A REVISION OF THE AUSTRALASIAN SPECIES OF THE GENUS TETRAGNATHA (ARANEAE, TETRAGNATHIDAE)

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1 九州大学学術情報リポジトリ Kyushu Uiversity Istitutioal Repository A REVISION OF THE AUSTRALASIAN SPECIES OF THE GENUS TETRAGNATHA (ARANEAE, TETRAGNATHIDAE) Okuma, Chiyoko 出版情報 :ESAKIA. 25, pp.37-96, 九州大学農学部昆虫学教室バージョン :published 権利関係 :

2 ESAKIA,(25): A REVISION OF THE AUSTRALASIAN SPECIES OF THE GENUS TETRAGNATHA (ARANEAE, TETRAGNATHIDAE)* CHIYOKO OKUMA Etomological Laboratory, Faculty of Agriculture, Kyushu Uiversity, Fukuoka 812, Japa Abstract The Australasia species of the geus Tetragatha are revised based o the specimes from Australia, New Guiea ad eighborig islads. Thirty-oe species are recogized. New species, ew syoymies ad ew records of distributio are as follows : T. T. T. T. T. T, T. T. T. T. T. T. T. T. T. T. T. T. amoea sp. ov. from New Guiea. biseriata Thorell, = T. airesis Strad, New syoymy. = T. valoka Chrysathus, New syoymy. New record from Australia. bituberculata L. Koch, = T. heatwolei Chrysathus, New syoymy. ceyloica Cambridge, = T. eitupesis Strad, New syoymy. chauliodus (Thorell, 1890). New record from New Guiea. demissa L. Koch, = T. quadrides Dodale, New syoymy. eumorpha sp. ov. from New Guiea. gressittomcm sp. ov. from New Guiea. hirashimai sp. ov. from New Guiea. isularis sp. ov. from Lord Howe Is. isulicola sp. ov. from Lord Howe Is. jaculator Tullgre, = T. sp. B (Okuma 1968 & 1981). New record from New Guiea. moticola sp. ov. from New Guiea. aa sp. ov. from New Guiea. oreobia sp. ov. from New Guiea. priamus sp. ov. from Solomo Is. serra Doleschall, New record from New Guiea. sigata sp. ov. from New Guiea. * Cotributio from the Etomological Laboratory, Faculty of Agriculture, Kyushu Uiversity, Fukuoka (Ser. 3, No. 220).

3 38 C.OKUMA The geus Tetragatha Latreille, 1804 has bee kow from 56 species i the Australasia regio (Roewer, 1942 ; Brigoli 1983). However, o detailed study of the geus has bee made i this vast geographical regio. Recetly I was able to study the Australasia tetragathid spiders based o the collectio of the several museums ad istitutios. As a result 31 species were recogized from Australia, New Guiea ad eighborig islads. Of these, 3 species are already kow as the cosmopolita species (Okuma, 1983). The preset paper gives the descriptios of 11 ew species ad the redescriptios of 17 kow species. Illustratios of importat characters as well as measuremets of the relative legths of legs of the 31 species are also preseted. The umber of materials examied, ad the abbreviatios of the museums ad the istitutios are as follows : BMH : Berice P. Bishop Museum, Hoolulu, 356 specimes. AMS : Australia Museum, Sydey, 181 specimes. NSMT : Natioal Sciece Museum, Tokyo, 17 specimes. KUF : Etomological Laboratory, Faculty of Agriculture, Kyushu Uiversity, Fukuoka, 72 specimes. Termiology of the cheliceral armature of Tetragatha was established by Wiehle (1939). Later, Locket & Millidge (1953) adopted his termiology ad traslated it i Eglish i their book, British Spiders. I the preset paper, the terms are used accordig to Locket & Millidge. I additio, some ew terms are provided. These are summarized as follows (a asterisk shows a ew term) : a : A apophysis, used for lockig the female s fag durig copulatio. Gu : The guide tooth of upper row. *Gl : The guide tooth of lower row. *e : A extra tooth foud betwee (Gu) ad (T) i the female of some species. T : The first (largest) tooth of the row proper. *rsu : The upper row of small teeth. *rsl : The lower row of small teeth. *U2 : The ext tooth of (Gu). *L2 : The ext tooth of (Gl). *s : A spur or promiece, preset i the female of some species. t : A tooth or promiece, foud i the male of some species. sl : A tooth which usually slopes towards the base of the segmet i the male of some species. *AXu : A auxiliary guide tooth of upper row, preset i some species. *AX1 : A auxiliary guide tooth of lower row, preset i some species. EX : A excrescece foud o the outer edge of the fag, ear its base, i some species. Ackowledgemet I wish to express my hearty thaks to Dr. JoA Teorio of the B. P. Bishop Museum, Hoolulu, Dr. M. R. Gray of the Australia Museum, Sydey, Dr. H. Oo of the Natioal Sciece Museum, Tokyo ad Dr. Y. Nishikawa of the Ohtemo Gakui Uiversity, Osaka for the loa of may valuable specimes. My hearty thaks are due to Prof. Y. Hirashima of Kyushu Uiversity for his helpful suggestio ad ecouragemet throughout the course of this work. I am also grateful to Assoc. Prof. K. Morimoto ad Dr. 0. Tadauchi for their kidess i may ways. KEY TO THE SPECIES-GROUPS OF THE AUSTRALASIAN Tetragathu 1. Posterior row of eyes evidetly arrower tha aterior row of eyes

4 TETRAGNATHA OF THE AUSTRALASIA 39 - Posterior row of eyes ot arrower tha aterior row of eyes Male chelicera as log as or loger tha carapace... ~b~~et~>-group - Male chelicera shorter tha carapace Lateral eyes more separated tha media eyes Lateral eyes as close as or closer tha media eyes Abdome without a tail... jaculator-group - Abdome with a tail overhagig spierets Male chelicera with (a), (AXu), (sl) ad (T)... caudifera-group - Male chelicera without (AXu) ad (sl) ; abdome elogate... micmra_group 6. Posterior row of eyes wider tha aterior row of eyes Two rows of eyes equal or subequal i width Lateral eyes closer tha media eyes, situated o a tubercle...bi,t&rcu~~_group - Lateral eyes ot as above , Abdome about times as log as broad... radiata-group - Abdome more tha 3 times as log as broad Abdome more tha 6 times as log as broad... cylidm ca-group - Abdome about 3-5 times as log as broad......lo 10. Female chelicera with (s) ear outer apex.....serra-grou p - Female chelicera without (s)....i1 11. Male _ Female Chelicera with (sl) betwee (Gu) ad (T)... valida-group - Chelicera ot as above... I3 13. Chelicera with three strog upper apical teeth... ites-group - Chelicera ot as above (a) bifurcated at tip... maxillosa-group - (a) ot bifurcated at tip... I5 15. Tip of (a) prologed ad sickle-like (Fig. 29, A & B)... papuaa-group - Tip of (a) simply poited.....madibulata-grou p 16. Chelicera with very strog (AXl) ad (Gl)... madibulata-group _ Chelicera ot as above Chelicera with somewhat strog (AXl) ad small (Gl) : fag with strog (EX) ***.** ites-group - Chelicera ot as above... I8 18. Chelicera without (T).....papuaa-grou p - Chelicera with (T)....1g 19. Chelicera with (e) betwee (Gl) ad (T)...v&i&-group - Chelicera without (e)...ṁail/osa_group The rubrivetris-group Diagosis. Posterior row of eyes evidetly arrower tha aterior row of eyes ; male chelicera as log as or loger tha carapace. K EY TO THE SPECIES 1. Lateral eyes more separated tha media eyes i both sexes...y&m uetyis - Lateral eyes as close as media eyes i both sexes Female chelicera without (s)... p~eo&ctyla

5 40 C. OKUMA - Female chelicera with small (s) ear outer apex Teeth o female chelicera irregularly arraged as show i Fig. 3, A & B a--me-.amoea sp. ov. - Teeth o female chelicera rather regularly arraged as show i Fig. 4, G & H eumo@ha sp. av. Tetragatha rubrivetris Doleschall, 1857 Tetragathu rubtivetris Doleschall, 1857, Naturk. Tijdschr. Neel. Id., 13 : 410 ; Thorell, 1878, A. Mus. Civ. Geova, 13 : 105 ; Thorell, 1881, A. Mus. Civ. Geova, 17 : 131; Simo, 1894, Hist. Nat. Araig., 1 (3) : 718 ; Hogg, 1915, Tras Sot. Lod., 20 (14) : 454 ; Gravely, 1921, Rec. Id. Mus., 22 : 432 ; Roewer, 1942, Katalog der Araeae, 1: 985 ; Chrysathus, 1975, Verhadl., 140 : 11. Tetragatha lupata L. Koch, 1871, Arach. Austral., 1 : 170. Eugatha mbrivetris Keyserlig, 1887, Arach. Austral., 2 : 225. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head ; posterior row of eyes cosiderably arrower tha aterior oe (68-78 : 100) ; viewed from above, aterior row moderately recurved, posterior row somewhat strogly recurved, so that lateral eyes more separated tha media eyes; cetral ocular quadragle early square ; ratio of diameter of AME : ALE : PME : PLE as 88 : 75 : 75 : 75 (fl). Chelicerae. Log, basal segmet as log as or slightly loger tha carapace; (a) simple, ot bifurcated at tip, (t) ad (sl) abset ; (Gu) preset, (U2) log ad somewhat curved forward, (rsu) 6-8 i umber ; (rsl) 4-6 i umber ; two additioal teeth preset iside of fag furrow o distal portio ; fag log, uarmed. Legs. Leg formula ; all legs with may ad rather short spies, first femora with about 20 spies ; relative legths of legs as show i Table 1. Palpi. Paracymbium log, blutly rouded at distal ed; both coductor ad embolus early straight ad hook-shaped at their tips. Abdome. Log, about 5-6 times as log as broad ; distal ed of abdome somewhat overhagig spierets. Color i alcohol. Cosiderably variable, but geerally light yellowish brow to dark reddish brow ; some specimes with a reddish dorsal markig from head to thorax ad abdome, Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm ; width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet as log as or slightly shorter tha carapace, with small (s) at outer side ear apex ; (Gu) larger tha (UZ), (rsu) 6-7 i umber ; (Gl) ormal, (rsl) 8-10 i umber ; fag uarmed. Legs. Nearly as i male ; relative legths of legs as show i Table 1. Abdome. Similar to male ; geital fold as show i Fig. 1, I. Color i alcohol. Nearly as i male. Specimes examied : NEW GUINEA (IRIAN JAYA) : 19, Fak fak, 1. VI. 1959, J. L. Gressitt (BMH) ; lb, Miwri, 16. II. 1943, T. Niimura (NSMT-Ar. 1086) ; 10, Aggaki, 23. IV. 1943, T. Niimura (NSMT-Ar. 552) ; lb, Aggaki, Yamor, 29. IV. 1943, T. Niimura (NSMT-Ar. 551) ; lb, Siimi, Yamor, 30. IV. 1943, T. Niimura (NSMT-Ar. 555). NEW GUINEA (PNG) : 19, Biake, West Sepik, 16. VII. 1957, D. Elmo Hardy (BMH) ; 10, 20, Brow River, Cetral Prov., 2. IX. 1959, T. Maa (BMH) ;

