; margin; this Ыа segmeпt segment 6ilobate bilobate with догва1 dorsal 1оЬе lobe (апtеnпа1 (antenna' punctata). pгocess process in other species) slightly longer loпger and more narrow Нгап than veпtral ventral one. опе. Arista dark brown, bare. Ьаге. Mouthparts Моиннрагнв АСКЫОЕ.ЕООЕМЕГГ8. ACKNOWLEDGEMENTS. I I wish to express ехргезз brown; ра1рг palpi Ыаск black with apical аргса1 long 1опу dark bristles along ту my sincere thanks to to the the following follawing persons регзолз for vcntral ventral тargin. margin. 3 ог, or, 2 vt, pvt р'т present. ргесепё. Thorax Тнагах black. Ыаск. иа1иаы valuable е help Ье!р in my ту work ог1 and алс! for loans of оf specimens: Mesonotum Mesonatиm shining, without withoиt obvious оь'топ short pale ра1е hairs. 2 Dr. E.P. Е.Р. Nartshuk, Dr. Dr. V.V. Z Z1oЬin lobin (both (ЬоЬ from (гош пр1, npl, 9 дс, dc, ae ас in й two rows. го '. Scuteilum синеl1ит bare Ьаге with a а pair рагг of Ы Zoological Zanlogical Iлstitute, Institute, St.Petersburg), Ре(егзЬог Dr. K. К. Elberg Е1Ьег stout bristles Ьгistles apically аргса11у and апд a а pair рагг of ' weak еаё hairs basally. Ьава1l у. in hind part and sternopleuron of zoology and botany, Tartu), Ms. E.M. Mesopleuron й hiпd рагн апд sternopleuroп with light (Institute оf оо1оу апд Ьо(апу, Тагни), М. Е.М. hairs. Legs Lеgs Ы black; асk; fore fоге соха coxa yellow with Ыаск black laterally, Апнопвча, Antonova, Dr. A.L. Ozerov Оего (Ьо(Ь (both from fгат Zoological тгддlе middle соха coxa уе11аш, yellow, hind coxa соха black; Ыаск; fore tibiae and Museum, Мозсот Moscow State (а(е University), Dr. Р. P. Arnaud Агпаид tarsi yellow; middle and апд hind hiпd tibiae yellow уе11ош apically; аргса11у; (California (Cali(ornia Academy Асас1ету ое of Science, 8ал San Prancisco), Francisco), Dr. middle midd]е апд and hind tarsi yellow except ехсерн for two Ёо 1аЁ last А. A. Freidberg, Dr. I. I. Yarom Уагот (both (Ьонн from Tel Те1 Aviv segments. Hind нтосьапнет trochanter with thorn-like apendage арепдауе оп University), Universi(y), Dr. F. F. Menzel Мепге1 (Deutsches Entomolo- Епногпо1о-