6 TETRAGNATHA OF THE AUSTRALASIA 41 Fig. 1. Tetragatha rubrivetris Doleschall. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view. D ad E : Distal portio of coductor ad embolus of male. F : Paracymbium of male. G : Left chelicera of female, upper view. H : Ditto, lower view. I : Geital fold of female. J : Eye group of female. K : Abdome of female, lateral view. 19, Cape Rodey, Cetral Prov., 2. XI. 1960, J. L. Gressitt (BMH) ; 19, Cetral Prov., 1. VII. 1962, J. L. Gressitt (BMH) ; 19, Roua m, Cetral Prov., XI N. L. H. Krauss (BMH). AUS- TRALIA : 19, Helevale, N. QLD, XI. 1975, M. Gray (AMS-KS 0419) ; 1Q (AMS-KS 5346) ; (AMS-KS 12830) ; 19, Edmoto, N. Q., 14. IX. 1969, N. C. Colema (AMS-KS 14079) ; 19 (AMS-KS 14095) ; 19 (AMS-KS 14118) ; (AMS-KS 14126) ; 10 19, Gordo Vale, N. Q. (AMS-KS 14127) ; 19, Edmoto, N. Q. (AMS-KS 14139) ; 10, Edmoto, N. Q., 28. VIII. 1976, R. Mascord (AMS-KS 14140) ; 29, Edmoto, N. Q., 16. IX. 1976, R. Mascord (AMS-KS 14141) ; 19, Edmoto, N. Q., 18. IX. 1971, R. Mascord (AMS-KS 14142) ; 19, Edmoto, N. Q., 17. IX. 1971, R. Mascord (AMS-KS 14156) ; lcl (AMS-KS 14209) ; 19, Strat food, Cairs, 20. IV. 1974, Y. Nishikawa (NSMT-Ar 1088) ; 29, NE of Mooroobal Peak, 19. IV. 1974, Y. Nishikawa (NSMT-Ar 1087 & 1089). Distributio : New Guiea ad Australia. Note : This species is related to T. phueodactyh Kulczyski, T. amoea sp. ov. ad T. eumoq%z sp. ov., but it may be easily distiguished from them by the arragemet of eyes, i. e., the lateral eyes more separated tha media eyes (lateral eyes as close as media eyes i phaeodactyla ad others). The shapes of chelicerae i both sexes are also differet from those of the latters. Tetragatha phaeodactyla Kulczyski, 1911 I I Tetragath phaeodactyh Kulczyski, 1911, Nova Guiea, 9 : 115 ; Roewer, 1942, Katalog der

7 42 C. OKUMA Araeae, 1: 987. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head ; posterior row of eyes evidetly arrower tha aterior oe (82-83 : 100) ; viewed from above, two rows early parallel ad recurved ; cetral ocular quadragle with posterior ocular width (measured from outer margis) smaller tha aterior ocular width (84-87 : loo), ad early equal to atero-posterior ocular legth (measured from outer margis) ; ratio of diameter of AME : ALE : PME : PLE as 150 : 75 : 95 : 95 (p). Chelicerae. Log ad sleder, basal segmet somewhat loger tha carapace; (a) small ad simple, ot bifucated at tip ; (Gu) ad (U2) somewhat log, (rsu) 6-8 i umber ; (Gl) preset, (L2) ad (L3) more separated tha (Gl) ad (L2), (rsl) 4-5 i umber ; fag log, uarmed. Legs. Ijured. Palpi. Paracymbium blutly rouded at tip ; coductor somewhat spiraloid ad the tip exteded as a termial hook. Abdome. Log, 4-5 times as log as broad ; distal ed of abdome slightly overhagig spierets. Color i alcohol. Somewhat variable ; legs, carapace, sterum ad mouth parts pale yellowish brow to brow ; legs i some specimes with dusky rigs o distal part of each segmet or irregular flecks o femora ; dorsal abdomial markigs abset. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet about times as log as carapace, somewhat swolle at the middle ; without (s) o the outer side ear apex ; (Gu) as log as or slightly shorter tha (U2), (rsu) 6-9 i umber ; (Gl) very small, (rsl) 6-9 i umber ; fag uarmed. Legs. Leg formula ; all legs with spies, somewhat fewer ad loger tha those of Fig. 2. Tetragathu phaeodactyla Kulczyski. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view. D : Left chelicera of female, upper view. E : Ditto, lower view. F ad G : Distal portio of coductor ad embolus of male. H : Paracymbium of male. I : Geital fold of female. J : Eye group of female. K : Abdome of female, lateral view.

8 TETRAGNATHA OF THE AUSTRALASIA 43 &rive&-is, first femora with lo-16 spies ; relative legths of legs as show i Table 2. Abdome. Similar to male ; geital fold as show i Fig. 2, I. Color i alcohol. Nearly as i male. Specimes examied : NEW GUINEA (IRIAN JAYA) : 19. Fak fak, Bomberi, 6-8. VI. 1959, T. Maa (BMH) ; 16, Orasbar, N. W. Geelyig Bay sealevel, 10. XI. 1963, J. Sedlacek (BMH). NEW GUINEA (PNG) : 19, Bubia, Markham, 50 m, Morobe Prov., 17. IX. 1955, J. L. Gressitt (BMH) ; 16, Daradai, r. Javarere, Mosgrove R. 100 m, C. P., 5. X. 1958, J. L. Gressitt (BMH) ; lb, Bisiaum to Catalia, Cetral Prov., 3. IX. 1959, T. Maa (BMH) ; 26 29, Daradai Platatio, Cetral Prov., 4-7. IX. 1959, T. Maa (BMH) ; 19, Cape Redey, Cetral Prov., 2. Xl. 1960, J. L. Gressitt (BMH) ; 19, Oriomo, West District, 29. X. 1960, J. L. Gressitt (BMH) ; 19, Wisselmere Moaemai, Kamo, 1,500 m, 14. VIII. 1965, J. Sedlacek (BMH) ; 19, Wau l,loo-1,200 m, Morobe Prov., V. 1968, N. L. H. Krauss (BMH). Distributio : New Guiea. Tetragatha amoea sp. av. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head ; posterior row of eyes evidetly arrower tha aterior oe (85 : 100) ; viewed from above, two rows early parallel ad moderately recurved ; cetral ocular quadragle early square,. ratio of diameter of AME : ALE : PME : PLE as 125 : 100: 100 : 100 (p), Chelicerae. Well developed. Basal segmet about 0.7 times as log as carapace ; (s) preset at lower outer side ear apex, very small ; (Gu) ad (U2) early equal i size, close together ear distal ed which are widely separated from (US), (rsu) 6-9 i umber ; (Gl) very small, (L2) very log, widely separated from (Gl), (L3) close to (L2), which are widely separated from (L4), (rsl) 6-7 i umber ; fag log, slightly siuous, uarmed. Legs. Leg formula ; all legs with may ad rather short spies, first femora with about 15 spies; relative legth of legs as show i Table 3. Abdome. Moderately log, about 4 times as log as broad ; geital fold as show i Fig. 3, D ; distal ed of abdome somewhat overhagig spierets. Color i alcohol. Legs pale yellowish brow with dusky rigs ear distal parts of each segmet ; sterum ad maxillae pale yellowish brow ; carapace, chelicerae ad fags light yellowish brow to light brow; abdome pale gray ad covered with may small whitish yellow spagles ; dorsal abdomial markigs abset. Fig. 3. TEdragmath amoea sp. ov. A : Left chelicera of female, upper view. B : Ditto, lower P C: Eye group of female. D : Geital fold of female. E : Abdome of female, lateral view.

9 44 C. OKUMA Type material : Holotype 0 (BISHOP 13631) NEW GUINEA (PNG), Kassam, E. Highlads, l- 9. XI. 1959, T. Maa ad J. L. Gressitt (BMH). Paratypes : NEW GUINEA (PNG) : 19 (12. XII. 1961), 19 (29. X. 1964) ad 19 (23. VI. 1965), Wau 1,200 m, Morobe Prov., J. Sedlacek (BMH) ; 19, Bulolo, Morobe Prov., 24. VIII. 1965, J. Sedlacek (BMH). Type depository : The holotype is preserved i the collectio of BMH. Distributio : New Guiea. Note : This ew species is related to T. mbrivetris Doleschall, T. phaeodactyla Kulczyski ad T. eumo@ha sp. ov., but it may be easily distiguished from them by the characteristic arragemet of teeth o the female chelicera as show i Fig. 3, A & B. Tetragatha eumorpha sp. ov. Male. Body legth, exclusive of chelicerae, 10.5 mm ; carapacial legth mm, width mm ; abdomial legth 7.5 mm, width 1.2 mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head ; posterior row of eyes evidetly arrower tha aterior oe (93 : 100) ; viewed from above, two rows early parallel ad recurved ; cetral ocular quadragle with posterior ocular width smaller tha aterior ocular width (79 : loo), ad smaller tha atero-posterior ocular legth (80 : loo), ratio of diameter of AME : ALE : PME : PLE as 200 : 95 : 125 : 95 (p). Chelicerae. Log ad sleder, basal segmet about 1.4 times as log as carapace ; (a) situated ear apex, small ad simple, ot bifurcated at tip ; (Gu) very log, about 2 times as log as (U2), slightly directed backward, (rsu) 778 i umber ; (Gl) very small, (L2) very log, about equal to (Gu), slightly directed forward, (rsl) 3-4 i umber ; fag log, slightly siuous, uarmed. Legs. Leg formula ; all legs with may ad rather short spies, first femora with about 15 spies ; relative legths of legs as show i Table 4. Palpi. Paracymbium blutly rouded at tip ; both coductor ad embolus log ; coductor bird head-shaped at its distal ed. Abdome. Log, early 6 times as log as broad ; distal ed of abdome slightly overhagig spierets. Color i alcohol. Cosiderably variable ; legs, carapace, sterum ad mouth parts light yellowish brow to dark brow ; abdome, geerally, pale gray ad covered with may small whitish yellow spagles ; abdomial markigs abset. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet as log as or slightly shorter tha carapace, (s) very small, o lower outer side ear apex ; (Gu) robust ad largest, (rsu) S-10 i umber ; (Gl) small, (rsl) 8-10 i umber ; fag log, uarmed. Legs. Nearly as i male ; relative legths of legs as show i Table 4. Abdome. Similar to male ; geital fold as show i Fig. 4, I. Color i alcohol. Nearly as i male. Type material : Holotype 0 (BISHOP 13632) NEW GUINEA (IRIAN JAYA), Holladia-Bie 50 m, 12. VII. 1957, D. Elmo Hardy (BMH). Paratypes : NEW GUINEA (IRIAN JAYA) : 29, same data as holotype ; 19, Cyclops m, 17. IV. 1959, J. L. Gressitt (BMH) ; 19, Oboe Ifar m, J. L. Gressitt (BMH). NEW GUINEA (PNG) : 10 19, Wauma SOO-l,OOOm, Adelbert Mts., Madag Prov., 24, 25. X. 1958, J. L. Gressitt (BMH) ; 19, Agora, Madag Prov., VIII. 1969, Y. Hirashima (KUF) ; 19, Madag, Madag Prov., IX. 1969, Y. Hirashima (KUF).

10 TETRAGNATHA OF THE AUSTRALASIA 45 Fig. 4. Tetragatha eumorpha sp. ov. A : Left chelicera of male, upper view. B : Ditto, lateral view. C: Ditto, lower view. D ad E : Distal portio of coductor ad embolus of male. F : Paracymbium of male. G : Left chelicera of female, upper view. H : Ditto, lower view. I : Geital fold of female. J : Eye group of male. K : Abdome of female, lateral view. Type depository : The holotype is preserved i the collectio of BMH. Distributio : New Guiea. Note : This ew species is related to T. rubrivetris Doleschall, T. phaeodactyla Kulczyski ad T. amoea sp. ov., but it may be easily distiguished from them by the very log (Gu) ad the very log (L2) i male chelicera. The shapes of male coductor ad female geital fold are also differet from those of the latters. The bituberculata-group Diagosis. Posterior row of eyes wider tha aterior row of eyes ; lateral eyes closer tha media eyes, situated o a tubercle ; atero-media eyes closer tha postero-media eyes ; male chelicerae with (a), (t), (sl) ad (T) ; female chelicera with (e) betwee (Gu) ad (T). KEY TO THE SPECIES Male....2 Female Tip of coductor twisted, broadeed...ḋgmissa Tip of coductor twisted, crescet-shaped or very arrowed.... Paracymbium with tip poited, ad subapical margi serrate... ceyloica Paracymbium with tip rouded, ad subapical margi etire... 4 (sl) ad (T) ear apex of upper margi of fag furrow... priamus sp. ov.

11 46 C. OKUMA (~1) ad (T) widely separated from apex of upper margi of fag furrow.... Chelicera about 0.5 times as log as carapace ; (sl) small ad directed forward.-...yalo m Chelicera about times as log as carapace ; (sl) log, ad its tip directed backward... bituberculata (AXl) large ad thick, ad its tip rouded... oreobia sp. ov. (AXl) small or abset... 7 (e) large ad thick, ad its tip rouded... priamfxs sp. ov. (e) simple, ad its tip poited... 8 (Gu) large, ad its tip directed backward ; (e) smaller tha (T)... bitubercuhta (Gu) ot as above ; (e) ot smaller tha (T)... 9 (Gu) larger tha (e), ad its tip directed forward ; small (AXl) preset... ceyloica (Gu) smaller tha (e), ad its tip directed forward ; (AXl) abset...&m&a Tetragatha bituberculata L. Koch, 1867 Tetragatha bituberculata L. Koch, 1867, Verh. Zool.-Bot. Ges. Wie, 17: 184 ; L. Koch, 1871, Arach. Austral., 1: 183 ; Keyserlig, 1887, Arach. Austral., 2 : 220 ; Roewer, 1942, Katalog der Araeae, 1: 985. Tertragatha heatwolei Chrysathus, 1975, Zoo]. Verhadl., 140 : 13. New syoymy. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Viewed from above, posterior row of eyes wider tha aterior row of eyes ( : 100) ; postero-media eyes more separated tha atero-media eyes, ad lateral eyes closer tha media eyes, situated o a tubercle ; cetral ocular quadragle with posterior ocular width larger tha aterior ocular width ( : loo), ad larger tha atero-posterior ocular legth ( : 100) ; ratio of diameter of AME : ALE : PME : PLE as 120 : 63 : 88 : 100 (p). Chelicerae. Basal segmet about times as log as carapace ; (a) bifurcated at tip ; (t), (sl) ad (T) preset, (rsu) 4-7 i umber ; (AXl) ad (Gl) preset, (rsl) 4-7 i umber ; fag uarmed. Legs. Leg formula ; all legs with somewhat log spies, first femora with about 6 spies ; relative legths of legs as show i Table 8. Palpi. Paracymbium rather sleder, blutly rouded at distal ed ; distal ed of coductor of embolus atteuated ad twisted. Abdome. Moderately log about 4 times as log as broad, ot exteded posteriorly to spierets. Color i alcohol. Chelicerae, carapace, sterum, maxillae, labium ad legs brow, fag of chelicerae somewhat dark ; abdome geerally grayish brow, with idistict dusky markigs ad covered with dusky silvery spagles dorsally ; with a cetral dark gray stripe vetrally. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet, times as log as carapace ; (Gu) large, its poit directed somewhat backward, (e) smaller tha (T) ; (rsu) 4-7 i umber ; (AXl) abset ; (rsl) 4-7 i umber ; fag robust, with (EX) o outer side ear base. Legs. Nearly as i male ; relative legths of legs as show i Table 8. Abdome. Moderately log, about 3 times as log as broad ; dorsum i some specimes (full grow?) gibbous at the middle ; ot exteded posteriorly to spierets ; geital fold as show i

12 TETRAGNATHA OF THE AUSTRALASIA \\ AXl._&+W$, EX :* J Fig. 5. Tetragatha bituberculata L. Koch. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view. D : Distal portio of coductor ad embolus of male. E : Paracymbium of male. F : Left chelicera of female, upper view. G : Ditto, lower view. H : Geital fold of female. I : Eye group of male. J : Abdome of female, lateral view. K : Female, dorsal view.

13 48 C. OKUMA Fig. 5, H. Color i alcohol. Geerally paler tha male ; abdome with more cospicuous markigs as show i Fig. 5, J & K. Specimes examied : AUSTRARIA : 10 (AMS-KS 12831) ; 19 (AMS-KS 8909) ; 10 (AMS-KS 9475) ; 20 (AMS-KS 12831); 19 (AMS-KS 14102) ; 20 (AMS-KS 14103); 1Q (AMS-KS 14104) ; 19, Gordo Vale, N. Q. (AMS-KS 14129) ; 10 (AMS-KS 14174); 19 (AMS-KS 14177) ; Id (AMS-KS 14210) ; lq(ams-ks 14215) ; 19, Lizard I., IX. 1967, H. Heatwole (AMS-KS 0025). Distributio : Australia ad eighborig Islads. Note: This species was described from Queeslad ad New South Wales, ad has bee recorded oly from Australia. All the specimes metioed above very well agree with L. Koch s descriptios ad figures. T. heatwolei Chrysathus, 1975 from Lizard I. (holotype, 0, (AMS-KS 0025) ) is idetical with this species. The eyes of T. heatwolei figured by the origial author do ot agree with those of the holotype. Tetragatha ceyloica Cambridge, 1869 Meta gracilis Stoliczka, 1869, Jour. Asiat. Sot. Begal, 38 : 244. Tetragatha ceyloica Cambridge, 1869, Jour. Li. Sot. Zool., 10 : 394 ; Roewer, 1942, Katalog der Araeae, 1: 984 ; Okuma, 1968, Mushi, 42 : 99 ; Chu ad Okuma, 1970, Mushi, 44 : 73. Tetragatha Zat$ros Thorell, 1877, A. Mus. Civ. Geova, 10 : 434. Tetragatha frot0 Thorell, 1890, A. Mus. Civ. Geova, 28 : 214. Tetragatha trides Thorell, 1898, A. Mus. Civ. Geova, 39 : 328. Tetragatka gracilis Pocock, 1900, Fau. Brit. Id. Arach., : 214 ; Meria, 1911, Jahrb. Syst., 31 (2) : 181 ; Gravely, 1921, Rec. Id. Mus., 22 : 427, 436. Tetragatha eitapesis Strad, 1913, Arch. Naturg., 79 : 115 ; Strad, 1914, Abh. Seckebg. Ges., 36 (2) : 196 ; Chrysathus, 1975, Verhadl., 140 : 6. New syoymy. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width 1.4 mm ; cheliceral legth mm. Eyes, Viewed from above, posterior row of eyes wider tha aterior row of eyes ( : 100) ; postero-media eyes more separated tha atero-media eyes ; lateral eyes closer tha media eyes, situated o a tubercle; cetral ocular quadragle with posterior ocular width larger tha aterior ocular width ( : loo), ad larger tha atero-posterior ocular legth ( : 100) ; ratio of diameter of AME : ALE : PME : PLE as 150 : 75 : 125 : 125 (p). Chelicerae. Basal segmet, about times as log as carapace ; (a) obliquely trucate at its tip ; (t), (sl) ad (T) preset, (t) comparatively large, (sl) situated subapical ed of upper margi of fag furrow, (rsu) 5-6 i umber ; (AXl) ad (Gl) preset, (Gl) cosiderably large, (rsl) 4-5 i umber ; fag uarmed. Legs. Leg formula ; all legs with may ad somewhat log spies, first femora with about 10 spies; relative legths of legs as show i Table 9. Palpi. Paracymbium termiated i a uique curved process ad with a serrate margi ear apex (Fig. 6, H) ; distal ed of coductor of embolus atteuated ad twisted. Abdome. Moderately log, about 3.5 times as log as broad, ot exteded posteriorly to spierets. Color i alcohol. Somewhat variable ; carapace, sterum ad chelicerae light brow to dark brow ; legs yellowish brow ; abdome with idistict dusky markigs ad covered with sparse dusky silvery spagles dorsally, with a cetral dark gray stripe vetrally. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width

14 TETRAGNATHA OF THE AUSTRALASIA 49 Fig. 6. Tetragatha ceyloica Cambridge. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view. D : Left chelicera of female, upper view. E : Ditto, lower view. F : Eye group of male. G : Distal portio of coductor ad embolus of male. H : Paracymbium of male. I : Geital fold of female mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet about times as log as carapace ; (Gu) somewhat robust, ad its tip directed forward, (rsu) 6-7 i umber ; (AXl) preset, (rsl) 5-7 i umber ; fag robust, (EX) abset i New Guiea specimes, preset (although very variable i size, ofte oly a mark) i S. E. Asia specimes. Legs. Nearly as i male ; relative legths of legs as show i Table 9. Abdome. Moderately log, about 3 times as log as broad ; ot exteded posteriorly to spierets ; geital fold as show i Fig. 6, I. Color i alcohol. Similar to male but abdome with more cospicuous markigs as usual for the group* Specimes examied. NEW GUINEA (PNG) : 19, Adelbert Mts., Madag Prov., 23. X. 1958, J. L. Gressitt (BMH) ; 19, Wau, 1,200 m, Morobe Prov., 16. XII. 1961, J. H. Sedlacek (Br\/IH). NEW BRITAIN : 10, Nakaai Mts., 28. VII. 1956, J. L. Gressitt (BMH) ; 10, Mt. Kobuga, Bagao, 15. II. 1957, Y. Kodo (BMH). ADMIRALTY Is... 19, Rosse, Maus I., 29. VI. 1959, J. L. Gressitt (BMH) ; lb, Kairiru I. (AMS-KS 7971). Distributio : South East Asia ad New Guiea. Note : This species is related to T. bituberculata L. Koch, T. demissa L. Koch ad T. priamus sp. ov., but it is separable from them by the shape of the male palpal paracymbium, i. e., the presece of the uique curved process o the apex ad the margi ear its apex is serrate. T. eitapesis Strad,

15 50 C. OKUMA 1913 from New Guiea is udoubtedly idetical with this species, ad is syoymized with T. ceyloica i the preset paper. Ad T. crassichelate Chrysathus, 1975 from New Guiea ad T. earmra Levi, 1981 from Florida may be syoyms of this species. Tetragatha demissa L. Koch, Tetragatha demissa L. Koch, , Arach. Austral., 1 : 185 ; Keyserlig, 1887, Arach. Austral., 2 : 221 ; Roewer, 1942, Katalog der Araeae, 1 : 985. Tetragatha quadm des Dodale, 1966, Aust. J. Zool., 14 : 1157 New syoymy. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width l.o- 1.6 mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Viewed from above, posterior row of eyes slightly wider tha aterior row of eyes ( : 100) ; postero-media eyes more separated tha atero-media eyes ; lateral eyes closer tha media eyes, situated o a tubercle ; cetral ocular quadragle with posterior ocular width larger tha aterior ocular width ( : loo), ad larger tha atero-posterior ocular legth i early same ratio as the latter ; ratio of diameter of AME : ALE : PME : PLE as 125 : 63 : 75 : 75 (p). Chelicerae. Basal segmet about times as log as carapace ; (a) bifurcated at tip ; (t), (sl) ad (T) preset, (rsu) 6-9 i umber ; (AXl) ad (Gl) preset, (rsl) 4-7 i umber ; fag uarmed. Legs. Leg formula ; all legs with may spies, first femora with about 11 spies ; relative legths of legs as show i Table 10. Fig. 7. Tetragatha demissa L. Koch. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view. D : Distal portio of coductor ad embolus of male. E ad F : Paracymbium of male. G : Left chelicera of female, upper view. H : Ditto, lower view. I : Abdome of female. lateral view. J : Geital fold of female. K : Eye group of male.

16 TETRAGNATHA OF THE AUSTRALASIA 51 Palpi. Paracymbium rather log, i some specimes with a beaked projectio at its tip, or rouded iwardly at tip ; tip of coductor twisted, broadeed. Abdome, Moderately log, about 4-5 times as log as broad, ot exteded posteriorly to spierets. Color i alcohol. Somewhat variable ; geerally light grayish brow to yellowish brow ; mouth parts somewhat dark ; legs with small flecks at base of each spie ; abdome with idistict dusky markigs dorsally, with a cetral dark gray stripe vetrally. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet about 0.5 times as log as carapace ; (Gu) smaller tha (e), (rsu) 5-6 i umber ; (AXl) abset, (rsl) 5-6 i umber ; fag with (EX) o outer side ear base. Legs. Nearly as i male ; relative legths of legs as show i Table 10. Abdome. Moderately log, about 4 times as log as broad ; ot exteded posteriorly to spierets ; geital fold as show i Fig. 7, J. Color i alcohol. Similar to male, but abdome with more cospicuous markigs as show i Fig. 7, I. Specimes examied : AUSTRALIA: 10 (AMS-KS 6690) ; 19 (AMS-KS 6691) ; 3Q (AMS-KS 14081) ; 26 (AMS-KS 14098) ; 10 2Q (AMS-KS 14105) ; 19 (AMS-KS 14106) ; 10 (AMS-KS 14153) ; ld(ams-ks 14173) ; 19, Queeslad, 20. IV. 1974, Y. Nishikawa (NSMT-Ar. 1090) ; 19, Queeslad, 19. IV. 1974, Y. Nishikawa (NSMT-Ar. 1091). GILBERT & ELLJCE Is. : 1$?,16. V. 1972, P. D. Maser (BMH). TONGA : 19 (other data lost) (BMH) ; Distributio : Australia ad eighborig Islads. Note : This species was described i 1871 from Australia based o a female specime, but i 1887 Keyserlig redescribed based o may specimes i Godeffroy Museum. I 1966, Dodale described a ew species, Tetragatha quadrides, from Australia. However, I have coviced that quadrides is a syoym of demissa. I additio T. margi&a (Thorell, 1890) from S. E. Asia ad T. Zatro Tullgre, 1910 from Africa may be syoymous with T. demissa. This species is related to T. bituberculata L. Koch, 1867 kow also from Australia but is separable from the latter by the male palpal coductor ad paracymbium ad the female (Gu) o the chelicera. The relative legths of legs are also somewhat differet each other (Tables 8 & 10). It is worthwhile to ote that Keyserlig (1887) icorrectly figured the chelicera ad palpus of demissa ad the chelicera of protesa. Namely, the oe show i Taf. XX, Fig. 3 as demissa is protesa, ad the other oe show i Fig. 4 i the same plate as protesa is demissa. Tetragatha oreobia sp. av. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth 3.0 mm, width mm ; abdomial legth mm, width mm ; cheliceral legth 1.9 mm. Eyes. Viewed from above, posterior row of eyes slightly wider tha aterior row of eyes ( : 100) ; postero-media eyes more separated tha atero-media eyes ; lateral eyes closer tha media eyes, situated o a tubercle; cetral ocular quadragle with posterior ocular width larger tha aterior ocular width ( : loo), ad larger tha atero-posterior ocular legth ( : 100) ; ratio of diameter of AME : ALE : PME : PLE as 150 : 50 : 100 : 100 (p). Chelicerae. Basal segmet about 0.6 times as log as carapace, robust, somewhat swolle i the middle, somewhat gibbous o upper-apical outer side ; (AXu) preset, (rsu) 6-8 i umber ; (AXl) very robust, its tip rouded, (rsl) 5-7 i umber; fag robust, uarmed.

17 52 OKUMA Fig. 8. Tetragatha oreobia sp. ov. A : Left chelicera of female, upper view. B : Ditto, lower view. C : Ditto, lateral view. D : Eye group of female. E : Geital fold of female. F : Abdome of female, lateral view. Legs. Leg formula ; all legs with may ad somewhat log spies, first femora with about 7-9 spies ; relative legths of legs as show i Table 11. Abdome. Moderately log, about 4 times as log as broad, ot exteded posteriorly to spierets ; geital fold as show i Fig. 8, E. Color i alcohol. Legs, carapace ad chelicerae yellowish brow ; lip, sterum, fag ad teeth somewhat dark ; abdome covered with may dusky yellowish silvery spagles ad dorsal cardiac area has a trasparet stripe ; veter somewhat dark. Type material : Holotype, 9 (BISHOP 13633) NEW GUINEA (PNG), Kudiawa, 1,400-1,500 m, Chimbu Prov., XI. 1971, N. L. H. Krauss (BMH). Paratype : NEW GUINEA (PNG), 19, Wau, l,loo- 1,200 m, Morobe Prov., V. 1968, N. L. H. Krauss (BMH). Type depository : The holotype is preserved i the collectio of BMH. Distributio : New Guiea. Note : This ew species is related to T. ceyloica Cambridge, T. bitubercukzta L. Koch, T. demissa L. Koch ad T. priamus sp. ov., but it is separable from them i the shape of female chelicera, i. e., a blut promiece situated o the lower-apical portio is very strikig. Tetragatha priamus sp. ov. Male. Body legth, exclusive of chelicerae, 7.6 mm ; carapacial legth 2.2 mm, width 1.3 mm ; abdomial legth 5.4 mm, width 0.9 mm ; cheliceral legth mm. Eyes. Viewed from above, posterior row of eyes slightly wider tha aterior oe ( : 100) ; postero-media eyes more separated tha atero-media eyes ; lateral eyes closer tha media eyes, situated o a tubercle ; cetral ocular quadragle with posterior ocular width larger tha aterior ocular width (131 : loo), ad larger tha atero-posterior ocular legth i early same ratio as the latter ; ratio of diameter of AME : ALE : PME : PLE as 125 : 55 : 125 : 125 (~1). Chelicerae. Cospicuously developed, basal segmet about 0.8 times as log as carapace ; (a) robust, obliquely trucate at tip ; (sl) ad (T) cotiguosly situated ear apex of upper margi of fag furrow, (t) preset betwee (a) ad (T) ; (rsu) 5-6 i umber ; (AXl) preset, (Gl) cosiderably log, (rsl) 4-5 i umber; fag slightly siuous, uarmed. Legs. Leg formula ; all legs with may spies, first femora with about 10 spies ; relative legths of legs as show i Table 12.

18 TETRAGNATHA OF THE AUSTRALASIA &GU e a L Fig. 9. Tetragatha priamus sp. ov. A ad B : Left chelicera of male, upper view. C : Ditto, lateral view. D : Ditto, lower view. E : Distal portio of coductor ad embolus of male. F : Paracymbium of male, G : Left chelicera of female, ier view. H : Ditto, upper view. I : Ditto, lateral view. J : Ditto, lower view. K : Geital fold of female. L : Eye group of male. M : Abdome of female, lateral view.

19 54 C. OKUMA Palpi. Paracymbium ot acumiated at tip ; coductor of embolus atteuated ad twisted at tip. Abdome. Moderately log, about 6 times as log as broad, ot exteded posteriorly to spierets. Color i alcohol. Carapace ad chelicerae brow ; legs, sterum, maxillae ad lip yellowish brow ; abdome geerally pale grayish brow, with idistict dusky markigs ad covered with dusky silvery spagles dorsally, with a cetral dark gray stripe vetrally. Female. Body legth, exclusive of chelicerae, mm, carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet about 0.7 times as log as carapace ; basal segmet large, robust, somewhat swolle i the middle, somewhat gibbous o subapical lower side ad outer side ; (Gu) ad (T) early equal i size, (Gu) directed somewhat backward, (e) robust, its tip rouded, (rsu) 7-10 i umber ; (AXl) preset, (rsl) 6-7 i umber ; fag uarmed. Legs. Nearly as i male ; relative legths of legs as show i Table 12. Abdome. Moderately log, about 4 times as log as broad, ot exteded posteriorly to spierets ; geital fold as show i Fig. 9, K. group. Color i alcohol. Similar to male but abdome with more cospicuous markigs as usual for the Type material : Holotype, 0 (BISHOP 13634) Solomo Is., Kudurumbagara 60 m, 15. XI. 1963, J. L. Gressitt. Paratypes : SOLOMON Is. : 19, Bougaiville (S.), 12. VI. 1956, E. J. Ford, Jr. ; 29, Kolombagara I., VII. 1959, J. L. Gressitt ; 19, Buka I., XII. 1959, T. Maa ; 19, Kudurumbagara 80 m, P. Shaaha ; Id, Kolombagara I., 15. I. 1964, P. Shaaha (all BMH). Type depository : The holotype is preserved i BMH. Distributio : Solomo Is. Note : This ew species is related to T. ceyloica Cambridge, T. bituberculata L. Koch, ad T. demissa L. Koch, but it is separable from them by the shape of male chelicera, particular-y the (~1) ad (T) which are early coalescet at the base i this species. Ad the female chelicera is provided with (e) which is robust, ad rouded at tip i this species. Tetragatha yalom Chrysathus, 1975 Tetragath yalom Chrysathus, 1975, Verhadl., 140 : 18. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width l.l- 1.2 mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Viewed from above, posterior row of eyes slightly wider tha aterior row of eyes ( : 100) ; postero-media eyes more separated tha atero-media eyes ; lateral eyes closer tha media eyes, situated o a tubercle ; cetral ocular quadragle with posterior ocular width larger tha aterior ocular width (128 : loo), ad larger tha atero-posterior ocular legth i early same ratio as the latter ; ratio of diameter of AME : ALE : PME : PLE as 100 : 50 : 75 : 75 (,Y). Chelicerae. Basal segmet about 0.5 times as log as carapace, obliquely trucate at apex ; (a), (t), (~1) ad (T) preset, (rsu) 4 i umber ; (AXl) abset, (Gl) somewhat log, (rsu) 4-6 i umber ; fag uarmed. Legs. Leg formula ; all legs with may spies, first femora with 7-8 spies ; relative legths of legs as show i Table 13. Palpi. Paracymbium rather robust, blutly rouded at tip ; distal ed of coductor similar to that of T. ceyloica or T. bituberculata. Abdome. Moderately log, about 4 times as log as broad, ot exteded posteriorly to

20 --l_m_-lli_li1l---i ll l_---_l _ TETRAGNATHA OF THE AUSTRALASIA b D E 55 Fig. 10. Tetragathu yalom Chrysathus. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view. D : Distal portio of coductor ad embolus of male. E : Paracymbium of male. F : Eye group of male. G : Abdome of male, lateral view. spierets. Color i alcohol. Carapace, chelicerae ad legs yellowish brow ; maxillae, labium ad sterum brow; abdome greyish yellow with some darker irregular logitudial lies; uderside greyish brow; veter with a cetral dark gray stripe. Female ukow. Specimes examied : NEW GUINEA (PNG) : lb, J. L. Gressitt (BMH) ; 10, Get River, 7. VIII. 1983, 0. Tadauchi (KUF). AUSTRALIA : Id, N. QLD, XI. 1975, M. Gray (AMS-KS 0420). Distributio : New Guiea ad Australia. Note : This species is related to T. ceyloica Cambridge ad T. bituberculata L. Koch, but it is separable from them by the shape of male chelicera, i. e., the basal segmet obliquely trucated i this species as show i Fig. 10, A & C. The jaculator-group Diagosis. Posterior row of eyes evidetly arrower tha aterior row of eyes ; lateral eyes more separated tha media eyes ; cheliceral legth shorter tha carapace i both sexes ; distal ed of abdome slightly overhagig spierets. Tetragatha jaculator Tullgre, 1910 Tetragatha jacukztor Tullgre, 1910, Sjijstedt Kilimad. Exped., 3: 150 ; Lawrece, 1936, A. Trasv. Mus., 17 (2) : 152 ; Roewer, 1942, Katalog der Araeae, 1: 978 ; Okuma, 1984, Esakia, (22) : 87. Tetragathu sp. B. Okuma, 1968, Mushi, 42 : 103 ; Okuma, 1981, Jap. J. Appl. Et. Zool., 25 : 297. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width l.o- 1.4 mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head ; posterior row of eyes somewhat arrower tha aterior oe (90-97 : 100) ; viewed from above aterior row moderately recurved, posterior row somewhat strogly recurved, so that lateral eyes more separated tha media eyes; cetral ocular quadragle with posterior ocular width larger tha aterior ocular width (123 : 100) ad slightly larger tha atero-posterior ocular legth (107 : 100) ; ratio of diameter of AME : ALE : PME : PLE as 113 : 55 : 75 : 75 (p).

21 56 C. OKUMA Chelicerae. Basal segmet, about 0.6 times as log as carapace ; (AXu), (a), (t), (sl) ad (T) preset, (AXu) variable i size, (rsu) 4-5 i umber ; (AXl) preset, (Gl) somewhat log, (rsl) 6-8 i umber ; fag uarmed. Legs. Leg formula ; all legs with spies, first femora with about 6-9 spies ; relative legths of legs as show i Table 5. Palpi. Paracymbium somewhat cubical ad complicated as show i Fig. 11, G ; distal ed of coductor ad embolus simple ; spherical tegulum distictly large, oval ad slatig. Abdome. Moderately log, about 4 times as log as broad ; distal ed of abdome somewhat overhagig spierets. Color i alcohol. Legs, carapace ad chelicerae geerally yellowish brow ; maxillae, lip, ad sterum somewhat dusky brow ; abdome geerally gray, covered with yellowish silvery spagles dorsally, ad gray vetrally. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width 1.L 1.5 mm ; abdomial legth mm ; width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet about 0.5 times as log as carapace ; (AXu) preset, (rsu) 4-5 i umber ; (rsl) 6-7 i umber ; fag uarmed. Legs. Nearly as i male ; relative legths of legs as show i Table 5. Abdome. Moderately log, about 3.5 times as log as broad ; distal ed of abdome slightly AXa Fig. 11. Tetmgathu jaculutor Tullgre. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view. D ad E: Left palpus of male. F : Distal portio of coductor ad embolus of male. G : Paracymbium of male. H : Left chelicera of female, upper view. I : Ditto, lower view. J : Geital fold of female. K : Eye group of female. L : Abdome of female, lateral view.

22 TETRAGNATHA OF THE AUSTRALASIA 57 overhagig spierets; geital fold as show i Fig. 11, J. Color i alcohol. Similar to male, but abdome with silvery lateral stripes as show i Fig. 11, L. Specimes examied : NEW GUINEA (IRIAN JAYA) : 26 29, Vogelkop, Sururai Vil. 1 area, W. shore Lake Agi Giji, 1,850 m, 26. VII. 1957, D. Elmo Hardy (BMH) ; 16, Vogelkop, Sucumi Camp, r. head of Sucumi R., 6. VIII 1975, D. Elmo Hardy (BMH) ; 20 19, Earatoli, 4. VII, 1962, N. Wilso (BMH) : NEW GUINEA (PNG) : , Myola 2,000-2,800 m, Cetral Prov., 1-5. XI. 1984, Y. Hirashima (KUF). Distributio : Africa, S. E. Asia ad New Guiea (ew record). Note : This species was described from Kilimajaro, Africa. A careful compariso of the above metioed specimes from New Guiea with a male specime from Africa ad Tullgre s descriptio ad figures has coviced me that they belog to the oe species, T. jaculator. Tetragatha sp. B (Okuma, 1968 ad 1981) from S. E. Asia is udoubtedly idetical with this species. Ad T. argyroides Mello-Leitao, 1945 from Argetia may be a syoym of this species. The caudifera-group Diagosis. Posterior row of eyes evidetly arrower tha aterior row of eyes ; lateral eyes more separated tha media eyes; chelicera shorter tha carapace i both sexes; a small tail preset behid spierets; male chelicera with (a), (AXu), (sl) ad (T). Tetragatha caudifera (Keyserlig, 1887) Eucta caudifera Keyserlig, 1887, Arach. Austral., 2 : 225 ; Roewer, 1942, Katalog der Araeae, 1 : 973. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width 1.9 mm ; abdomial legth mm, width 1.6 mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head ; posterior row of eyes slightly arrower tha aterior oe (88-93 : 100) ; viewed from above aterior row moderately recurved, posterior row somewhat strogly recurved, so that lateral eyes more saparated tha media eyes; cetral ocular quadragle with posterior ocular width larger tha aterior ocular width (130 : loo), ad larger tha atero-posterior ocular legth (115 : 110) ; ratio of diameter of AME : ALE: PME : PLE as 113 : 75 : 100 : 100 (/J). Chelicerae. Basal segmet about 0.7 times as log as carapace ; (a) bifurcated at tip ; (AXu), (sl) ad (T) preset, (rsu) 5-6 i umber ; (AXl) preset, (Gl) cosiderably log, (rsl) 6-7 i umber ; fag uarmed. Legs. Leg formula ; all legs with spies, first femora with about 5-9 spies ; relative legths of legs as show i Table 6. Palpi. Paracymbium somewhat bifurcated at tip,. coductor ad embolus simple. Abdome. Moderately log, 3-4 times as log as broad, with a small tail behid spierets. Color i alcohol. Legs, carapace ad chelicerae geerally yellowish or light brow ; maxillae, lip ad sterum somewhat dusky brow ; abdome geerally yellowish brow, covered with dusky silvery spagles dorsally, ad with a cetral dark gray stripe vetrally. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width 1.7 mm ; abdomial legth mm, width mm, cheliceral legth 1.7 mm. Eyes. Nearly as i male. Chelicerae. Basal segmet about 0.6 times as log as carapace ; (Gu), (e) ad (T) preset, (rsu)

23 58 C. OKUMA Fig. 12. Tetragathu caudifera (Keyserlig). A : Left chelicera of male, upper view. B : Ditto, lateral view, C : Ditto, lower view. D : Left chelicera of female, upper view. E : Ditto, lateral view. F : Ditto, lower view. G ad H : Distal portio of coductor ad embolus of male. I : Paracymbium of male. J : Geital fold of female. K : Eye group of female. L : Abdome of female, lateral view. 6-7 i umber ; (AXl) ad (Gl) preset, (Gl) robust ad largest, (rsl) 6-7 i umber ; fag uarmed. Legs. Nearly as i male, * relative legths of legs as show i Table 6. Abdome. Moderately log, more tha 4 times as log as broad, with a small tail behid spierets ; geital fold as show i Fig. 12, J. Color i alcohol. Similar to male, but abdome with silvery lateral stripes as show i Fig. 12, L. Specimes exmied : AUSTRALIA : 16, W. of Welshpool, H. Way (AMS-KS 9088) ; 10, W. of Welshpool, H. Way (AMS-KS 9098) ; 29, N. S. W., I. 1936, K. C. Mckoow (AMS-KS 14165). Distributio : Australia. Note : This species was described from New South Wales, i Sice the, o record has bee made. The preset specimes well agree with Keyserlig s descriptios ad figures. This species is readily separated from other Australia Tetragatha by the shorter abdome (about 3-4 times as log as broad) ad the presece of the tail behid the spierets; This species is similar to T. caudicula (Karsch, 1879) from Japa ad T. caudata Emerto, 1884 from America by the shape of abdome, but it may be easily distiguished from them by the simple coductor of the male (distictly modified i caudicula ad caudata). The shape of chelicerae of both sexes is also differet from the latter-s. The micrura-group Diagosis. Posterior row of eyes evidetly arrower tha aterior row of eyes ; lateral eyes more

24 TETRAGNATHA OF THE AUSTRALASIA 59 separated tha media eyes; chelicera shorter tha carapace i both sexes ; male chelicera without (0, (AXu), (Gu) ad (sl) ; abdome elogate, with a small tail behid spierets i both sexes. Tetragatha micrura Kulczyski, 1911 Tetragath micmra Kulczyski, 1911, Nova Guiea, 5 (Zool. 3) : 450 ; Roewer, 1942, Katalog der Araeae, 1 : 987 ; Chrysathus, 1975, Verhadl., 140 : 6. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width 1.0 mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head ; posterior row of eyes evidetly arrower tha aterior oe (77-78 : 100) ; viewed from above, aterior row moderately recurved, posterior row somewhat strogly recurved, so that lateral eyes more separated tha media eyes ; cetral ocular quadragle early square ; ratio of diameter of AME : ALE : PME : PLE as 175 : 113 : 113 : 113 (/1). Chelicerae. Basal segmet about 0.8 times as log as carapace ; (t), (AXu), (Gu) ad (sl) abset, (a) obliquely trucate at tip ; (T) moderately log, (rsu) 6-7 i umber ; (AXI) abset, (Gl) large ad robust, (rsl) 9-10 i umber; fag with a small ier cusp ear base. Legs. Leg formula ; all legs with may ad rather short spies, first femora with about 20 spies; relative legths of legs as show i Table 7. Palpi. Paracymbium relatively short ad broad except termial ; most part of coductor somewhat broad ad straight, the tip small, twisted. Abdome, Elogate, 9-12 times as log as broad, with a small tail behid spierets. Color i alcohol. Geerally yellowish pale brow ; fags ad teeth reddish brow ; abdome covered with yellowish silvery spagles dorsally ; i some specimes abdome with pale grayish dots dorso-laterally. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet about 0.5 times as log as carapace ; (Gu) small, (e) ad (T) subequal i size, (rsu) 4-5 i umber ; (Gl) preset, (rsl) 7-10 i umber ; fag uarmed. Legs. Nearly as i male ; relative legths of legs as show i Table 7. Abdome. Very log, more tha 8 times as log as broad, with a small tail behid spierets ; geital fold as show i Fig. 13, I. Color i alcohol. Similar to male. Specimes exmied. NEW GUINEA (PNG) : 19, VII-VIII. 1959, T. C. Maa (BMH) ; 19, Sogeri, Cetral Prov., 24. VIII. 1982, Y. Hirashima (KUF). NEW BRITAIN : Id, Gazelle Peisula, 9. IX. 1955, J. L. Gressitt (BMH) ; 10, Keravat, VI. 1959, T. Maa (BMH) ; 19, Gazelle Peisula, X. 1962, J. Sedlacek (BMH) ; 19, Gazelle Peisula, Mt. Siewit 900 m, XI. 1962, J. Sedlacek (BMH). SOLOMON Is... lb, Vella Lavella, Kudurumbagara 80 m, P. Shaaha (BMH) ; 19, Big Nggela, C. W. O Brie (BMH). BUKA I... 19, Ag. Statio, XII. 1959, T. Maa (BMH). NEW IRELAND : 19, Ridge above Camp Bishop 20 km up m, 8. VII. 1956, J. L. Gressitt (BMH). Distributio : New Guiea ad eighborig Islads. Note : This species is readily separated from all other species i the geus by the elogate abdome edig i a small tail, ad the male chelicera lackig (t), (AXu), (Gu) ad (~1). The female is recorded ad described for the first time.

25 60 C. OKUMA Fig. 13. Tetragathu micrwa Kulczyski. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view. D : Left chelicera of female, upper view. E : Ditto, lower view. F : Eye group of male. G : Paracymbium of male. H : Distal portio of coductor ad embolus of male. I : Geital fold of female. J : Abdome of male, lateral view. The cylidrica-group Diagosis. Aterior ad posterior rows of eyes equal or subequal i width; abdome very log, about 6-10 times as log as broad, tail abset. KEY TO THE SPECIES Male....2 Female.... f helicera with (a), (t), (~1) ad (T)....3 Chelicera without (t)... 4 Lateral eyes more separated tha media eyes, (sl) directed forward; (LZ) directed backward...cyl&dric a Lateral eyes as close as media eyes ; (sl) directed backward ; (L2) directed forward..a...ṃcileta Paracymbium sleder, poited at tip ; distal ed of coductor bird head-shaped......aa sp. ova Paracymbium ot sleder, ot poited at tip... 5 Paracymbium bifurcated at tip... isularis sp. ov. Paracymbium rouded at tip... biseriata Fag with (EX) ad a ier cusp.... Fag uarmed... 8 Chelicera shorter tha oe-half of carapace ; (EX) situated o upper side gyessittorum sp. av.

26 _. - l---_ I - l l_i._- _ I* ~ _. TETRAGNATHA OF THE AUSTRALASIA Chelicera about 0.6 times as log as carapace ; (EX) situated o lower side chauziodus Chelicera with (s) ear outer apex bis&ata Chelicera without (s) , (Gl) rouded at tip; lower margi of fag furrow with a remarkable promiece subbasally vacile ta (Gl) poited at tip ; fag furrow usually without above promiece Lateral eyes more separated tha media eyes cylidm ca Lateral eyes as close as media eyes Geital fold somewhat log, ratio of width ad legth as about 1: aa sp. ova Geital fold somewhat short, ratio of width ad legth as about 2 : 1 ~--~-~~--~~-isuzaris sp. ov. Tetragatha cylidrica Walckeaer, 1841 Tetragatha cylidrica Walckeaer, 1841, Hist. Nat. Is. Apt., 2 : 210 ; Keyserlig, 1865, Verh. Zool.-Bot. Ges. Wie, 15 : 842 ; L. Koch, 1871, Arach. Austral., 1 (1) : 181 ; Roewer, 1942, Katalog der Araeae, 1: 985. Eugathu cylidracea Keyserlig, 1887, Arach. Austral., 2 : 224. Eugatha cylidrica Keyserlig, 1887, Arach. Austral., 2 : 226. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head ; posterior row of eyes slightly arrower tha aterior oe (93-98 : 100) ; viewed from above, aterior row moderately recurved, posterior row somewhat strogly recurved, so that lateral eyes more separated tha media eyes ; cetral ocular quadragle early square,. ratio of diameter of AME : ALE : PME : PLE as 95 : 58 : 70 : 70 (p). Chelicerae. Basal segmet about times as log as carapace ; (a) bifurcated at tip ; (t), (sl) ad (T) preset, (sl) directed forward, (rsu) 4-5 i umber ; (Gl) slightly directed forward, (L2) directed backward, (rsl) 4-5 i umber ; fag uarmed. Legs. Leg formula ; all legs with several spies, first femora with about 5 spies ; relative legths of legs as show i Table 14. Palpi. Paracymbium rather sleder, moderately curved ad somewhat poited at tip ; coductor ad embolus subspiral at the middle. Abdome. Very log, about 8 times as log as broad, ot cotiued behid to spierets. Color i alcohol. Legs, carapace ad chelicerae geerally yellowish brow ; maxillae, lip, ad sterum somewhat dusky brow ; abdome geerally pale gray, covered with yellowish silvery spagles dorsally, somewhat dusky brow vetrally. Female. Body legth, exclusive of chelicerae, 10.2 mm ; carapacial legth 2.3 mm, width 1.1 mm ; abdomial legth 8.0 mm, width 1.0 mm ; cheliceral legth 0.9 mm. Eyes. Nearly as i male. Chelicerae. Basal segmet about 0.4 times as log as carapace ; (Gu) ad (T) preset, (rsu) 4 i umber ; (AXl) abset, (Gl) preset, (rsl) 4 i umber ; fag uamed. Legs. Nearly as i male ; relative legths of legs as show i Table 14. Abdome. Very log about 8 times as log as broad, ot cotiued behid to spierets ; geital fold as show i Fig. 14, I. Color i alcohol. Similar to male, but abdome with silvery lateral stripes as show i Fig. 14, K. Specimes examied : NEW GUINEA (PNG) : lb, Brow River, Cetral Prov., 21. V. 1956, J. Ford, Jr. (BMH). AUSTRALIA : 20, Mout Garet, N. Q., 23. II. 1972, N. C. Colema (AMS-KS ad 14161) ; 19, Bellagray, SE NSW (AMS-KS 9690).

27 62 C. OKUMA L-d D u J Fig. 14. Tetragatha cylidtica Walckeaer. A : Left chelicera of male, upper view. B : Ditto, lateral view. C: Ditto, lower view. D : Left palpus of male. E : Distal portio of coductor ad embolus of male. F : Paracymbium of male. G : Left chelicera of female, upper view. H : Ditto, lower view. I : Geital fold of female. J : Carapace of male. K : Abdome of female, lateral view. Distributio : New Guiea, Australia, Fiji ad Lord Howe Is. Note : This species was described from New Guiea ad has bee recorded from Australia ad eighborig islads. The preset specimes well agree with the origial descriptio ad figures. This species is related to T. macileta L. Koch ad T. biseriata Thorell, but it may be easily distiguished from them by the arragemet of eyes, i. e., lateral eyes more separated tha media eyes. Ad also the shape of male palpus is evidetly differet from those of macilata ad biseriata. Tetragatha chauliodus (Thorell, 1890) Limoxere chauliodus Thorell, 1890, A. Mus. Civ. Geeva, 30 : 292. Tetragatha chauliodus Gravely, 1921, Rec. Id. Mus., 22 : 425 ; Roewer, 1942, Katalog der Araeae, 1 : 981. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head, posterior row of eyes about as wide as aterior oe ; viewed from above, two rows early parallel ad moderately recurved ; cetral ocular quadragle with posterior ocular width slightly larger tha aterior ocular width ( : loo), ad slightly larger tha atero-posterior ocular legth i early same ratio as the latter; ratio of diameter of AME : ALE : PME : PLE as 125 : 93 : 100 : 100 (p). Chelicerae. Basal segmet about 0.6 times as log as carapace ; (Gu) ad (T) preset, (rsu) 5-6 i umber ; (AXl) abset, (Gl) preset, (rsl) 7-9 i umber ; fag robust, armed with (EX) o lower side, ad log ier tooth at about l/3 from base.

28 TETRAGA ATHA OF THE AUSTRALASIA 63 Legs. Leg formula ; all legs with a few spies, first femora with l-3 spies ; relative legths of legs as show i Table 15. Abdome. Very log, about times as log as broad, ot exteded posteriorly to spierets; geital fold as show i Fig. 15, D. Color i alcohol. Geerally light brow ; abdome light gray, with dusky spots dorso-laterally. Specimes examied : NEW GUINEA : 20, Holladia-Bie 50 m, 12. VII. 1957, D. Elmo Hardy (BMH). Distributio: Burma, Malaysia, Sigapore ad New Guiea (ew record). Note: This species is related to T. macileta L. Koch ad T. biseriata Thorell, but it may be easily distiguished from them by the female cheliceral fag which is armed with (EX) ad a additioal ier tooth (female fag of ma&&a ad that of biseriata uarmed). - I._ - -\ JE Fig. 15. Tetragatha chuuliodus (Thorell). A : Left chelicera of female, upper view. B : Ditto, lower view. C : Eye group of female. D : Geital fold of female. E : Abdome of female, lateral view. Tetragatha macileta L. Koch, 1871 Tetragatha macile ta L. Koch, 1871, Arach. Austral., l(1) : 192 ; Berlad, 1929, Isect Samoa, 8(2) : 60 ; Roewer, 1942, Katalog der Araeae, 1: 986 ; Chrysathus, 1975, Zool. Verhadl., 140 : 10. Male. Body legth, exclusive of chelicerae, mm,. carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head, posterior row of eyes about as wide as aterior oe ; viewed from above, two rows early parallel ad moderately recurved ; cetral ocular quadragle early square,* ratio of diameter of AME : ALE : PME : PLE as 105 : 63 : 88 : 63 (p). bifurcated at tip ; (t) small ; (sl) situated ear distal ed of upper margi of fag furrow, directed backward ; (T) somewhat robust, (rsu) 4-5 i umber ; (Gl) ad (L2) somewhat robust, (rsl) 5-6 i umber ; fag slightly siuous, uarmed. Legs. Leg formula ; all legs with oly a few spies, first femora with l-2 spies ; relative legths of legs as show i Table 16. Abdome. Very log, about 10 times as log as broad, ot exteded posteriorly to spierets. Color i alcohol. Chelicerae, carapace ad legs yellowish brow, labium ad sterum somewhat

29 64 C. OKUMA dusky brow ; abdome geerally grayish brow, covered with small yellowish silvery spagles dorsally, grayish brow vetrally. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet about times as log as carapace ; (Gu) acumiated at tip, (rsu) 5-7 i umber; (Gl) variable i size, but its tip usually blutly rouded, (rsl) 7-9 i umber, lower margi of fag furrow with a remarkable promiece subbasally ; fag slightly siuous, uarmed. Legs. Nearly as i male ; relative legths of legs as show i Table 16. Abdome. Very log, about 7 times as log as broad, ot exteded posteriorly to spierets ; geital fold as show i Fig. 16, I. Color i alcohol. Similar to male. Specimes examied : NEW GUINEA (PNG) : 19, Mij 1,700 m, 1. VII. 1975, D. Elmo Hardy (BMH) ; 19, W. Highlads, Goiburug, E. of Kor Farm 1,560-1,650 m, 16. X. 1958, J. L. Gressitt (BMH) ; 19, Bisiaumu, 3. IX. 1959, T. Maa (BMH) ; 16 19, E. Highlads, Kassam, l-9. XI. 1959, T. Maa & J. L. Gressitt (BMH) ; 19, Goroka 1,550 m, 10. VI. 1965, J. L. Gressitt (BMH). ADMIRALTY Is. : 20, Maus I., Loregau, 27. VI. 1959, J. L. Gressitt (BMH); 19, Maus I., 29. VI. 1959, J. L. Gressitt (BMH). SOLOMON Is. : 19, Malaita, 21. IX. 1957, J. L. Gressitt (BMH) ; 29, Muda, 20. VII. 1959, J. L. Gressitt (BMH) ; 19 Guadalcaal I., 15. V. 1960, C. W. O Brie (BMH) ; 19, Guadalcaal I., 5. V. 1964, J. H. Sedlacek (BMH) ; 49, Sata Isabel, Buala, O-100 m, XII. 1971, N. L. H. Krauss (BMH). SAMOA : 19, Upolu I., II. 1955, N. L. H. Krauss (BMH). TONGA: 10 19, Eua I. (BMH) ; Id 29, VaVau I. (BMH) ; 19, Nuku Alofa I. FIJI : Id, Viti Levu I. (BMH). NEW BRITAIN : 30 2?,28,29. Fig. 16. Tetragatha macileziz L. Koch. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view, D : Eye group of male. E : Distal portio of coductor ad embolus of male. F : Paracymbium of male. G : Left chelicera of female, upper view. H : Ditto, lower view. I : Geital fold of female. J : Abdome of female, lateral view.

30 TETRAGNATHA OF THE AUSTRALASIA 65 X. 1984, Y. Hirashima (KUF). AUSTRALIA : I@ 19, Queeslad, 20. IV. 1974, Y. Nishikawa (NSMT- Ar. 553). Distributio : Norfolk I., Samoa, Mariaas, Toga, Hawaii, Marquesas I., Solomo Is., New Guiea ad Australia. Note : This species is related to T. cylidtica Walckeaer ad 2. biseriata Thorell, but it may be easily distiguished from them by the shape of chelicera i both sexes. Ad the shapes of male palpal coductor ad embolus are also differet from those of cylidrica ad biseriata. Te tragrta tha gressi t torum sp. ov. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm ; width mm ; abdomial legth mm, width 1.0 mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head, posterior row of eyes about as wide as aterior oe ; viewed from above, two rows early parallel ad moderately recurved ; cetral ocular quadragle with posterior ocular width slightly larger tha aterior ocular width (118 : loo), ad slightly larger tha atero-posterior ocular legth (113 : 100) ; ratio of diameter of AME : ALE : PME : PLE as 88 : 63 : 88 : 75 (p). Chelicerae. Basal segmet about 0.4 times as log as carapace ; (Gu), ad (T) preset, (rsu) 4-5 i umber; (Gl) preset, (rsl) 3-4 i umber ad additioal miute deticles preset o its basal margi ; fag armed with (EX) o upper side, ad a small ier promiece preset at about oe-half from base. Legs. Leg formula ; all legs without spies except a fait spie o distal ed of each patella ; relative legths of legs as show i Table 17. Abdome. Log, about times as log as broad, ot exteded posteriorly to spierets ; geital fold as show i Fig. 17, E. Color i alcohol. Legs, carapace ad chelicerae geerally light yellowish brow ; labium ad sterum somewhat dark ; abdome geerally pale grayish brow, covered with small silvery spagles dorsally. Type material : Holotype 0 (BISHOP 13635), NEW GUINEA (IRIAN JAYA), Earatoli, 1,800-1,900 m, 26. VII. 1962, J. Sedlacek (BMH). Paratype : 19, Earatoli, 1,800 m, 2. VIII. 1955, J. L. Gressitt (BMH). Type depository : The holotype is preserved i the collectio of the BMH. Distributio : New Guiea (Iria Jaya). Note : This ew species is related to T. biseriata Thorell, T. isularis sp. ov. ad T. aa sp. ov. but it may be easily distiguished from them by the fag armature. The species is amed i hoor of the late Dr. ad Mrs. J. L. Gressitt. Fig. 17. Tetragatha gressittomm sp. ov. A : Left chelicera of female, upper view. B : Ditto, lower view. C : Abdome of female, lateral view. D : Eye group of female. E : Geital fold of female.

31 66 C. OKUMA Tetragatha biseriata Thorell, 1881 Tetragatha biswiata Thorell, 1881, A. Mus. Civ. Geova, 17 : 139 ; Gravely, 1921, Rec. Id. Mus., 22 : 424 ; Roewer, 1942, Katalog der Araeae, 1 : 985 ; Chrysathus, 1975,Zool. Verhadl., 140 : 4. Tetragatha modica Kulczyski, 1911, Nova Guiea, 5 ( ) : 449. Tetragath airesis Strad, 1914, Abh. Seckebg. Ges., 36 (2) : 196. New syoymy. Tetragatha valoka Chrysathus, 1975, Zool. Verhadl., 140 : 17. New syoymy. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head, posterior row of eyes about as wide as aterior oe ; viewed from above, two rows early parallel ad moderately recurved ; cetral ocular quadragle early square ; ratio of diameter of AME : ALE : PME : PLE as 138 : 75 : 100 : 88 (p). Chelicerae. Basal segmet about 0.7 times as log as carapace ; (a) bifurcated at tip ; (~1) ot slopig, directed forward, (T) moderately large, (rsu) 4-5 i umber ; (Gl) directed forward, (L2) large, directed somewhat backward, (rsl) 5-8 i umber ; fag uarmed. Legs. Leg formula ; all legs with oly a few spies, first femora with about 5 spies ; relative legths of legs as show i Table 19. Palpi. Paracymbium broad, blutly rouded at tip ; both coductor ad embolus getly twisted ; embolus variable i shape (Fig. 18, D & E), ofte the tip broke off durig copulatio (cf. Chrysathus, 1975). Abdome. Very log, about 6-8 times as log as broad, ot exteded posteriorly to spierets. Color i alcohol. Legs, carapace ad chelicerae yellowish brow to light brow ; fag, labium ad sterum somewhat dark ; abdome geerally pale grayish brow, covered with small silvery spagles dorsally, with a cetral dark gray strip vetrally ; i some specimes, abdome with pale blackish logitudial lies dorso-laterally. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Chelicerae. Basal segmet about times as log as carapace ; (s) preset o outer side of subapex ; small obtuse tubercle preset o lower side of subapex ; (Gu) ad (T) preset, (rsu) 4-5 i umber ; (Gl) preset, (rsl) 4-6 i umber ; fag uarmed. Legs. Nearly as i male ; relative legths of legs as show i Table 19. Abdome. Very log, about times as log as broad, ot exteded posteriorly to spierets ; geital fold as show i Fig. 18, I. Color i alcohol. Similar to male but abdome with more cospicuous lies as show i Fig. 18,K. Specimes examied : NEW GUINEA : A total of 490 ad 909, (all BMH). AUSTRALIA : 19 (AMS-KS 0418) ; 20 (AMS-KS 0468) ; 20 (AMS-KS 14094); 19 (AM%KS 14146) ; 19, N. Q., 30. IX. 1971, R. E. Mascord (AMS-KS 14159) ; 19, N. Q., III. 1979, W. D. Campbell (AMS-KS 14172) ; Id 39, N. E. of Mooroobool peak, W. of Cairs, QSL, 19. IV. 1974, Y. Nishikawa (NSMT-Ar. 1093). Distributio : New Guiea, Aru I. ad Australia (ew record). Note : This species seems to be commo i New Guiea. This is related to T. cylidtica Walckeaer ad T. macileta L. Koch, but it may be easily distiguished from them by the female chelicera which is provided with a small (s), ad the shapes of male palpal coductor ad embolus are also differet from those of cylidrica ad macileta. T. airesis Strad, 1914 ad T. valoka Chrysathus, 1975 are udoubtedly idetical with this species, ad are syoymized with biseriata i the preset paper.

32 TETRAGNATHA OF THE AUSTRALASIA 67 Fig. 18. Tetragatha biseriata Thorell. A : Left chelicera of male, upper view. B : Ditto, lateral view, C: Ditto, lower view. D ad E : Distal portio of coductor ad embolus of male. F: Paracymbium of male. G : Left chelicera of female, upper view. H : Ditto, lower view. I : Geital fold of female. J : Eye group of female. K : Abdome of female, lateral view. Tetragatha isularis sp. av. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head, posterior row of eyes about as wide as aterior oe ; viewed from above, two rows early parallel ad moderately recurved ; cetral ocular quadragle early square ; ratio of diameter of AME : ALE : PME : PLE as 113 : 75 : 88 : 88 (p). Chelicerae. Basal segmet slightly loger tha oe-half of carapace ; (a) ot bifurcated at tip ; (sl) directed forward, (T) ot so large, (rsu) 3-4 i umber ; (Gl) directed forward, (L2) directed backward, (rsl) 5 i umber ; fag uarmed. Legs. Leg formula ; all legs without spies except a fait spie at distal ed of patella (abset i some specimes) ; relative legths of legs as show i Table 20. Palpi. Paracymbium broad, slightly curved, largely bifurcated at tip ; both coductor ad embolus twisted as show i Fig. 19, D. Abdome. Very log, about 6-7 times as log as broad, ot exteded posteriorly to spierets. Color i alcohol. Legs, carapace ad chelicerae geerally light yellowish brow ; abdome geerally pale grayish yellow, covered with silvery spagles dorsally, with pale blackish logitudial lies ad spots laterally, ad with a cetral dark gray stripe vetrally. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male.

33 68 C. OKUMA Chelicerae. Basal segmet about 0.3 times as log as carapace ; (Gu), (e) ad (T) preset, (rsu) 4 i umber ; (Gl) preset, (rsl) 4 i umber ; fag uarmed. Legs. Nearly as i male ; relative legths of legs as show i Table 20. Abdome. Very log, about 6-9 times as log as broad, ot exteded posteriorly to spierets ; geital fold as show i Fig. 19, I. Color i alcohol, Similar to male but abdome i some specimes with silvery stripe laterally as usual for the group. Type material : Holotype 10 (AMS-KS 14025) Lord Howe Is., Grass m. Paratypes : 30 20, same data as holotype ; (AMS-KS 14190) ; lb CAMS-KS 14191) ; 19 CAMS-KS 14194) ; 19 (AMS-KS 14195) ; 20= (AMS-KS 14196) ; 1019 (AMS-KS 14197) ; lb CAMS-KS 14199) ; Id (AMS- KS 14201) ; 19 (AMS-KS 14202) ; 19 (AMS-KS 14203) all specimes collected from Lord Howe Is. Type depository : The holotype is preserved i the collectio of AMS. Distributio : Lord Howe Is. Note : This ew species is related to T. bisem ata Thorell but it may be easily distiguished from the latter by the characteristic male palpal coductor ad embolus as show i Fig. 19, D. Tetrugmztha aa sp. ov. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width 0.8 mm ; abdomial legth mm, width 0.55 mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head, posterior row of eyes about as wide as aterior oe ; viewed from above, two rows early parallel ad somewhat strogly recurved ; cetral ocular quadragle with posterior ocular width slightly larger tha aterior ocular width ( : loo), ad early equal to atero-posterior ocular legth ; ratio of diameter of AME : ALE : PME : PLE as 100 : 55 : 100 : 75 (p). Chelicerae. Basal segmet slightly loger tha oe-half of carapace ; (a) ot bifurcated at tip ; (Gu) preset, (sl) ad (T) early equal i size, slightly directed forward, (rsu) 3 i umber ; (Gl) somewhat robust, (rsl) 4 i umber; fag uarmed. Legs. Leg formula ; all legs without spies ; relative legths of legs as show i Table 18. Fig. 19. Tetrag~atka isularis sp. ov. A : Left chelicera of male, upper view. B : Ditto, lower view. C : Ditto, lateral view. D : Distal portio of coductor ad embolus of male. E : Paracymbium of male. F : Left chelicera of female, upper view. G : Ditto, lower view. H : Abdome of male, lateral view. I : Geital fold of female. J : Eye group of male.

34 TETRAGNATHA OF THE AUSTRALASIA 69 Palpi. Paracymbium sleder, straight ad poited at tip; coductor ad embolus uique, as show i Fig. 20, F & G. Abdome. Very log, about 10 times as log as broad, ot exteded posteriorly to spierets. Color i alcohol. Geerally pale grayish brow, without ay markigs. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet slightly shorter tha oe-half of carapace ; (Gu), (e) ad (T) preset, (rsu) 3-4 i umber ; (Gl) preset, (rsl) 8-10 i umber ; fag uarmed. Legs. Nearly as i male ; relative legths of legs as show i Table 18. Abdome. Very log, about 7.5 times as log as broad, ot exteded posteriorly to spierets ; geital fold as show i Fig. 20, I. Color i alcohol. Similar to male. Type material : Holotype B (BISHOP 13636), NEW GUINEA (IRIAN JAYA), Fak fak Bombarai, 6-8. VI. 1959, T. Maa (BMH). Paratypes : 20 79, same data as holotype. NEW GUINEA (PNG) : 10, Murua, Gulf Prov., 18. XII. 1964, J. L. Gressitt (BMH). Type depository : The holotype preserved i the collectio of BMH. Distributio : New Guiea. Note : This ew species is related to T. biseriata Thorell ad T. isuluris sp. ov. but it may be Fig. 20. Tetragatha aa sp. ov. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view. D : Left chelicera of female, upper view. E : Ditto, lower view. F : Left palpus of male. G : Distal portio of coductor ad embolus of male. H : Paracymbium of male. I : Geital fold of female. J : Carapace of male. K : Abdome of male, lateral view.

35 70 C. OKUMA easily distiguished from them by the characteristic male palpal coductor ad paracymbium as show i Fig. 20, F, G ad H. The radiata-group Diagosis. Posterior row of eyes about as wide as or slightly arrower tha aterior row of eyes ; lateral eyes as close as media eyes ; cheliceral legth shorter tha carapace i both sexes ; abdome oval, with white markigs. This group is composed of three species which are very distictive i the geus. K EY TO THE SPECIES 1. Color geerally light brow to grayish brow ; abdomial white markigs iterrupted ad idistict , rad&ta - Color geerally glossy brow, abdomial white markigs distict, etire or iterrupted Abdome with elogate markigs (stripes) etire - Abdome with elogate markigs iterrupted i the middle... * motico/a... * s@&2 Tetragatha radiata Chrysathus, 1975 sp. ov. sp. ov. Tetragatha radiata Chrysathus, 1975, Zool. Verhadl., 14 : 15. Male. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth 1.0 mm. Eyes. Aterior row of eyes occupyig full width of head, posterior row of eyes about as wide as or slightly arrower tha aterior oe ( : 100) ; viewed from above, aterior row somewhat strogly recurved ad posterior row somewhat getly recurved, so that lateral eyes somewhat closer tha media eyes ; cetral ocular quadragle with posterior ocular width slightly larger tha aterior ocular width (118 : loo), ad early equal to atero-posterior ocular legth ; ratio of diameter of AME : ALE : PME : PLE as 75 : 63 : 75 : 70 (p). Chelicerae. Basal segmet about 0.6 times as log as carapace ; (a) acumiated at tip, cospicuously robust at base, with a small basal tubercle as show i Fig. 21, A, B ad C ; (sl) slightly directed forward, (T) somewhat large, (rsu) 3-4 i umber ; (Gl) ormal, (L2) slightly curved backward, (rsl) 4 i umber ; fag uarmed. Legs. Leg formula ; all legs with spies, first femora with about 6 spies ; relative legths of legs as show i Table 21. Palpi. Paracymbium ot bifurcated at tip ; coductor ad embolus cosiderably complicated i structure ; coductor twisted ad spatulate at tip as show i Fig. 21, E, F ad G. Abdome. Not log, about 2 times as log as broad, ot exteded posteriorly to spierets. Color i alcohol. Chelicerae reddish brow ; media part of carapace reddish brow to brow, radiated as show i Fig. 21, D, lateral margis yellowish white ; maxillae, labium ad sterum dusky brow ; abdome geerally grayish brow, covered with dusky silvery spagles dorsally, ad with idistict white markigs dorso- ad vetro-laterally. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width 1.4 mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Nearly as i male. Chelicerae. Basal segmet about 0.6 times as log as carapace ; (Gu) ad (T) preset, (rsu) 4 i

36 TETRAGNATHA OF THE AUSTRALASIA 71 umber ; (Gl) preset, (rsl) 3-4 i umber, (L2) ad (L3) ofte coalescet as show i Fig. 21, K ; fag uarmed. Legs. Nearly as i male ; relative legths of legs as show i Table 21. Abdome. Nearly as i male ; geital fold as show i Fig. 21, I. Color i alcohol. Similar to male but abdomial white markigs more distict. Specimes examied : NEW GUINEA (PNG) : 29, Mt. Wilhelm, 3,600 m, 2. VI. 1955, J. L. E F... : : :... J.....,... K M Fig. 21. Tetragatha radiata Chrysathus. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view. D : Female. E, F ad G : Distal portio of coductor ad embolus of male, H : Paracybium of male. I : Geital fold of female. J : Left chelicera of female, upper view. K : Ditto, lower view. L : Eye group of male. M : Abdome of male, lateral view.

37 72 C. OKUMA Gressitt ; 20, Mt. Wilhelm, 3,800 m, 3. VII. 1963, J. Sedlacek ; 10, Mt. Wilhelm, m, 3, V ; 20 29, Mt. Hage, 1,650 m, 28. V. 1966, J. L. Gressitt ; lb 20, Mt. Giluwe N. side 300-3,300 m, J. L. Gressitt, (all BMH). Distributio : New Guiea. Tetragatha moticola sp. ov. Male. Body legth, exclusive of chelicerae, 4.1 mm ; carapacial legth 1.6 mm, width 1.0 mm ; abdomial legth 2.5 mm, width 1.0 mm ; cheliceral legth 1.2 mm. Eyes. Aterior row of eyes occupyig full width of head ad lateral eyes somewhat produced ; posterior row of eyes somewhat arrower tha aterior oe (96 : 100) ; viewed from above, aterior row somewhat strogly recurved ad posterior row rather getly recurved, so that lateral eyes slightly closer tha media eyes ; cetral ocular quadragle with posterior ocular width slightly larger tha aterior ocular width (109 : loo), ad larger tha atero-posterior ocular legth (120 : 100) ; ratio of diameter of AME : ALE : PME : PLE as 80 : 50 : 75 : 50 (p). Chelicerae. Basal segmet about 0.75 times as log as carapace ; (a) blutly at tip ; (sl) slightly directed forward, (T) large, slightly curved backward, (rsu) 5 i umber ; (Gl) small, (L2) cosiderably large, slightly curved backward, (rsl) oly 3 i umber; fag rather log, with a subbasal ier tubercle obtuse. Legs. Leg formula ; all legs with oly a few spies, first femora with 3 spies ear distal ed ; relative legths of legs as show i Table 22. Palpi. Paracymbium ot bifurcated at tip, a small lobe somewhat closer apex tha to base ; coductor rather sleder at distal ed, embolus distictly dilated at tip as show i Fig. 22, D & F. Abdome. Not so log, about 2.5 times as log as broad, ot exteded posteriorly to spierets. Color i alcohol. Legs, carapace ad chelicerae glossy brow ; maxillae, lip ad sterum glossy blackish brow ; abdome light glossy blackish brow, without spagles. Female. Body legth, exclusive of chelicerae, 4.5 mm ; carapacial legth 1.7 mm, width 1.0 mm ; abdomial legth 2.8 mm, width 1.2 mm ; cheliceral legth 1.1 mm. Eyes. Nearly as i male. Chelicerae. Basal segmet about 0.65 times as log as carapace ; somewhat swolle subapically ; (Gu) ad (T) preset, (rsu) 6 i umber ; (Gl) preset, (rsl) 5-6 i umber ; fag uarmed. Legs. Nearly as i male ; relative legths of legs as show i Table 22. Abdome. Nearly as i male ; geital fold as show i Fig. 22, I. Color i alcohol. Similar to male but abdome with a white stripe alog dorso-lateral margis. Type material : Holotype c? (BISHOP 13637), NEW GUINEA (PNG), Wau 1,200 m, Morobe Prov., 13. III. 1969, J. Sedlacek (BMH). Paratype : 19, NEW GUINEA (PNG), Mt. Kaidi, Morobe Prov., 17. VII. 1982, 0. Tadauchi (KUF). Type depository : The holotype is preserved i the collectio of BMH. Distributio : New Guiea (PNG). Note : This is a strikig species, which is closer to T. sigata sp. ov. rather tha to T. radiata, but it may be easily distiguished from them by the male chelicera ad palpus as show i Fig. 22. The abdomial markigs of the female of this ew species are also differet from those of sigata ad radiatu. Tetragatha sigata sp. ov. Male. Body legth, exclusive of chelicerae, 3.6 mm ; carapacial legth 1.6 mm, width 1.0 mm ;

38 TETRAGNATHA OF THE AUSTRALASIA 73 Fig. 22. Tetragathu moticola sp. ov. A : Left chelicera of male, upper view. B : Ditto, lower view. C : Ditto, lateral view. D : Distal portio of coductor ad embolus of male. E : Paracymbium of male. F : Left palpus of male. G : Eye group of male. H : Abdome of female, lateral view. I : Geital fold of female. J : Left chelicera of female, upper view. K : Ditto, ier view. abdomial legth 2.1 mm, width 1.0 mm ; cheliceral legth 1.0 mm. Eyes. Aterior row of eyes occupyig full width of head, lateral eyes somewhat produced ; posterior row of eyes somewhat smaller tha aterior oe (97 : 100) ; viewed from above, two rows early parallel ad getly recurved ; cetral ocular quadragle with posterior ocular width slightly larger tha aterior ocular width (120 : loo), ad larger tha atero-posterior ocular legth i early same ratio as the latter ; ratio of diameter of AME : ALE : PME : PLE as 80 : 75 : 75 : 75 (p). Chelicerae. Basal segmet about 0.6 times as log as carapace ; (a) cosiderably thick, bifurcated at tip ; (sl) slightly directed backward, (T) moderately large, (rsu) 6 i umber ; (Gl) directed iward, (rsl) 3-5 i umber ; fag thick, log, ad with a ier obtuse tubercle subbasally. Legs. Leg formula ; all legs with oly a few spies, first femora with 2 spies ear distal ed ; relative legths of legs as show i fable 23. Palpi. Paracymbium bifurcated at tip ; coductor ad embolus simple, sleder. Abdome, Not so log, about 2 times as log as broad, ot exteded posteriorly to spierets. Color i alcohol. Chelicerae, carapace ad sterum light brow, carapace with silvery white markigs laterally ; abdome geerally grayish brow, with two pairs of silvery white markigs dorso-laterally. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width about 1.0 mm ; abdomial legth mm, width mm ; cheliceral legth 0.9 mm.

39 74 COKUMA Eyes. Nearly as i male. Chelicerae. Basal segmet about 0.6 times as log as carapace, somewhat swolle o lower side subapically ; (Gu) ad (T) preset, (rsu) 5-6 i umber; (Gl) preset, (rsl) 4-5 i umber ; fag uarmed. Legs. Nearly as i male, first femora with oly oe spie ear distal ed ; relative legths of legs as show i Table 23. Abdome. Nearly as i male ; geital fold as show i Fig. 23, I. Color i alcohol. Similar to male. Type material : Holotype 0 (BISHOP 13638), NEW GUINEA (PNG), Wharto Rd., 2,100-2,850 m, 4. XI. 1965, J. Sedlacek (BMH). Paratypes: ZQ, same data as holotype ; 19, Garaia, 800 m, Morobe Prov., 16. I. 1968, J. Sedlacek (BMH). Type depository : The holotype is preserved i the collectio of BMH. Distributio : New Guiea (PNG). Note : This ew species is also very distictive i the geus, ad is related to T. moticola sp. ov., but it may be easily distiguished from the latter by the male chelicera ad palpus, ad the abdomial markigs of both sexes are also differet from those of the latter. The serra-group Diagosis. Posterior row of eyes slightly larger tha aterior oe ; cetral ocular quadragle early Fig. 23. Tetragatha sigata sp. ov. A : Left chelicera of male, upper view. B : Ditto, lateral view. C : Ditto, lower view. D : Left palpus of male. E : Paracymbium of male. F: Left chelicera of female, upper view. G : Ditto, lower view. H : Female. I : Geital fold of female. J : Eye group of male. K : Abdome of female, lateral view.

40 TETRAGNATHA OF THE AUSTRALASIA 75 square ; female chelicera with log (s) ear outer apex ; fag geiculate. KEY TO THE SPECIES (FEMALE) 1. Fag with (EX) ear base; (Gu) robust, directed iward * ser ya - Fag without (EX) ; (Gu) small, directed forward...*..+.-**.....* hiya_yhimi sp_ av. Tetragatha serra Doleschall, 1857 Tetragathu sem Doleschall, 1857, Naturk. Tijdschr. Ned. Id., 13 : 408 ; Thorell, 1878, A. Mus. Civ. Geova, 13 : 111 ; Meria, 1911, Jahrb. Syst., 31(Z) : 182 ; Gravely, 1921, Rec. Id. Mus., 22 : 433 ; Roewer, 1942, Katalog der Araeae, 1: 983. Female. Body legth, exclusive of chelicerae, 9.1 mm ; carapacial legth 3.7 mm, width 1.6 mm ; abdomial legth 5.5 mm, width 1.0 mm ; cheliceral legth 2.5 mm. Eyes. Aterior row of eyes occupyig full width of head ; posterior row of eyes slightly wider tha aterior oe (106 : 100) ; viewed from above, aterior row somewhat strogly recurved ad posterior row moderately recurved, so that lateral eyes somewhat closer tha media eyes ; cetral ocular quadragle early square ; ratio of diameter of AME : ALE : PME : PLE as 150 : 100 : 125 : 113 (/1). Chelicerae. Cospicuously developed, basal segmet about 0.7 times as log as carapace ; (s) preset at outer side ear apex, log, acumiate at tip ; (Gu) robust, directed iward, (T) large, slightly directed forward, (rsu) 5 i umber ; (Gl) ot so large, (rsl) 11 i umber ; fag somewhat geiculate, with (EX) ear base. Legs, Leg formula ; all legs with spies, first femora with about 15 spies ; relative legths of legs as show i Table 24. Abdome. Cosiderably log, about 5 times as log as broad, ot exteded posteriorly to spierets; geital fold as show i Fig. 24, E. Color i alcohol. Geerally yellowish brow ; abdome geerally grayish brow, with idistict logitudial brow markigs ad 5 or 6 pairs of dark brow spots dorso-laterally. Specime examied : NEW GUINEA (IRIAN JAYA) : 19, Nabire, H. Holtma Jugle, alog river, 20. IX (BMH). Distributio : Java, Sumatra, Celebes, Amboia ad New Guiea (ew record). Tetragatha hirashimai sp. ov. Male. Abdomial legth 7.1 mm, width 1.4 mm ; cheliceral legth 2.4 mm. Chelicerae. Cosiderably developed, basal segmet with (a), ad a small apophysis at outer side ear apex ; (Gu) ad (T) early equal i size, (rsu) 6 i umber ; (Gl) small, (L2) robust, largest, (rsl) 9 i umber ; fag log, slightly geiculate, uarmed. Legs. Ijured. Palpi. Paracymbium broad at the middle, bifurcated at tip, * coductor of embolus somewhat twisted at the middle ad hooked at tip. Abdome. Nearly as i female. Female. Body legth, exclusive of chelicerae, mm ; carapacial legth mm, width mm ; abdomial legth mm, width mm ; cheliceral legth mm. Eyes. Aterior row of eyes occupyig full width of head ; posterior row of eyes slightly larger tha aterior oe (105 : 100) ; viewed from above, aterior row somewhat strogly recurved ad

41 76 C. OKUMA Fig. 24. Tetragatha serra Doleschall. A : Left chelicera of female, upper view. B : Ditto, lateral view. C : Ditto, lower view. D : Eye group of female. E : Geital fold of female. F : Abdome of female, lateral view. posterior row moderately recurved, so that lateral eyes somewhat closer tha media eyes; cetral ocular quadragle early square ; ratio of diameter of AME : ALE : PME : PLE as 150 : 113 : 138 : 125 (p). Chelicerae. Cospicuously developed as i male, basal segmet about 0.8 times as log as carapace, lower side strogly swolle at 3/4 from base ; (s) preset o outer side ear apex, log, acumiate at tip, directed forward ; (T) widely separated from (Gu), (rsu) 7-8 i umber; (AXl) preset, very small, (Gl) large, directed forward, (rsl) i umber but some basal oes very miute ; fag strogly geiculate, uarmed basally, but with a uique projectio ear the middle. Legs, Leg formula ; all legs with may spies, first femora with about 15 spies ; relative legths of legs as show i Table 25. Abdome. Cosiderably log about 4-5 times as log as broad, ot exteded posteriorly to spierets ; geital fold as show i Fig. 26, I. Color i alcohol. Legs ad chelicerae grayish brow ; maxillae, lip, fag ad sterum dark brow ; carapace grayish brow, lateral margis blackish ; a media blackeed stripe o carapace ot reachig the apex of the latter; abdome geerally grayish brow, with logitudial blackish markigs ad yellowish silver spagles dorsally. Type material : Holotype 9 (BISHOP 13639), NEW GUINEA (PNG), Sogeri, Cetral Prov., 24. VIII. 1982, Y. Hirashima (BMH). Paratypes: Id 29, NEW GUINEA (PNG), Mioa Rubber Est. 9 mile E. Sogeri, 1,600 m, Cetral Prov., 19. VI. 1959, D. E. Hardy (BMH). Type depository : The holotype is preserved i the collectio of BMH. Distributio : New Guiea. Note : This ew species is related to T. sewa Doleschall, 1857, T. j7ugezzas Hasselt, 1882 ad T. delumbis Thorell, 1891, but is easily distiguished from them by the shape of characteristic chelicerae

42 TETRAGNATHA OF THE AUSTRALASIA 77 Fig. 25. Tetragatha hirashimai sp. ov. Female.